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<document id="B5523034C9CD9BBA8D6246694855D5BC" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D5D9218FDF7F997CADD2BF2" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius leucopsar Telnov 2020, sp. nov." docType="treatment" docUuid="6D5298A8-F6E1-4361-B7CA-62C55B85C1DB" docUuidSource="ZooBank" docVersion="10" lastPageNumber="68" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="66" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7849AC58371D5B4035B0B35768837206">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="C7EC5F18890705E33AD25423CAA68ECB" type="personal">
<mods:role id="552433E933FBB596C87F2D7BA55590BE">
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<mods:namePart id="E156CC438753736C40600755BD7FB806">Telnov, Dmitry</mods:namePart>
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<mods:title id="E79627C8A28255C9415175FE9547CB83">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9043D1B37CE30F15A6F1F0CC501E96C6">
<mods:date id="3EE54229908A1BA5AB8CB38074F43EB6">2020</mods:date>
<mods:detail id="4EA677F1FA2640023837CF8FEBE89F89" type="pubDate">
<mods:number id="A445679AB2E93E312851DB9DBB884A6E">2020-10-19</mods:number>
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<mods:detail id="188817D48E222A546C656EACF92126A5" type="volume">
<mods:number id="9B4494E0D0C3D91F263B0342F56C0F85">721</mods:number>
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<mods:classification id="F099AD976BD716B0492CC6ABA1D89425">journal article</mods:classification>
<mods:identifier id="D09B6B906114DFD43BEE246B22E1D796" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="2A749C0E36B56E9D9691CC59A6A59677" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="2CDBDF291370472BA1058B0C6E62AAEA" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="F239A4AC86145AB3CA853DC4E78156DF" type="Zenodo-Dep">4122118</mods:identifier>
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<treatment id="1A3787F69D5D9218FDF7F997CADD2BF2" ID-DOI="http://doi.org/10.5281/zenodo.4344399" ID-GBIF-Taxon="168648168" ID-Zenodo-Dep="4344399" LSID="urn:lsid:zoobank.org:act:6D5298A8-F6E1-4361-B7CA-62C55B85C1DB" httpUri="http://treatment.plazi.org/id/1A3787F69D5D9218FDF7F997CADD2BF2" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
<subSubSection id="DA84656B9D5D921AFDF7F997CB862E26" box="[612,976,1577,1604]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="922136E09D5D921AFDF7F997CB862E26" blockId="65.[363,1225,1577,1673]" box="[612,976,1577,1604]" pageId="65" pageNumber="66">
<heading id="C969818C9D5D921AFDF7F997CB862E26" box="[612,976,1577,1604]" centered="true" fontSize="11" level="2" pageId="65" pageNumber="66" reason="2">
<taxonomicName id="559E4D639D5D921AFDF7F997CB382E26" ID-CoL="7X3WZ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[612,878,1577,1604]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="leucopsar" status="sp. nov.">
<emphasis id="A0EAEAF29D5D921AFDF7F997CB382E26" bold="true" box="[612,878,1577,1604]" italics="true" pageId="65" pageNumber="66">Maechidius leucopsar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5D921AFCE5F994CB862E26" box="[886,976,1578,1604]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5D921AFEF8F9F5CC182EEB" pageId="65" pageNumber="66" type="description">
<paragraph id="922136E09D5D921AFEF8F9F5CC9F2E04" blockId="65.[363,1225,1577,1673]" box="[363,1225,1611,1638]" pageId="65" pageNumber="66">
<uri id="E60F3AE29D5D921AFEF8F9F5CC9F2E04" box="[363,1225,1611,1638]" pageId="65" pageNumber="66">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D5D921AFD02F9F5CC9F2E04" box="[657,1225,1611,1638]" pageId="65" pageNumber="66">6D5298A8-F6E1-4361-B7CA-62C55B85C1DB</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D5D921AFE75F9D0CC182EEB" blockId="65.[363,1225,1577,1673]" box="[486,1102,1646,1673]" pageId="65" pageNumber="66">
<figureCitation id="0AA52A659D5D921AFE75F9D0CA162EEB" box="[486,576,1646,1673]" captionStart="Figs 50–53" captionStartId="62.[189,241,1927,1953]" captionTargetBox="[326,1248,265,1865]" captionTargetId="figure-65@62.[324,1254,249,1883]" captionTargetPageId="62" captionText="Figs 50–53. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 50. M. lapsus sp. nov., holotype, ♂ (BMNH). 51. M. legalovi sp. nov., holotype, ♂ (BMNH). 52. M. leucopsar sp. nov., holotype, ♂ (IECA). 53. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122148" httpUri="https://zenodo.org/record/4122148/files/figure.png" pageId="65" pageNumber="66">Figs 52</figureCitation>
,
<figureCitation id="0AA52A659D5D921AFDDFF9D0CA2D2EEA" box="[588,635,1646,1672]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="65" pageNumber="66">132</figureCitation>
,
<figureCitation id="0AA52A659D5D921AFD14F9D0CAE02EEB" box="[647,694,1646,1673]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="65" pageNumber="66">225</figureCitation>
,
<figureCitation id="0AA52A659D5D921AFD52F9D0CAA62EEA" box="[705,752,1646,1672]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="65" pageNumber="66">316</figureCitation>
,
<figureCitation id="0AA52A659D5D921AFD6FF9D0CB7D2EEB" box="[764,811,1646,1673]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="65" pageNumber="66">357</figureCitation>
, 389, 485, 550, 643–645
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5D921AFF2EF910C9D02F8A" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="922136E09D5D921AFF2EF910C99F2EAA" blockId="65.[189,457,1710,1736]" box="[189,457,1710,1736]" pageId="65" pageNumber="66">
<heading id="C969818C9D5D921AFF2EF910C99F2EAA" bold="true" box="[189,457,1710,1736]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="A0EAEAF29D5D921AFF2EF910C99F2EAA" bold="true" box="[189,457,1710,1736]" pageId="65" pageNumber="66">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5D921AFF2EF962C9D02F8A" blockId="65.[189,1399,1756,2024]" pageId="65" pageNumber="66">
This new species is undoubtedly very close to
<taxonomicName id="559E4D639D5D921AFC9AF962CBA82E94" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[777,1022,1756,1782]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D5D921AFC9AF962CBA82E94" box="[777,1022,1756,1782]" italics="true" pageId="65" pageNumber="66">Maechidius suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5D921AFB99F963CC3C2E95" box="[1034,1130,1757,1783]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
from
<collectingRegion id="505AF8029D5D921AFB28F962CD202E94" box="[1211,1398,1756,1782]" country="Indonesia" name="Sulawesi Utara" pageId="65" pageNumber="66">North Sulawesi</collectingRegion>
(see description of this species below) and other congeners. It differs primarily in the male genitalia (cf.
<figureCitation id="0AA52A659D5D921AFF7CF89FC9C02F5E" box="[239,406,1825,1852]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="65" pageNumber="66">Figs 643–645</figureCitation>
,
<figureCitation id="0AA52A659D5D921AFE36F89FCA472F5E" box="[421,529,1825,1852]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="65" pageNumber="66">726–729</figureCitation>
), the shape of the male metatibial terminal spurs, the absence of glossy elytral carinae (two flat glossy glabrous carinae on each elytron in
<taxonomicName id="559E4D639D5D921AFC5FF8FACC052F3F" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[972,1107,1860,1885]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D5D921AFC5FF8FACC052F3F" box="[972,1107,1860,1885]" italics="true" pageId="65" pageNumber="66">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5D921AFBC9F8FACCEE2F3C" box="[1114,1208,1860,1886]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
), a less coarsely punctured pronotum (punctures of pronotal disc smaller, ovoid, deeper in
<taxonomicName id="559E4D639D5D921AFBB2F8D9CCEA2FE2" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1057,1212,1894,1920]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="leucopsar" status="sp. nov.">
<emphasis id="A0EAEAF29D5D921AFBB2F8D9CCEA2FE2" box="[1057,1212,1894,1920]" italics="true" pageId="65" pageNumber="66">M. leucopsar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5D921AFB57F8D8CD482FE2" box="[1220,1310,1894,1920]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
than in any
<collectingRegion id="505AF8029D5D921AFF63F836C90C2FC0" box="[240,346,1928,1954]" country="Indonesia" name="Sulawesi Utara" pageId="65" pageNumber="66">Sulawesi</collectingRegion>
species), the punctures of the elytral disc are circular and less regular (elytral punctures ovoid, substriate in
<taxonomicName id="559E4D639D5D921AFE3BF812CA662FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[424,560,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D5D921AFE3BF812CA662FA7" box="[424,560,1964,1989]" italics="true" pageId="65" pageNumber="66">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5D921AFDABF815CAC12FA7" box="[568,663,1963,1989]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
), and the comparatively deeper emarginate anterior margin of the labroclypeus.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5E9219FF2EFEB7CA3B29FB" pageId="66" pageNumber="67" type="etymology">
<paragraph id="922136E09D5E9219FF2EFEB7C9122941" blockId="66.[189,324,265,291]" box="[189,324,265,291]" pageId="66" pageNumber="67">
<heading id="C969818C9D5E9219FF2EFEB7C9122941" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="A0EAEAF29D5E9219FF2EFEB7C9122941" bold="true" box="[189,324,265,291]" pageId="66" pageNumber="67">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5E9219FF2EFE86CA3B29FB" blockId="66.[189,1398,312,410]" pageId="66" pageNumber="67">
Named after
<emphasis id="A0EAEAF29D5E9219FECAFE86C9832933" box="[345,469,312,337]" italics="true" pageId="66" pageNumber="67">Leucopsar</emphasis>
Stresemann, 1912, the monotypical sturnid genus of the famous and critically endangered
<collectingRegion id="505AF8029D5E9219FEDCFEE5C9D62917" box="[335,384,347,373]" country="Indonesia" name="Bali" pageId="66" pageNumber="67">Bali</collectingRegion>
endemic
<emphasis id="A0EAEAF29D5E9219FE6BFEE2CAAA2917" box="[504,764,347,373]" italics="true" pageId="66" pageNumber="67">Leucopsar rothschildi</emphasis>
Stresemann, 1912, widely known as the
<collectingRegion id="505AF8029D5E9219FB60FEE5CD722917" box="[1267,1316,347,373]" country="Indonesia" name="Bali" pageId="66" pageNumber="67">Bali</collectingRegion>
myna, Rothschild’s mynah, or
<collectingRegion id="505AF8029D5E9219FE46FEC1CA5029FB" box="[469,518,383,409]" country="Indonesia" name="Bali" pageId="66" pageNumber="67">Bali</collectingRegion>
starling.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5E9219FF2EFE7CCB1D2ADB" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="922136E09D5E9219FF2EFE7CC93A29B9" blockId="66.[189,364,449,476]" box="[189,364,449,476]" pageId="66" pageNumber="67">
<heading id="C969818C9D5E9219FF2EFE7CC93A29B9" bold="true" box="[189,364,449,476]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="A0EAEAF29D5E9219FF2EFE7CC93A29B9" bold="true" box="[189,364,449,476]" pageId="66" pageNumber="67">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D5E9219FF79FE4FCB582A31" ID-GBIF-Occurrence="2898878567" collectingDate="2004-11" collectionCode="IECA" country="Indonesia" elevation="1200" location="Isl. Bedugul Region" pageId="66" pageNumber="67" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<paragraph id="922136E09D5E9219FF79FE4FC90A2A69" blockId="66.[189,1399,497,597]" box="[234,348,497,523]" pageId="66" pageNumber="67">
<emphasis id="A0EAEAF29D5E9219FF79FE4FC90A2A69" bold="true" box="[234,348,497,523]" pageId="66" pageNumber="67">
<typeStatus id="4D2588429D5E9219FF79FE4FC90A2A69" box="[234,348,497,523]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D5E9219FF2EFDABCB582A31" blockId="66.[189,1399,497,597]" pageId="66" pageNumber="67">
<collectingCountry id="EA8976709D5E9219FF2EFDABC90D2A4D" box="[189,347,533,559]" name="Indonesia" pageId="66" pageNumber="67">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D5E9219FEE2FDABC9DF2A4D" box="[369,393,533,559]" pageId="66" pageNumber="67" type="male">♂</specimenCount>
; “Indonesia,
<collectingRegion id="505AF8029D5E9219FDB2FDABCA042A4D" box="[545,594,533,559]" country="Indonesia" name="Bali" pageId="66" pageNumber="67">Bali</collectingRegion>
<location id="9741603B9D5E9219FDC4FDABCB152A4D" LSID="urn:lsid:plazi:treatment:1A3787F69D5D9218FDF7F997CADD2BF2:9741603B9D5E9219FDC4FDABCB152A4D" box="[599,835,533,559]" country="Indonesia" name="Isl. Bedugul Region" pageId="66" pageNumber="67" stateProvince="Bali">Isl. Bedugul Region</location>
, Tamblingan Lakes N.R.
<date id="E62010209D5E9219FBF5FDABCC9C2A4D" box="[1126,1226,533,559]" pageId="66" pageNumber="67" value="2004-11">
<collectingDate id="F664E9C89D5E9219FBF5FDABCC9C2A4D" box="[1126,1226,533,559]" pageId="66" pageNumber="67" value="2004-11">XI.2004</collectingDate>
</date>
,
<quantity id="55669B059D5E9219FB47FDABCD7A2A4D" box="[1236,1324,533,559]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="66" pageNumber="67" unit="m" value="1200.0">
<elevation id="19B3D1D39D5E9219FB47FDABCD7A2A4D" box="[1236,1324,533,559]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="66" pageNumber="67" unit="m" value="1200.0">1200 m</elevation>
</quantity>
// coll.
<collectionCode id="F48FAE259D5E9219FF2EFD87C9562A31" box="[189,256,569,595]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="66" pageNumber="67" type="Other institutional type">IECA</collectionCode>
České Budějovice, Czech Republic”;
<collectionCode id="F48FAE259D5E9219FD51FD87CB5C2A31" box="[706,778,569,595]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="66" pageNumber="67" type="Other institutional type">IECA</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D5E9219FF79FDC5C90A2AF7" blockId="66.[189,843,635,697]" box="[234,348,635,661]" pageId="66" pageNumber="67">
<emphasis id="A0EAEAF29D5E9219FF79FDC5C90A2AF7" bold="true" box="[234,348,635,661]" pageId="66" pageNumber="67">
<typeStatus id="4D2588429D5E9219FF79FDC5C90A2AF7" box="[234,348,635,661]" pageId="66" pageNumber="67" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D5E9219FF2EFD21CB1D2ADB" blockId="66.[189,843,635,697]" box="[189,843,670,697]" pageId="66" pageNumber="67">
<materialsCitation id="22F63CBD9D5E9219FF2EFD21CB102ADB" ID-GBIF-Occurrence="2898878389" box="[189,838,670,697]" collectingDate="2004-11" collectionCode="IECA" country="Indonesia" elevation="1200" location="INDONESIA" pageId="66" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Bali" typeStatus="paratype">
<collectingCountry id="EA8976709D5E9219FF2EFD21C90D2ADB" box="[189,347,671,697]" name="Indonesia" pageId="66" pageNumber="67">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D5E9219FEE7FD21C9F42ADB" box="[372,418,671,697]" pageId="66" pageNumber="67" type="male">1 ♂</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D5E9219FD6DFD21CB102ADB" box="[766,838,671,697]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="66" pageNumber="67" type="Other institutional type">IECA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5E9218FF2EFD5ECA9B2AEC" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="description">
<paragraph id="922136E09D5E9219FF2EFD5EC9182A98" blockId="66.[189,334,736,762]" box="[189,334,736,762]" pageId="66" pageNumber="67">
<heading id="C969818C9D5E9219FF2EFD5EC9182A98" bold="true" box="[189,334,736,762]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="A0EAEAF29D5E9219FF2EFD5EC9182A98" bold="true" box="[189,334,736,762]" pageId="66" pageNumber="67">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5E9219FF2EFCAEC97B2B10" blockId="66.[189,1399,784,882]" pageId="66" pageNumber="67">
MEASUREMENTS.
<typeStatus id="4D2588429D5E9219FEEFFCAEC9BA2B48" box="[380,492,784,810]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D5E9219FD54FCAECB652B48" box="[711,819,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="66" pageNumber="67" unit="mm" value="6.6">6.60 mm</quantity>
. Head
<quantity id="55669B059D5E9219FC16FCAECBB82B48" box="[901,1006,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="66" pageNumber="67" unit="mm" value="1.3">1.30 mm</quantity>
long, across eyes
<quantity id="55669B059D5E9219FB55FCAECD792B48" box="[1222,1327,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="66" pageNumber="67" unit="mm" value="1.47">1.47 mm</quantity>
wide. Pronotum
<quantity id="55669B059D5E9219FEA4FC8AC9C92B2C" box="[311,415,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="66" pageNumber="67" unit="mm" value="1.5">1.50 mm</quantity>
long, maximum width
<quantity id="55669B059D5E9219FD3EFC8ACB412B2C" box="[685,791,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="66" pageNumber="67" unit="mm" value="2.5">2.50 mm</quantity>
. Elytral length
<quantity id="55669B059D5E9219FC54FC8ACC642B2C" box="[967,1074,820,846]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="66" pageNumber="67" unit="mm" value="3.8">3.80 mm</quantity>
, maximum combined width
<quantity id="55669B059D5E9219FF2EFCE6C97F2B10" box="[189,297,856,882]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="66" pageNumber="67" unit="mm" value="3.26">3.26 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D5E9218FF2EFC24CBB429D2" blockId="66.[189,1399,922,2025]" lastBlockId="67.[189,1399,265,432]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
Dorsum uniformly black with reddish brown labroclypeus, mouthparts, antennae, legs and all of venter. Head transverse, glossy to subopaque dorsally and ventrally, somewhat convex between eyes and vaguely impressed on labroclypeus anterior to each compound eye. Compound eye large, occupying about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D5E9219FD56FBBBCB7F2C42" box="[709,809,1029,1056]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="66" pageNumber="67">Fig. 132</figureCitation>
) very shallowly broadly emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views, anterolateral angles weakly protruding, obtuse in dorsal view, bent up at ~80–90° to frons in lateral view. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head dorsal punctures circular to ovoid, deep, smaller on labroclypeus than on frons; inner margin of some punctures with delicate membrane. Intervening spaces glossy to very delicately microreticulate, glabrous, variably large. Suberect thickened moderately long scale-like seta rises from anterior margin of each puncture, surpassing length of corresponding puncture. Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 trapezoid, transverse. Pronotum glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate, anterolateral angles protruding anteriad. Vague dorsal impression beyond anterolateral angle on either side. Basal and lateral margins broadly rounded. Crenulae of lateral margin delicate, a long suberect seta present between every two crenulae (
<figureCitation id="0AA52A659D5E9219FE2AFA66CA752D91" box="[441,547,1496,1523]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="66" pageNumber="67">Fig. 225</figureCitation>
). Lateral margin of pronotum nearly straight in lateral view. Pronotal punctures ovoid, very deep, variably large; inner margin of some punctures with delicate membrane. Intervening spaces glossy and glabrous, in part wrinkled, generally smaller than punctures except near lateral margins. Setae shorter than those on frons. Lateral and basal margins, antero- and posterolateral angles partly covered with dense microscopical velvety pubescence (
<figureCitation id="0AA52A659D5E9219FC62F9D9CC0C2EE0" box="[1009,1114,1639,1666]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="66" pageNumber="67">Fig. 357</figureCitation>
). Hypomeron shallowly emarginate and very long setose on anterior margin which is flange-like. Antennal pocket deep. Median anterior process of prosternum long brushy setose, short, moderately raised. Scutellar shield rounded apically. Elytra slightly widened in posterior half, maximum width across midlength, glossy dorsally, with distinct humeri. Punctures of elytral disc irregular in shape and size, generally much shallower than those on pronotal disc; on periphery partly or completely encircled with dense microscopical velvety pubescence (
<figureCitation id="0AA52A659D5E9219FEC4F881C9942F38" box="[343,450,1855,1882]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="66" pageNumber="67">Fig. 316</figureCitation>
). Setae inconspicuous, in part minute, in part moderately large and surpassing length of corresponding punctures. Epipleuron all along with very short appressed setae. Male pygidium slightly convex dorsally, circularly punctate, subopaque microreticulate on intervening spaces, spaces generally smaller than punctures, with moderately long suberect seta that rises from each puncture (
<figureCitation id="0AA52A659D5E9219FF55F870C9672F8B" box="[198,305,1998,2025]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="66" pageNumber="67">Fig. 485</figureCitation>
). Venter covered with sparse small shallow punctures, each provided with very short seta. Legs long and slender. Male protibia nearly straight on external margin, delicately crenulate in basal third, with almost complete dorsal fovea. Two acute rather large distal teeth on male protibia, distal longer than basal one (Fig. 389). Male protibial terminal spur large, curved. Male metatibial terminal spurs obtuse, the lower stronger curved than upper one. Tarsal claws with large pulvilli (in male only?). Spiculum gastrale as in
<figureCitation id="0AA52A659D5F9218FE46FE2BCA6D29D2" box="[469,571,405,432]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="67" pageNumber="68">Fig. 550</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D5F9218FCA9FE2BCB8829D2" box="[826,990,405,432]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="67" pageNumber="68">Figs 643–645</figureCitation>
.
</paragraph>
<paragraph id="922136E09D5F9218FF2EFE68C9E62992" blockId="67.[189,432,470,496]" box="[189,432,470,496]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFE68C9E62992" bold="true" box="[189,432,470,496]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFE68C9E62992" bold="true" box="[189,432,470,496]" pageId="67" pageNumber="68">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFDBAC9FD2A7C" blockId="67.[189,427,516,542]" box="[189,427,516,542]" pageId="67" pageNumber="68">Female is unknown.</paragraph>
<paragraph id="922136E09D5F9218FF2EFDFBC9152A3D" blockId="67.[189,323,581,607]" box="[189,323,581,607]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFDFBC9152A3D" bold="true" box="[189,323,581,607]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFDFBC9152A3D" bold="true" box="[189,323,581,607]" pageId="67" pageNumber="68">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFDCACA9B2AEC" blockId="67.[189,717,628,654]" box="[189,717,628,654]" pageId="67" pageNumber="68">
<typeStatus id="4D2588429D5F9218FF2EFDCAC9722AEC" box="[189,292,628,654]" pageId="67" pageNumber="68" type="paratype">Paratype</typeStatus>
elytral disc less coarsely punctured.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F9218FF2EFD0BCA292A9C" box="[189,639,693,766]" pageId="67" pageNumber="68" type="biology_ecology">
<paragraph id="922136E09D5F9218FF2EFD0BC9762AAD" blockId="67.[189,288,693,719]" box="[189,288,693,719]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFD0BC9762AAD" bold="true" box="[189,288,693,719]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFD0BC9762AAD" bold="true" box="[189,288,693,719]" pageId="67" pageNumber="68">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFD5ACA292A9C" blockId="67.[189,639,739,766]" box="[189,639,739,766]" pageId="67" pageNumber="68">
Occurs at an altitude of about
<quantity id="55669B059D5F9218FD8CFD5ACA2D2A9C" box="[543,635,740,766]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="67" pageNumber="68" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F9218FF2EFC9ACADD2BF2" pageId="67" pageNumber="68" type="distribution">
<paragraph id="922136E09D5F9218FF2EFC9AC9002B5C" blockId="67.[189,342,804,830]" box="[189,342,804,830]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFC9AC9002B5C" bold="true" box="[189,342,804,830]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFC9AC9002B5C" bold="true" box="[189,342,804,830]" pageId="67" pageNumber="68">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFCEDCADD2BF2" blockId="67.[189,1399,851,912]" pageId="67" pageNumber="68">
Hitherto only known from
<collectingRegion id="505AF8029D5F9218FD9EFCEDCA172B0F" box="[525,577,851,877]" country="Indonesia" name="Bali" pageId="67" pageNumber="68">Bali</collectingRegion>
, the westernmost of the
<collectingRegion id="505AF8029D5F9218FCE5FCEDCC2A2B0F" box="[886,1148,851,877]" country="Indonesia" name="Nusa Tenggara Timur" pageId="67" pageNumber="68">Lesser Sunda Islands</collectingRegion>
. This is hitherto the westernmost record of the
<taxonomicName id="559E4D639D5F9218FE65FCC8CADE2BF2" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[502,648,886,912]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="5" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Maechidiini">Maechidiini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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