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<document id="EEAA2216D8423C53ABEBF74FF90BF329" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D6A9223FDC6FE41C9FD2AAD" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius tarsalis Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="121" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="119" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="85C3A21C0E39B24E10B2FBA84673A584">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="609DBED04AD39312FB3C9AF54E5FDCF6">Telnov, Dmitry</mods:namePart>
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<mods:title id="7C5E2E05591496F7E362FF4D66DD75E8">European Journal of Taxonomy</mods:title>
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<mods:date id="7247C61FE868967F6FCE2595D7162DEA">2020</mods:date>
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<mods:number id="EE48FB80E469B2DF489E8B55E0B5DC74">2020-10-19</mods:number>
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<mods:number id="0E5DE2FA5CEF5E24BE7A2747C2F6F4A6">721</mods:number>
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<mods:identifier id="798729C48353C63616F4C68D84D1DCF6" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D6A9223FDC6FE41C9FD2AAD" ID-DOI="http://doi.org/10.5281/zenodo.4344233" ID-GBIF-Taxon="168648176" ID-Zenodo-Dep="4344233" LSID="urn:lsid:plazi:treatment:1A3787F69D6A9223FDC6FE41C9FD2AAD" httpUri="http://treatment.plazi.org/id/1A3787F69D6A9223FDC6FE41C9FD2AAD" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119">
<subSubSection id="DA84656B9D6A922DFDC6FE41CB882A78" box="[597,990,511,538]" pageId="118" pageNumber="119" type="nomenclature">
<paragraph id="922136E09D6A922DFDC6FE41CB882A78" blockId="118.[457,1131,511,574]" box="[597,990,511,538]" pageId="118" pageNumber="119">
<heading id="C969818C9D6A922DFDC6FE41CB882A78" box="[597,990,511,538]" centered="true" fontSize="11" level="2" pageId="118" pageNumber="119" reason="2">
<taxonomicName id="559E4D639D6A922DFDC6FE41CB882A78" ID-CoL="3XFLP" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[597,990,511,538]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="A0EAEAF29D6A922DFDC6FE41CB152A78" bold="true" box="[597,835,511,538]" italics="true" pageId="118" pageNumber="119">Maechidius tarsalis</emphasis>
<bibRefCitation id="F60F4B119D6A922DFCD9FDBECB882A78" author="Arrow G. E." box="[842,990,512,538]" pageId="118" pageNumber="119" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A922DFE5AFD9DCC3D2A5F" box="[457,1131,547,574]" pageId="118" pageNumber="119" type="description">
<paragraph id="922136E09D6A922DFE5AFD9DCC3D2A5F" blockId="118.[457,1131,511,574]" box="[457,1131,547,574]" pageId="118" pageNumber="119">
<figureCitation id="0AA52A659D6A922DFE5AFD9DCA752A5F" box="[457,547,547,573]" captionStart="Figs 90–93" captionStartId="119.[189,241,1928,1954]" captionTargetBox="[329,1264,264,1883]" captionTargetId="figure-60@119.[321,1264,256,1905]" captionTargetPageId="119" captionText="Figs 90–93. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 90. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 91. M. trivialis sp. nov., holotype, ♂ (BMNH). 92. M. ursus sp. nov., holotype, ♀ (NME). 93. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122169" httpUri="https://zenodo.org/record/4122169/files/figure.png" pageId="118" pageNumber="119">Figs 90</figureCitation>
,
<figureCitation id="0AA52A659D6A922DFDBCFD9DCA082A5F" box="[559,606,547,573]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="118" pageNumber="119">186</figureCitation>
,
<figureCitation id="0AA52A659D6A922DFDFAFD9DCACE2A5C" box="[617,664,547,574]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="118" pageNumber="119">259</figureCitation>
,
<figureCitation id="0AA52A659D6A922DFD37FD9DCA852A5C" box="[676,723,547,574]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="118" pageNumber="119">350</figureCitation>
, 422, 435, 437, 526, 565, 730–732
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A922DFF2EFDDBCD212BC2" pageId="118" pageNumber="119" type="materials_examined">
<paragraph id="922136E09D6A922DFF2EFDDBC93A2A1C" blockId="118.[189,364,612,639]" box="[189,364,612,639]" pageId="118" pageNumber="119">
<heading id="C969818C9D6A922DFF2EFDDBC93A2A1C" bold="true" box="[189,364,612,639]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="A0EAEAF29D6A922DFF2EFDDBC93A2A1C" bold="true" box="[189,364,612,639]" pageId="118" pageNumber="119">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6A922DFF79FD2DCA3B2ACC" blockId="118.[189,1399,659,792]" box="[234,621,659,686]" pageId="118" pageNumber="119">
<emphasis id="A0EAEAF29D6A922DFF79FD2DC9302ACF" bold="true" box="[234,358,659,685]" pageId="118" pageNumber="119">
<typeStatus id="4D2588429D6A922DFF79FD2DC9302ACF" box="[234,358,659,685]" pageId="118" pageNumber="119" type="lectotype">Lectotype</typeStatus>
</emphasis>
(herewith designated)
</paragraph>
<paragraph id="922136E09D6A922DFF2EFD09CA182B7A" blockId="118.[189,1399,659,792]" pageId="118" pageNumber="119">
<materialsCitation id="22F63CBD9D6A922DFF2EFD09CA182B7A" ID-GBIF-Occurrence="2898878468" collectingDate="1933-08" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="366" location="Type" pageId="118" pageNumber="119" specimenCount="1" typeStatus="syntype">
<collectingCountry id="EA8976709D6A922DFF2EFD09C9882AB3" box="[189,478,695,721]" name="Papua New Guinea" pageId="118" pageNumber="119">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D6A922DFE6FFD09CA422AB3" box="[508,532,695,721]" pageId="118" pageNumber="119">♂</specimenCount>
; “
<location id="9741603B9D6A922DFDBDFD09CA3D2AB3" LSID="urn:lsid:plazi:treatment:1A3787F69D6A9223FDC6FE41C9FD2AAD:9741603B9D6A922DFDBDFD09CA3D2AB3" box="[558,619,695,721]" country="Papua New Guinea" name="Type" pageId="118" pageNumber="119">
<typeStatus id="4D2588429D6A922DFDBDFD09CA3D2AB3" box="[558,619,695,721]" pageId="118" pageNumber="119">Type</typeStatus>
</location>
[p, label circular, red frame] //
<typeStatus id="4D2588429D6A922DFC73FD09CC322AB3" box="[992,1124,695,721]" pageId="118" pageNumber="119" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] //
<specimenCount id="8498FD699D6A922DFEBDFD65C9122A97" box="[302,324,731,757]" pageId="118" pageNumber="119">♂</specimenCount>
[p]//
<collectionCode id="F48FAE259D6A922DFE14FD64C9BF2A96" box="[391,489,730,756]" pageId="118" pageNumber="119">PAPUA</collectionCode>
: Kokoda.
<quantity id="55669B059D6A922DFDF7FD64CA952A96" box="[612,707,730,756]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="118" pageNumber="119" unit="ft" value="1200.0">
<elevation id="19B3D1D39D6A922DFDF7FD64CA952A96" box="[612,707,730,756]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="118" pageNumber="119" unit="ft" value="1200.0">1,200ft.</elevation>
</quantity>
<date id="E62010209D6A922DFD51FD64CB7A2A96" box="[706,812,730,756]" pageId="118" pageNumber="119" value="1933-08">
<collectingDate id="F664E9C89D6A922DFD51FD64CB7A2A96" box="[706,812,730,756]" pageId="118" pageNumber="119" value="1933-08">viii.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D6A922DFCAAFD64CBBB2A96" box="[825,1005,730,756]" pageId="118" pageNumber="119">L.E.Cheesman.</collectorName>
B.M.1933-577. [p] //
<taxonomicName id="559E4D639D6A922DFB7DFD64C9432B75" authorityName="Arrow" authorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="A0EAEAF29D6A922DFB7DFD64C9432B75" italics="true" pageId="118" pageNumber="119">Maechidius tarsalis</emphasis>
</taxonomicName>
,
<typeStatus id="4D2588429D6A922DFEB7FD40C9012B7A" box="[292,343,766,792]" pageId="118" pageNumber="119">type</typeStatus>
Arrow [h]”;
<collectionCode id="F48FAE259D6A922DFE7DFD40CA1C2B7A" box="[494,586,766,792]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="118" pageNumber="119">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D6A922DFF79FC81CACB2B38" blockId="118.[189,1399,831,928]" box="[234,669,831,858]" pageId="118" pageNumber="119">
<emphasis id="A0EAEAF29D6A922DFF79FC81C9C12B3B" bold="true" box="[234,407,831,857]" pageId="118" pageNumber="119">
<typeStatus id="4D2588429D6A922DFF79FC81C9C12B3B" box="[234,407,831,857]" pageId="118" pageNumber="119" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(herewith designated)
</paragraph>
<paragraph id="922136E09D6A922DFF2EFCDCCD212BC2" blockId="118.[189,1399,831,928]" pageId="118" pageNumber="119">
<materialsCitation id="22F63CBD9D6A922DFF2EFCDCCD252BC2" ID-GBIF-Occurrence="2898878592" collectingDate="1933-08" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="366" location="PAPUA NEW GUINEA" pageId="118" pageNumber="119" specimenCount="1" typeStatus="syntype">
<collectingCountry id="EA8976709D6A922DFF2EFCDCC98D2B1E" box="[189,475,866,893]" name="Papua New Guinea" pageId="118" pageNumber="119">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D6A922DFE64FCDCCA702B1F" box="[503,550,866,893]" pageId="118" pageNumber="119" type="male">1 ♂</specimenCount>
; “
<typeStatus id="4D2588429D6A922DFDACFCDCCA902B1E" box="[575,710,866,892]" pageId="118" pageNumber="119" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] //
<specimenCount id="8498FD699D6A922DFBDFFCDDCC342B1F" box="[1100,1122,867,893]" pageId="118" pageNumber="119">♂</specimenCount>
[p] //
<collectionCode id="F48FAE259D6A922DFB37FCDCCD502B1E" box="[1188,1286,866,892]" pageId="118" pageNumber="119">PAPUA</collectionCode>
: Kokoda.
<quantity id="55669B059D6A922DFF2EFC3BC94A2BFD" box="[189,284,901,927]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="118" pageNumber="119" unit="ft" value="1200.0">
<elevation id="19B3D1D39D6A922DFF2EFC3BC94A2BFD" box="[189,284,901,927]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="118" pageNumber="119" unit="ft" value="1200.0">1,200ft.</elevation>
</quantity>
<date id="E62010209D6A922DFE8FFC3BC9D32BFD" box="[284,389,901,927]" pageId="118" pageNumber="119" value="1933-08">
<collectingDate id="F664E9C89D6A922DFE8FFC3BC9D32BFD" box="[284,389,901,927]" pageId="118" pageNumber="119" value="1933-08">viii.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D6A922DFE1DFC3BCA142BFD" box="[398,578,901,927]" pageId="118" pageNumber="119">L.E.Cheesman.</collectorName>
B.M.1933-577. [p]//
<taxonomicName id="559E4D639D6A922DFCA9FC3BCC762BFD" authorityName="Arrow" authorityYear="1941" box="[826,1056,901,927]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="A0EAEAF29D6A922DFCA9FC3BCC762BFD" box="[826,1056,901,927]" italics="true" pageId="118" pageNumber="119">Maechidius tarsalis</emphasis>
</taxonomicName>
, co-type Arrow [h]”;
<collectionCode id="F48FAE259D6A922DFA84FC38CD252BC2" box="[1303,1395,902,928]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="118" pageNumber="119">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A922DFF2EFC79CA222C29" pageId="118" pageNumber="119" type="reference_group">
<paragraph id="922136E09D6A922DFF2EFC79CA222C29" blockId="118.[189,1399,967,1100]" pageId="118" pageNumber="119">
<bibRefCitation id="F60F4B119D6A922DFF2EFC79C9F52B80" author="Arrow G. E." box="[189,419,967,994]" pageId="118" pageNumber="119" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow (1941: 454)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D6A922DFD48FC76CB2B2B83" ID-CoL="3XFH6" authorityName="Heller" authorityYear="1910" box="[731,893,968,993]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D6A922DFD48FC76CB2B2B83" box="[731,893,968,993]" italics="true" pageId="118" pageNumber="119">M. aenescens</emphasis>
</taxonomicName>
on a number of specimens, although not explicitly stated, and gives a range of sizes. The
<typeStatus id="4D2588429D6A922DFD60FC54CB092C66" box="[755,863,1002,1028]" pageId="118" pageNumber="119" type="lectotype">lectotype</typeStatus>
is designated in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other (non-conspecific)
<typeStatus id="4D2588429D6A922DFF2EFB8CC9722C2E" box="[189,292,1074,1100]" pageId="118" pageNumber="119" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A9223FF2EFBCCC9FD2AAD" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119" type="description">
<paragraph id="922136E09D6A922DFF2EFBCCC9182CEE" blockId="118.[189,334,1138,1164]" box="[189,334,1138,1164]" pageId="118" pageNumber="119">
<heading id="C969818C9D6A922DFF2EFBCCC9182CEE" bold="true" box="[189,334,1138,1164]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="A0EAEAF29D6A922DFF2EFBCCC9182CEE" bold="true" box="[189,334,1138,1164]" pageId="118" pageNumber="119">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6A9223FF2EFB1CCC7F2A3D" blockId="118.[189,1399,1185,2025]" lastBlockId="120.[189,1399,265,608]" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119">
Dorsum and venter uniformly brown, margins of labroclypeus and legs somewhat paler. Head flattened dorsally, shallowly impressed at either anterolateral angle of labroclypeus. Male labroclypeus (
<figureCitation id="0AA52A659D6A922DFA98FB7BCD382C82" box="[1291,1390,1221,1248]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="118" pageNumber="119">Fig. 186</figureCitation>
) broadly shallowly emarginate on anterior margin, its lateral margins strongly sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, slightly protruding. Canthus broadly rounded in dorsal view. Head punctures irregularly hexagonal, horseshoe-shaped on frons and vertex, very deep and dense. Intervening spaces much smaller than punctures, glossy on forehead, in part microreticulate on frons. Head setae inconspicuous, suberect, slightly surpassing length of corresponding punctures. Pronotum strongly transverse, subopaque dorsally and laterally, with shallow circular dorsal impression on either lateral third. Anterior margin of pronotum slightly sinuous with anterolateral angles protruding. Basal margin of pronotum broadly rounded. Lateral margin of pronotum in dorsal view in anterior half nearly straight to slightly widened towards middle, strongly emarginate in prebasal third, strongly crenulate all along except in emargination area (
<figureCitation id="0AA52A659D6A922DFD68F998CB332E23" box="[763,869,1574,1601]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="118" pageNumber="119">Fig. 259</figureCitation>
); intervals between crenulae deep and rather large. Inconspicuous erect short seta present between every two crenulae. Lateral margin of pronotum arched in lateral view. Hypomeron separated from prosternum by moderately high nearly straight carina which is obtusely dentate medially, with long setae on anterolateral margin. Antennal pocket deep. Pronotal punctures irregularly hexagonal, very deep and dense, in part horseshoe-shaped. Intervening spaces much smaller than punctures, glossy to microreticulate. Pronotal setae inconspicuous, rise from anterior margin of each puncture; not or slightly surpassing length of corresponding puncture. Lateral and basal margins, antero- and posterolateral angles and hypomeron covered with microscopical velvety pubescence. Scutellar shield covered with microscopical velvety pubescence, narrowly rounded apically. Elytra slightly widened postmedially, opaque dorsally. Elytron with elevated and broadly interrupted tracks of four longitudinal carinae, including sutural one. Elytral punctures sinuous (elongate and narrow), incision-shaped, moderately deep and dense (
<figureCitation id="0AA52A659D6A922DFC31F815CC472FA4" box="[930,1041,1963,1990]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="118" pageNumber="119">Fig. 350</figureCitation>
). Intervening spaces densely microreticulate; reticulation somewhat less prominent on elevated parts, forming tracks of elytral carinae. Elytral setae inconspicuous, suberect; each seta rises from anterior margin of each incision, not or slightly surpassing its length. Stronger erect not much longer setae present in irregular longitudinal rows along remnants of carinae. Abdominal sternites medially each with bunch of erect golden setae (
<figureCitation id="0AA52A659D649223FF55FECCC96629EF" box="[198,304,370,397]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="120" pageNumber="121">Fig. 435</figureCitation>
). Male pygidium flattened dorsally, with large dense shallow annular punctures (Fig. 526). Intervening spaces much smaller than punctures, densely microreticulate. Setae of pygidium sparse, suberect, surpassing length of corresponding punctures; each seta rises either from anterior margin or centre of corresponding puncture. Male protibia with three distal teeth on external margin, of which distal one prolonged and basal one inconspicuous and strongly obtuse (Fig. 422). Male basal metatarsomere leaf-like, flat and slightly convex dorsally, with brush of long dense setae on either lateroventral margin (
<figureCitation id="0AA52A659D649223FF55FDFBC9782A3D" box="[198,302,581,607]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="120" pageNumber="121">Fig. 437</figureCitation>
). Spiculum gastrale as in
<figureCitation id="0AA52A659D649223FDCAFDFBCAE92A02" box="[601,703,581,608]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="120" pageNumber="121">Fig. 565</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D649223FC12FDFBCC732A3D" box="[897,1061,581,607]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="120" pageNumber="121">Figs 730–732</figureCitation>
.
</paragraph>
<caption id="C6E166689D6B922CFF2EF836CC512F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122169" ID-Zenodo-Dep="4122169" httpUri="https://zenodo.org/record/4122169/files/figure.png" pageId="119" pageNumber="120" startId="119.[189,241,1928,1954]" targetBox="[329,1264,264,1883]" targetPageId="119">
<paragraph id="922136E09D6B922CFF2EF836CC512F8A" blockId="119.[189,1399,1928,2025]" pageId="119" pageNumber="120">
<emphasis id="A0EAEAF29D6B922CFF2EF836C91B2FC0" bold="true" box="[189,333,1928,1954]" pageId="119" pageNumber="120">Figs 90–93.</emphasis>
Papuan
<taxonomicName id="559E4D639D6B922CFE28F836CB552FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[443,771,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D6B922CFE28F836CA122FC0" box="[443,580,1928,1954]" italics="true" pageId="119" pageNumber="120">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D6B922CFDC3F836CB552FC0" author="Macleay W. S." box="[592,771,1928,1954]" pageId="119" pageNumber="120" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D6B922CFB8AF836CC602FC0" bold="true" box="[1049,1078,1928,1954]" pageId="119" pageNumber="120">90</emphasis>
.
<taxonomicName id="559E4D639D6B922CFBDAF837CD272FC0" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1097,1393,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="A0EAEAF29D6B922CFBDAF837CC9B2FC0" box="[1097,1229,1928,1954]" italics="true" pageId="119" pageNumber="120">M. tarsalis</emphasis>
<bibRefCitation id="F60F4B119D6B922CFB44F836CD272FC0" author="Arrow G. E." box="[1239,1393,1928,1954]" pageId="119" pageNumber="120" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D6B922CFE5FF815C9BF2FA7" bold="true" box="[460,489,1963,1989]" pageId="119" pageNumber="120">91</emphasis>
.
<taxonomicName id="559E4D639D6B922CFE65F812CA2E2FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[502,632,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="trivialis" status="sp. nov.">
<emphasis id="A0EAEAF29D6B922CFE65F812CA2E2FA7" box="[502,632,1963,1989]" italics="true" pageId="119" pageNumber="120">M. trivialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D6B922CFDEDF812CA8C2FA4" box="[638,730,1964,1990]" pageId="119" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D6B922CFC7CF815CC5A2FA7" bold="true" box="[1007,1036,1963,1989]" pageId="119" pageNumber="120">92</emphasis>
.
<taxonomicName id="559E4D639D6B922CFB89F812CCD62FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1050,1152,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="A0EAEAF29D6B922CFB89F812CCD62FA7" box="[1050,1152,1964,1989]" italics="true" pageId="119" pageNumber="120">M. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D6B922CFB16F812CCB72FA4" box="[1157,1249,1964,1990]" pageId="119" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (NME).
<emphasis id="A0EAEAF29D6B922CFEB3F870C96B2F8A" bold="true" box="[288,317,1998,2024]" pageId="119" pageNumber="120">93</emphasis>
.
<taxonomicName id="559E4D639D6B922CFEDFF871CA342F8A" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[332,610,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="A0EAEAF29D6B922CFEDFF871C9902F8A" box="[332,454,1999,2024]" italics="true" pageId="119" pageNumber="120">M. vicinus</emphasis>
<bibRefCitation id="F60F4B119D6B922CFE5DF870CA342F8A" author="Heller K. M." box="[462,610,1998,2024]" pageId="119" pageNumber="120" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, lectotype, ♀ (SNSD). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D649223FF2EFD38C9E62AC2" blockId="120.[189,432,646,672]" box="[189,432,646,672]" pageId="120" pageNumber="121">
<heading id="C969818C9D649223FF2EFD38C9E62AC2" bold="true" box="[189,432,646,672]" fontSize="11" level="3" pageId="120" pageNumber="121" reason="3">
<emphasis id="A0EAEAF29D649223FF2EFD38C9E62AC2" bold="true" box="[189,432,646,672]" pageId="120" pageNumber="121">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D649223FF2EFD0BC9FD2AAD" blockId="120.[189,427,693,719]" box="[189,427,693,719]" pageId="120" pageNumber="121">Female is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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