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<document id="95A0F55111406C3EAE8E32C57B196CF2" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D68922DFDFDFC70CAE129A3" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius suwawa Telnov 2020, sp. nov." docType="treatment" docUuid="1B489847-69C1-47B5-90F7-AC360EC1E7D2" docUuidSource="ZooBank" docVersion="10" lastPageNumber="119" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="117" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CBCFFA724EB5FE3A8347E287F5E3A24C">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="ABDD6E82CCD346BC7717780F4946FE4A" type="personal">
<mods:role id="60FC96CEDDD0FF877434BA17325381C9">
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<mods:namePart id="A223132B466BF6ABEA40A65BAEAEF4FE">Telnov, Dmitry</mods:namePart>
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<mods:title id="21D4E6C4C9D110555B17FAE4CE262AAD">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="BBBB1B6BE75252ED2FAA056B4059416C">2020</mods:date>
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<mods:number id="63FA67BEB88A71603A3D676388251F85">2020-10-19</mods:number>
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<mods:number id="A7D00AF6D624F253D02C63CE966F84EC">721</mods:number>
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<mods:classification id="7D4F8BA1B41A17896C06BD9518E4D463">journal article</mods:classification>
<mods:identifier id="C16B94E81418D63ADE1F9E7CC7C8F2E4" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="081BF3D513E0CDBC438EA4227F53835A" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<mods:identifier id="6C6E98B0C6EDF5DCDF8457A26810C6A9" type="Zenodo-Dep">4122118</mods:identifier>
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<treatment id="1A3787F69D68922DFDFDFC70CAE129A3" ID-DOI="http://doi.org/10.5281/zenodo.4344367" ID-GBIF-Taxon="168648177" ID-Zenodo-Dep="4344367" LSID="urn:lsid:zoobank.org:act:1B489847-69C1-47B5-90F7-AC360EC1E7D2" httpUri="http://treatment.plazi.org/id/1A3787F69D68922DFDFDFC70CAE129A3" lastPageId="118" lastPageNumber="119" pageId="116" pageNumber="117">
<subSubSection id="DA84656B9D68922FFDFDFC70CB902B8B" box="[622,966,974,1001]" pageId="116" pageNumber="117" type="nomenclature">
<paragraph id="922136E09D68922FFDFDFC70CB902B8B" blockId="116.[370,1218,974,1074]" box="[622,966,974,1001]" pageId="116" pageNumber="117">
<heading id="C969818C9D68922FFDFDFC70CB902B8B" box="[622,966,974,1001]" centered="true" fontSize="11" level="2" pageId="116" pageNumber="117" reason="2">
<taxonomicName id="559E4D639D68922FFDFDFC70CB352B8B" ID-CoL="7X3XT" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[622,867,974,1001]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D68922FFDFDFC70CB352B8B" bold="true" box="[622,867,974,1001]" italics="true" pageId="116" pageNumber="117">Maechidius suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D68922FFCF8FC71CB902B8B" box="[875,966,975,1001]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D68922FFEE1FC4DCC182C53" pageId="116" pageNumber="117" type="description">
<paragraph id="922136E09D68922FFEE1FC4DCC942C6C" blockId="116.[370,1218,974,1074]" box="[370,1218,1011,1038]" pageId="116" pageNumber="117">
<uri id="E60F3AE29D68922FFEE1FC4DCC942C6C" box="[370,1218,1011,1038]" pageId="116" pageNumber="117">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D68922FFD0BFC4DCC942C6C" box="[664,1218,1011,1038]" pageId="116" pageNumber="117">1B489847-69C1-47B5-90F7-AC360EC1E7D2</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D68922FFE75FBA9CC182C53" blockId="116.[370,1218,974,1074]" box="[486,1102,1047,1074]" pageId="116" pageNumber="117">
<figureCitation id="0AA52A659D68922FFE75FBA9CA162C53" box="[486,576,1047,1073]" captionStart="Figs86–89" captionStartId="112.[189,241,1928,1954]" captionTargetBox="[282,1293,267,1862]" captionTargetId="figure-66@112.[278,1296,259,1877]" captionTargetPageId="112" captionText="Figs86–89. PapuanandWallaceanMaechidiusMacleay, 1819,habitus, dorsalview.86.M.speciosus(Frey, 1969) comb. nov., ♀ (BMNH). 87. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 88. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 89. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122167" httpUri="https://zenodo.org/record/4122167/files/figure.png" pageId="116" pageNumber="117">Figs 89</figureCitation>
,
<figureCitation id="0AA52A659D68922FFDDFFBA9CAE02C50" box="[588,694,1047,1074]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="116" pageNumber="117">184–185</figureCitation>
,
<figureCitation id="0AA52A659D68922FFD52FBA9CAA62C50" box="[705,752,1047,1074]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="116" pageNumber="117">258</figureCitation>
,
<figureCitation id="0AA52A659D68922FFD6FFBA9CB7D2C53" box="[764,811,1047,1073]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="116" pageNumber="117">349</figureCitation>
, 421, 525, 564, 726–729
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D68922FFF2EFBE7CCD62C89" pageId="116" pageNumber="117" type="diagnosis">
<paragraph id="922136E09D68922FFF2EFBE7C99F2C11" blockId="116.[189,457,1113,1139]" box="[189,457,1113,1139]" pageId="116" pageNumber="117">
<heading id="C969818C9D68922FFF2EFBE7C99F2C11" bold="true" box="[189,457,1113,1139]" fontSize="11" level="3" pageId="116" pageNumber="117" reason="3">
<emphasis id="A0EAEAF29D68922FFF2EFBE7C99F2C11" bold="true" box="[189,457,1113,1139]" pageId="116" pageNumber="117">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D68922FFF2EFB37CCD62C89" blockId="116.[189,1399,1161,1260]" pageId="116" pageNumber="117">
The new species is most similar to
<taxonomicName id="559E4D639D68922FFDCEFB34CB862CC1" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[605,976,1161,1187]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D68922FFDCEFB34CB542CC1" box="[605,770,1162,1187]" italics="true" pageId="116" pageNumber="117">M. peregrinus</emphasis>
<bibRefCitation id="F60F4B119D68922FFC99FB37CB862CC1" author="Lansberge J. W. van" box="[778,976,1161,1187]" pageId="116" pageNumber="117" pagination="131 - 137" refId="ref98256" refString="Lansberge J. W. van 1886. Description des quelques scarabaeides des Indes neerlandaises, appartenant au Musee du Leyde. Notes from the Leyden Museum 8: 131 - 137." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="559E4D639D68922FFC4EFB34CCD62CC1" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[989,1152,1161,1187]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D68922FFC4EFB34CCD62CC1" box="[989,1152,1161,1187]" italics="true" pageId="116" pageNumber="117">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D68922FFB1BFB34CCB32CC6" box="[1160,1253,1162,1188]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="559E4D639D68922FFB61FB34CD212CC1" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1266,1399,1161,1187]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D68922FFB61FB34CD212CC1" box="[1266,1399,1161,1187]" italics="true" pageId="116" pageNumber="117">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D68922FFF2EFB10C94F2CAA" box="[189,281,1198,1224]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="559E4D639D68922FFEC5FB10C98B2CA5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[342,477,1197,1223]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D68922FFEC5FB10C98B2CA5" box="[342,477,1197,1223]" italics="true" pageId="116" pageNumber="117">M. legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D68922FFE75FB10CA152CAA" box="[486,579,1198,1224]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
(all from
<collectingRegion id="505AF8029D68922FFD2DFB13CB7B2CAA" box="[702,813,1197,1224]" country="Indonesia" name="Sulawesi Utara" pageId="116" pageNumber="117">Sulawesi</collectingRegion>
). This new species readily differs from them and other congeners only in the shape of the male genital organs and the labroclypeus.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D68922FFF2EFAAACB6E2DE0" pageId="116" pageNumber="117" type="etymology">
<paragraph id="922136E09D68922FFF2EFAAAC9122D4C" blockId="116.[189,324,1300,1326]" box="[189,324,1300,1326]" pageId="116" pageNumber="117">
<heading id="C969818C9D68922FFF2EFAAAC9122D4C" bold="true" box="[189,324,1300,1326]" fontSize="11" level="3" pageId="116" pageNumber="117" reason="3">
<emphasis id="A0EAEAF29D68922FFF2EFAAAC9122D4C" bold="true" box="[189,324,1300,1326]" pageId="116" pageNumber="117">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D68922FFF2EFAFDCB6E2DE0" blockId="116.[189,1398,1347,1410]" pageId="116" pageNumber="117">
The name derives from Suwawa, one of the native languages spoken in the Bogani Nani Wartabone National park, northern
<collectingRegion id="505AF8029D68922FFE44FAD6CA132DE0" box="[471,581,1384,1410]" country="Indonesia" name="Sulawesi Utara" pageId="116" pageNumber="117">Sulawesi</collectingRegion>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D68922FFF2EFA14C9012F8B" pageId="116" pageNumber="117" type="materials_examined">
<paragraph id="922136E09D68922FFF2EFA14C93A2DA6" blockId="116.[189,364,1450,1476]" box="[189,364,1450,1476]" pageId="116" pageNumber="117">
<heading id="C969818C9D68922FFF2EFA14C93A2DA6" bold="true" box="[189,364,1450,1476]" fontSize="11" level="3" pageId="116" pageNumber="117" reason="3">
<emphasis id="A0EAEAF29D68922FFF2EFA14C93A2DA6" bold="true" box="[189,364,1450,1476]" pageId="116" pageNumber="117">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D68922FFF79FA64C90A2D96" blockId="116.[189,1399,1498,1633]" box="[234,348,1498,1524]" pageId="116" pageNumber="117">
<emphasis id="A0EAEAF29D68922FFF79FA64C90A2D96" bold="true" box="[234,348,1498,1524]" pageId="116" pageNumber="117">
<typeStatus id="4D2588429D68922FFF79FA64C90A2D96" box="[234,348,1498,1524]" pageId="116" pageNumber="117" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D68922FFF2EFA40C9932E03" blockId="116.[189,1399,1498,1633]" pageId="116" pageNumber="117">
<materialsCitation id="22F63CBD9D68922FFF2EFA40C9932E03" ID-GBIF-Occurrence="2898878333" collectingDate="1985-04" collectionCode="R, BMNH" collectorName="H. Barlow" country="Indonesia" elevation="1200" location="SULAWESI UTARA" pageId="116" pageNumber="117" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<collectingCountry id="EA8976709D68922FFF2EFA40C90D2E7A" box="[189,347,1534,1560]" name="Indonesia" pageId="116" pageNumber="117">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D68922FFEE0FA40C9DD2E7A" box="[371,395,1534,1560]" pageId="116" pageNumber="117">♂</specimenCount>
; “
<collectingCountry id="EA8976709D68922FFE31FA40CA152E7A" box="[418,579,1534,1560]" name="Indonesia" pageId="116" pageNumber="117">INDONESIA</collectingCountry>
:
<collectingRegion id="505AF8029D68922FFDCAFA40CB012E7A" box="[601,855,1534,1560]" country="Indonesia" name="Sulawesi Utara" pageId="116" pageNumber="117">SULAWESI UTARA</collectingRegion>
, Dumoga-Bone N.P.
<date id="E62010209D68922FFBDFFA40CC992E7B" box="[1100,1231,1534,1561]" pageId="116" pageNumber="117" value="1985-04">
<collectingDate id="F664E9C89D68922FFBDFFA40CC992E7B" box="[1100,1231,1534,1561]" pageId="116" pageNumber="117" value="1985-04">April 1985</collectingDate>
</date>
.//
<collectingMethod id="4BDF4EF79D68922FFB79FA40C97C2E5E" pageId="116" pageNumber="117">Rothamsted light trap</collectingMethod>
, site
<quantity id="55669B059D68922FFEF1F99CC9922E5E" box="[354,452,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="116" pageNumber="117" unit="m" value="1200.0">
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1,
<specimenCount id="8498FD699D68922FFEEEF99CC9922E5E" box="[381,452,1570,1596]" pageId="116" pageNumber="117" type="generic">200m</specimenCount>
</elevation>
</quantity>
.
<collectorName id="3F6B53369D68922FFE5EF99CCA172E5E" box="[461,577,1570,1596]" pageId="116" pageNumber="117">H.Barlow</collectorName>
//
<collectionCode id="F48FAE259D68922FFDCBF99CCA3B2E5E" box="[600,621,1570,1596]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="116" pageNumber="117">R</collectionCode>
.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985-10// 44.16 [light green label]”;
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.
</materialsCitation>
</paragraph>
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<typeStatus id="4D2588429D68922FFF79F937C9312EC1" box="[234,359,1673,1699]" pageId="116" pageNumber="117" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D68922FFEEBF937CA5D2EC1" box="[376,523,1672,1699]" pageId="116" pageNumber="117" type="generic">4 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D68922FFF2EF913C9012F8B" blockId="116.[189,1399,1672,2025]" pageId="116" pageNumber="117">
<materialsCitation id="22F63CBD9D68922FFF2EF913C9432F72" ID-GBIF-Occurrence="2898878488" collectingDate="1985-02" collectionCode="R, BMNH" collectorName="Tumpah Transect J. D. Holloway" country="Indonesia" location="Ent. Soc. Lond." municipality="Dumoga-Bone" pageId="116" pageNumber="117" specimenCount="492" stateProvince="Sulawesi Utara">
<collectingCountry id="EA8976709D68922FFF2EF913C90D2EA5" box="[189,347,1709,1735]" name="Indonesia" pageId="116" pageNumber="117">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D68922FFEEBF913C9FE2EA5" box="[376,424,1709,1735]" pageId="116" pageNumber="117" type="male">1 ♂</specimenCount>
; “
<collectingCountry id="EA8976709D68922FFE52F913CA342EA5" box="[449,610,1709,1735]" name="Indonesia" pageId="116" pageNumber="117">INDONESIA</collectingCountry>
:
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,
<collectingMunicipality id="7245AC9A9D68922FFC1FF913CC6F2EA5" box="[908,1081,1709,1735]" pageId="116" pageNumber="117">Dumoga-Bone</collectingMunicipality>
N.P.
<date id="E62010209D68922FFBEFF913CD7B2EAA" box="[1148,1325,1709,1736]" pageId="116" pageNumber="117" value="1985-02">
<collectingDate id="F664E9C89D68922FFBEFF913CD7B2EAA" box="[1148,1325,1709,1736]" pageId="116" pageNumber="117" value="1985-02">February 1985</collectingDate>
</date>
. // Site
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9,
<specimenCount id="8498FD699D68922FFF4EF96FC9782E8E" box="[221,302,1745,1772]" pageId="116" pageNumber="117" type="generic">492 m</specimenCount>
</quantity>
.
<collectorName id="3F6B53369D68922FFEAFF96FCAE12E89" box="[316,695,1745,1772]" pageId="116" pageNumber="117">Tumpah Transect J.D.Holloway</collectorName>
//
<collectionCode id="F48FAE259D68922FFD5CF96FCAB22E89" box="[719,740,1745,1771]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="116" pageNumber="117">R</collectionCode>
.
<location id="9741603B9D68922FFD7BF96FCBC52E89" LSID="urn:lsid:plazi:treatment:1A3787F69D68922DFDFDFC70CAE129A3:9741603B9D68922FFD7BF96FCBC52E89" box="[744,915,1745,1771]" country="Indonesia" municipality="Dumoga-Bone" name="Ent. Soc. Lond." pageId="116" pageNumber="117" stateProvince="Sulawesi Utara">Ent.Soc.Lond.</location>
PROJECT WALLACE B.M. 1985-10”;
<collectionCode id="F48FAE259D68922FFF2EF948C9432F72" box="[189,277,1782,1808]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="116" pageNumber="117">BMNH</collectionCode>
</materialsCitation>
•
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<specimenCount id="8498FD699D68922FFEA4F94BC93F2F72" box="[311,361,1781,1808]" pageId="116" pageNumber="117" type="male">1 ♂</specimenCount>
; “
<collectingCountry id="EA8976709D68922FFE17F94BCA732F6D" box="[388,549,1781,1807]" name="Indonesia" pageId="116" pageNumber="117">INDONESIA</collectingCountry>
:
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,
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N.P. //
<collectingMunicipality id="7245AC9A9D68922FFBC9F94BCCDE2F6D" box="[1114,1160,1781,1807]" pageId="116" pageNumber="117">Site</collectingMunicipality>
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8,
<specimenCount id="8498FD699D68922FFB26F94BCD502F72" box="[1205,1286,1781,1808]" pageId="116" pageNumber="117" type="generic">440 m</specimenCount>
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.
<collectorName id="3F6B53369D68922FFA85F94BC98F2F51" pageId="116" pageNumber="117">Tumpah Transect J.D.Holloway</collectorName>
<date id="E62010209D68922FFE7DF8A7CADA2F56" box="[494,652,1817,1844]" pageId="116" pageNumber="117" value="1985-02-12" valueMax="1985-02-13" valueMin="1985-02-12">
<collectingDate id="F664E9C89D68922FFE7DF8A7CADA2F56" box="[494,652,1817,1844]" pageId="116" pageNumber="117" value="1985-02-12" valueMax="1985-02-13" valueMin="1985-02-12">12-13.ii.1985</collectingDate>
</date>
//
<collectionCode id="F48FAE259D68922FFD37F8A7CAEF2F51" box="[676,697,1817,1843]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="116" pageNumber="117">R</collectionCode>
.
<location id="9741603B9D68922FFD2EF8A7CB3E2F51" LSID="urn:lsid:plazi:treatment:1A3787F69D68922DFDFDFC70CAE129A3:9741603B9D68922FFD2EF8A7CB3E2F51" box="[701,872,1817,1843]" country="Indonesia" county="Dumoga-Bone" municipality="Site" name="Ent. Soc. Lond." pageId="116" pageNumber="117" stateProvince="Sulawesi Utara">Ent.Soc.Lond.</location>
PROJECT WALLACE B.M. 1985-10”;
<collectionCode id="F48FAE259D68922FFF2EF880C9432F3A" box="[189,277,1854,1880]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="116" pageNumber="117">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D68922FFEB4F880C9192FC2" ID-GBIF-Occurrence="2898878341" collectingDate="1985-02-06" collectingDateMax="1985-02-07" collectingDateMin="1985-02-06" collectionCode="R, BMNH" collectorName="Tumpah Transect J. D. Holloway" country="Indonesia" location="Ent. Soc. Lond." municipality="Dumoga-Bone" pageId="116" pageNumber="117" specimenCount="300" stateProvince="Sulawesi Utara">
<specimenCount id="8498FD699D68922FFEB4F880C9042F3A" box="[295,338,1854,1880]" pageId="116" pageNumber="117" type="male">1 ♂</specimenCount>
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:
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,
<collectingMunicipality id="7245AC9A9D68922FFC8FF880CB9F2F3A" box="[796,969,1854,1880]" pageId="116" pageNumber="117">Dumoga-Bone</collectingMunicipality>
N.P.
<date id="E62010209D68922FFB91F880CC972F3A" box="[1026,1217,1853,1880]" pageId="116" pageNumber="117" value="1985-02-07">
<collectingDate id="F664E9C89D68922FFB91F880CC972F3A" box="[1026,1217,1853,1880]" pageId="116" pageNumber="117" value="1985-02-07">7 February 1985</collectingDate>
</date>
.// Site
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5,
<specimenCount id="8498FD699D68922FFABBF880CD242F3A" box="[1320,1394,1854,1880]" pageId="116" pageNumber="117" type="generic">300 m</specimenCount>
</quantity>
.
<collectorName id="3F6B53369D68922FFF2EF8DCCA602F1E" box="[189,566,1890,1916]" pageId="116" pageNumber="117">Tumpah Transect J.D.Holloway</collectorName>
<date id="E62010209D68922FFDD3F8DCCAFE2F1F" box="[576,680,1890,1917]" pageId="116" pageNumber="117" value="1985-02-06">
<collectingDate id="F664E9C89D68922FFDD3F8DCCAFE2F1F" box="[576,680,1890,1917]" pageId="116" pageNumber="117" value="1985-02-06">6.ii.1985</collectingDate>
</date>
[sic
<specimenCount id="8498FD699D68922FFD71F8DFCAB02F1E" box="[738,742,1889,1916]" pageId="116" pageNumber="117">♂</specimenCount>
] //
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.
<location id="9741603B9D68922FFC8CF8DCCB9C2F1E" LSID="urn:lsid:plazi:treatment:1A3787F69D68922DFDFDFC70CAE129A3:9741603B9D68922FFC8CF8DCCB9C2F1E" box="[799,970,1890,1916]" country="Indonesia" municipality="Dumoga-Bone" name="Ent. Soc. Lond." pageId="116" pageNumber="117" stateProvince="Sulawesi Utara">Ent.Soc.Lond.</location>
PROJECT WALLACE B.M. 1985- 10”;
<collectionCode id="F48FAE259D68922FFF64F838C9192FC2" box="[247,335,1926,1952]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="116" pageNumber="117">BMNH</collectionCode>
</materialsCitation>
•
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<specimenCount id="8498FD699D68922FFEFAF838C9C02FC2" box="[361,406,1926,1952]" pageId="116" pageNumber="117" type="female">1 ♀</specimenCount>
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, Dumoga-Bone N.P.
<date id="E62010209D68922FFBF3F838CD592FC3" box="[1120,1295,1926,1953]" pageId="116" pageNumber="117" value="1985-02">
<collectingDate id="F664E9C89D68922FFBF3F838CD592FC3" box="[1120,1295,1926,1953]" pageId="116" pageNumber="117" value="1985-02">February 1985</collectingDate>
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.// Site
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5,
<specimenCount id="8498FD699D68922FFF2EF814C95C2FA7" box="[189,266,1962,1989]" pageId="116" pageNumber="117" type="generic">300 m</specimenCount>
</elevation>
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.
<collectorName id="3F6B53369D68922FFE87F814CADF2FA6" box="[276,649,1962,1989]" pageId="116" pageNumber="117">Tumpah Transect J.D.Holloway</collectorName>
<date id="E62010209D68922FFD03F814CAAF2FA7" box="[656,761,1962,1989]" pageId="116" pageNumber="117" value="1985-02-06">
<collectingDate id="F664E9C89D68922FFD03F814CAAF2FA7" box="[656,761,1962,1989]" pageId="116" pageNumber="117" value="1985-02-06">6.ii.1985</collectingDate>
</date>
//
<collectionCode id="F48FAE259D68922FFC83F814CB732FA6" box="[784,805,1962,1988]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="116" pageNumber="117">R</collectionCode>
.
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PROJECT WALLACE B.M. 1985- 10”;
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.
</paragraph>
</subSubSection>
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<emphasis id="A0EAEAF29D69922EFF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="117" pageNumber="118">Description</emphasis>
</heading>
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<paragraph id="922136E09D69922EFF2EFE86CC502980" blockId="117.[189,1399,312,482]" pageId="117" pageNumber="118">
MEASUREMENTS (exposed abdominal ventrites not included). Male
<typeStatus id="4D2588429D69922EFC50FE86CC7B2930" box="[963,1069,312,338]" pageId="117" pageNumber="118" type="holotype">holotype</typeStatus>
, total body length
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. Head
<quantity id="55669B059D69922EFE97FEE2C9382914" box="[260,366,348,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="117" pageNumber="118" unit="mm" value="1.3">1.30 mm</quantity>
long, across eyes
<quantity id="55669B059D69922EFDD9FEE5CAE22914" box="[586,692,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="117" pageNumber="118" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D69922EFC10FEE5CBBB2914" box="[899,1005,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="117" pageNumber="118" unit="mm" value="1.5">1.50 mm</quantity>
long, maximum width
<quantity id="55669B059D69922EFA96FEE5CD252914" box="[1285,1395,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="117" pageNumber="118" unit="mm" value="2.6">2.60 mm</quantity>
. Elytral length
<quantity id="55669B059D69922EFEE3FE3EC9B729F8" box="[368,481,384,410]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="117" pageNumber="118" unit="mm" value="3.9">3.90 mm</quantity>
, maximum combined width
<quantity id="55669B059D69922EFCD5FEC1CBE129F8" box="[838,951,383,410]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="117" pageNumber="118" unit="mm" value="3.15">3.15 mm</quantity>
. Female
<typeStatus id="4D2588429D69922EFBB8FE3ECCC529F8" box="[1067,1171,384,410]" pageId="117" pageNumber="118" type="paratype">paratype</typeStatus>
, total body length
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. Head
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long, across eyes
<quantity id="55669B059D69922EFD57FE1DCB7829DC" box="[708,814,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="117" pageNumber="118" unit="mm" value="1.8">1.80 mm</quantity>
wide. Pronotum
<quantity id="55669B059D69922EFC6EFE1DCC3129DC" box="[1021,1127,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="117" pageNumber="118" unit="mm" value="1.8">1.80 mm</quantity>
long, maximum width
<quantity id="55669B059D69922EFF2EFE79C97F2980" box="[189,297,455,482]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="117" pageNumber="118" unit="mm" value="2.9">2.90 mm</quantity>
. Elytral length
<quantity id="55669B059D69922EFE4FFE79CA112980" box="[476,583,455,482]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="117" pageNumber="118" unit="mm" value="4.7">4.70 mm</quantity>
, maximum combined width
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.
</paragraph>
<paragraph id="922136E09D69922EFF2EFDB4CA9C2F69" blockId="117.[189,1399,521,1804]" pageId="117" pageNumber="118">
Dorsum uniformly brown with castaneous labroclypeus, mouthparts, legs and venter. Head transverse, trapezoid, slightly convex on vertex in lateral view, glossy dorsally and ventrally, with large not prominent compound eyes occupying more than half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D69922EFBDDFDEFCCE22A09" box="[1102,1204,593,619]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="117" pageNumber="118">Fig. 184</figureCitation>
) almost straight anteriorly, its lateral margins smooth, slightly sinuous in both dorsal and lateral views. Anterolateral angles shortly protruding anteriad, obtuse, bent up almost perpendicularly with regard to axis of frons (in lateral view). Female labroclypeus (
<figureCitation id="0AA52A659D69922EFDF2FD03CA932ABA" box="[609,709,701,728]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="117" pageNumber="118">Fig. 185</figureCitation>
) shallowly emarginate on anterior margin, with obtuse and not protruding anterolateral angles. Head punctures irregularly shaped, somewhat smaller in anterior, larger in posterior half. Intervening spaces glossy, smaller than punctures. Pubescence dirty yellow, rather long and erect but sparse, rising from each puncture. Setae somewhat hook-like curved apically, sparse, moderately long. Labroclypeus laterally and frontally with more delicate and shorter yellowish setae. Antenna 9-segmented. Scape large, widened on upper side in distal half, provided with few long erect setae on its posterior and distal margin. Antennomere 2 short and transverse. Club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum evenly broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D69922EFA91FBBECD3F2C79" box="[1282,1385,1024,1051]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="117" pageNumber="118">Fig. 258</figureCitation>
), nearly straight in lateral view. Long erect seta present between each of lateral crenulae and all along basal margin. Pronotal punctures circular to ovoid, deep and coarse. Intervening spaces glossy, smaller than punctures, in part wrinkled. Pronotal setae long, suberect, clavate. Hypomeron smooth, sparse punctured with circular punctures, with large emargination opposite to compound eye (to receive canthus separating compound eye) covered with long setae. Antennal pocket deep. Scutellar shield triangular, pointed apically. Elytra cylindrical, maximum combined width across median third, glossy and slightly convex dorsally, with rounded humeri and elevated omoplates. Vague tracks of two flat longitudinal carinae on each elytron. Sutural carinae shortly indicated near apices. Rather large obtuse hump on each elytron near apex. Disc and lateral sides of elytra densely punctured; punctures ovoid, less coarse and shallower than those of pronotum, arranged in irregular longitudinal rows (
<figureCitation id="0AA52A659D69922EFBB0FAD9CCDC2DE3" box="[1059,1162,1383,1409]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="117" pageNumber="118">Fig. 349</figureCitation>
). Intervening spaces glossy, generally smaller than punctures. An inconspicuous erect to suberect scale-like seta rises from anterior margin of each puncture, surpassing or not its length. Few longer stronger erect scale-like setae scattered over elytral disc and denser on lateral sides. Epipleuron with row or two (in broad anterior part) of suberect setae. Male and female pygidium with large shallow circular punctures (Fig. 525). Abdominal ventrites glossy, covered with small annular punctures, each provided with long suberect to erect seta. Legs long and slender, femora and tibiae covered with delicate yellowish setae. Outer margins of meso- and metatibiae densely denticulate. Two strong teeth on external margin of male protibia (Fig. 421). Male lower meso- and metatibial terminal spur strongly curved. Distal margin of metatibia in both sexes with projection over basal metatarsomere. Metatibial terminal spurs shorter than basal metatarsomere in male, nearly as long as it in female. Tarsal claws with pulvilli. Spiculum gastrale as in
<figureCitation id="0AA52A659D69922EFF2EF94FC9752F6E" box="[189,291,1777,1804]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="117" pageNumber="118">Fig. 564</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D69922EFDB1F94FCA902F69" box="[546,710,1777,1803]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="117" pageNumber="118">Figs 726–728</figureCitation>
.
</paragraph>
<paragraph id="922136E09D69922EFF2EF88DC9E62F2F" blockId="117.[189,432,1843,1869]" box="[189,432,1843,1869]" pageId="117" pageNumber="118">
<heading id="C969818C9D69922EFF2EF88DC9E62F2F" bold="true" box="[189,432,1843,1869]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
<emphasis id="A0EAEAF29D69922EFF2EF88DC9E62F2F" bold="true" box="[189,432,1843,1869]" pageId="117" pageNumber="118">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D69922EFF2EF8DDCA6E2F8A" blockId="117.[189,1399,1891,2024]" pageId="117" pageNumber="118">Female is generally larger; protibial spur slightly curved distally in male, straight in female; metatibial spurs somewhat longer in female than in male; lower metatibial spur curved in male, straight in female; female pulvilli narrower and shorter than in male; female lower meso- and metatibial terminal spur slender, slightly curved apically.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A922DFF2EFEB7CB512933" box="[189,775,265,337]" pageId="118" pageNumber="119" type="biology_ecology">
<paragraph id="922136E09D6A922DFF2EFEB7C9762941" blockId="118.[189,288,265,291]" box="[189,288,265,291]" pageId="118" pageNumber="119">
<heading id="C969818C9D6A922DFF2EFEB7C9762941" bold="true" box="[189,288,265,291]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="A0EAEAF29D6A922DFF2EFEB7C9762941" bold="true" box="[189,288,265,291]" pageId="118" pageNumber="119">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6A922DFF2EFE89CB512933" blockId="118.[189,775,311,337]" box="[189,775,311,337]" pageId="118" pageNumber="119">Occurs in lowland rainforests. Possibly nocturnal.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6A922DFF2EFEC6CAE129A3" box="[189,695,376,449]" pageId="118" pageNumber="119" type="distribution">
<paragraph id="922136E09D6A922DFF2EFEC6C90029F0" blockId="118.[189,342,376,402]" box="[189,342,376,402]" pageId="118" pageNumber="119">
<heading id="C969818C9D6A922DFF2EFEC6C90029F0" bold="true" box="[189,342,376,402]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="A0EAEAF29D6A922DFF2EFEC6C90029F0" bold="true" box="[189,342,376,402]" pageId="118" pageNumber="119">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6A922DFF2EFE19CAE129A3" blockId="118.[189,695,423,449]" box="[189,695,423,449]" pageId="118" pageNumber="119">
Hitherto only known from
<collectingRegion id="505AF8029D6A922DFE6AFE19CAE529A3" box="[505,691,423,449]" country="Indonesia" name="Sulawesi Utara" pageId="118" pageNumber="119">North Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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