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<document id="62504E020E36D32EC03DED8F8223EDDA" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D5C921AFDFCFEB6CA9E2D8E" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius legalovi Telnov 2020, sp. nov." docType="treatment" docUuid="ABC509EC-B963-4A6D-8CC7-804615AB794D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="66" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="65" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8E4809F2CC0DCED761D1B1D868920470" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85C94780987A3A5A2D8D850581BD672D">
<mods:title id="62ABD8BF214AE714BC3A25D3A0771598">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="8FD93F0429F93A0537489F19F18F6D65" type="personal">
<mods:role id="189180D0D8705E4FEDC7BA87D4359A96">
<mods:roleTerm id="DB11BE02F7D3F105F1E20CF0034AF251">Author</mods:roleTerm>
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<mods:namePart id="FDA6F57C905F004CC476C198064C61C8">Telnov, Dmitry</mods:namePart>
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<mods:titleInfo id="3F339F9FECE088EFCE751D888F477CEB">
<mods:title id="27B2A3FA37DB5F644234D8433916A19F">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="2BF59926E9686FA71E4830130F25EAC4">
<mods:date id="BDD3B4EBC6667C28549657AEDE8F12A4">2020</mods:date>
<mods:detail id="AFE4700A724952D1796B172D00EFE41A" type="pubDate">
<mods:number id="D13E8B2DB599BF85C22E2EF0C28BA256">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="F93A0BFBD1ACF7E73F74D71EE0F1E6A3" type="volume">
<mods:number id="52BF8BF16546E4EE509D4FC4052FCFDC">721</mods:number>
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<mods:classification id="EF3F94D1E7D2C421CCA714A3CE5AF160">journal article</mods:classification>
<mods:identifier id="2F245521C02CE6D2F31E8E2EB5936D6F" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="EE9CE870F1B5F0956B40EAC06ACE9108" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="FCFC660D6A76E8BF9E4331733821F3E9" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="D89AE376E745EE95ECA25E55902E11B7" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="0D8F320768A43611B52C6173CCA38425" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
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<treatment id="1A3787F69D5C921AFDFCFEB6CA9E2D8E" ID-DOI="http://doi.org/10.5281/zenodo.4344361" ID-GBIF-Taxon="168648169" ID-Zenodo-Dep="4344361" LSID="urn:lsid:zoobank.org:act:ABC509EC-B963-4A6D-8CC7-804615AB794D" httpUri="http://treatment.plazi.org/id/1A3787F69D5C921AFDFCFEB6CA9E2D8E" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<subSubSection id="DA84656B9D5C921BFDFCFEB6CB932941" box="[623,965,264,291]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="922136E09D5C921BFDFCFEB6CB932941" blockId="64.[358,1229,264,363]" box="[623,965,264,291]" pageId="64" pageNumber="65">
<heading id="C969818C9D5C921BFDFCFEB6CB932941" box="[623,965,264,291]" centered="true" fontSize="11" level="2" pageId="64" pageNumber="65" reason="2">
<taxonomicName id="559E4D639D5C921BFDFCFEB6CB352941" ID-CoL="7X3WY" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[623,867,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D5C921BFDFCFEB6CB352941" bold="true" box="[623,867,264,291]" italics="true" pageId="64" pageNumber="65">Maechidius legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5C921BFCF9FEB7CB932941" box="[874,965,265,291]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5C921BFEF5FE93CC662909" pageId="64" pageNumber="65" type="description">
<paragraph id="922136E09D5C921BFEF5FE93CC9B292A" blockId="64.[358,1229,264,363]" box="[358,1229,301,328]" pageId="64" pageNumber="65">
<uri id="E60F3AE29D5C921BFEF5FE93CC9B292A" box="[358,1229,301,328]" pageId="64" pageNumber="65">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D5C921BFD10FE93CC9B292A" box="[643,1229,301,328]" pageId="64" pageNumber="65">ABC509EC-B963-4A6D-8CC7-804615AB794D</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D5C921BFD90FEEFCC662909" blockId="64.[358,1229,264,363]" box="[515,1072,336,363]" pageId="64" pageNumber="65">
<figureCitation id="0AA52A659D5C921BFD90FEEFCA0B2909" box="[515,605,336,363]" captionStart="Figs 50–53" captionStartId="62.[189,241,1927,1953]" captionTargetBox="[326,1248,265,1865]" captionTargetId="figure-65@62.[324,1254,249,1883]" captionTargetPageId="62" captionText="Figs 50–53. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 50. M. lapsus sp. nov., holotype, ♂ (BMNH). 51. M. legalovi sp. nov., holotype, ♂ (BMNH). 52. M. leucopsar sp. nov., holotype, ♂ (IECA). 53. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122148" httpUri="https://zenodo.org/record/4122148/files/figure.png" pageId="64" pageNumber="65">Figs 51</figureCitation>
,
<figureCitation id="0AA52A659D5C921BFDFAFEEFCACE2909" box="[617,664,337,363]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="64" pageNumber="65">136</figureCitation>
,
<figureCitation id="0AA52A659D5C921BFD37FEEFCA852909" box="[676,723,337,363]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="64" pageNumber="65">224</figureCitation>
,
<figureCitation id="0AA52A659D5C921BFD4CFEEECB582909" box="[735,782,336,363]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="64" pageNumber="65">315</figureCitation>
, 388, 483, 549, 640–642
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5C921BFF2EFE2DCCEE2A2A" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="922136E09D5C921BFF2EFE2DC99F29CF" blockId="64.[189,457,403,429]" box="[189,457,403,429]" pageId="64" pageNumber="65">
<heading id="C969818C9D5C921BFF2EFE2DC99F29CF" bold="true" box="[189,457,403,429]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A0EAEAF29D5C921BFF2EFE2DC99F29CF" bold="true" box="[189,457,403,429]" pageId="64" pageNumber="65">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5C921BFF2EFE7CCCEE2A2A" blockId="64.[189,1399,450,584]" pageId="64" pageNumber="65">
This new species is most similar to
<taxonomicName id="559E4D639D5C921BFDF3FE7DCB8529BE" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[608,979,450,476]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D5C921BFDF3FE7DCB5029BE" box="[608,774,451,476]" italics="true" pageId="64" pageNumber="65">M. peregrinus</emphasis>
<bibRefCitation id="F60F4B119D5C921BFC9EFE7CCB8529BE" author="Lansberge J. W. van" box="[781,979,450,476]" pageId="64" pageNumber="65" pagination="131 - 137" refId="ref98256" refString="Lansberge J. W. van 1886. Description des quelques scarabaeides des Indes neerlandaises, appartenant au Musee du Leyde. Notes from the Leyden Museum 8: 131 - 137." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="559E4D639D5C921BFC4CFE7DCCD429BE" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[991,1154,450,476]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D5C921BFC4CFE7DCCD429BE" box="[991,1154,450,476]" italics="true" pageId="64" pageNumber="65">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5C921BFB1AFE7DCCB029BF" box="[1161,1254,451,477]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="559E4D639D5C921BFB61FE7DCD2129BE" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1266,1399,450,476]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D5C921BFB61FE7DCD2129BE" box="[1266,1399,450,476]" italics="true" pageId="64" pageNumber="65">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5C921BFF2EFE58C94F2A62" box="[189,281,486,512]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="559E4D639D5C921BFEC0FE59C98D2A62" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[339,475,487,512]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D5C921BFEC0FE59C98D2A62" box="[339,475,487,512]" italics="true" pageId="64" pageNumber="65">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5C921BFE70FE58CA682A62" box="[483,574,486,512]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
(all from
<collectingRegion id="505AF8029D5C921BFD24FE58CB702A63" box="[695,806,486,513]" country="Indonesia" name="Sulawesi Utara" pageId="64" pageNumber="65">Sulawesi</collectingRegion>
). It readily differs from them and other congeners in the widened and flattened male basal metatarsomere, the shape of the male genital organs and the labroclypeus, which is entire and without protruding anterolateral angles in both sexes.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5C921BFF2EFDCECAF12ABC" pageId="64" pageNumber="65" type="etymology">
<paragraph id="922136E09D5C921BFF2EFDCEC9122AE8" blockId="64.[189,324,624,650]" box="[189,324,624,650]" pageId="64" pageNumber="65">
<heading id="C969818C9D5C921BFF2EFDCEC9122AE8" bold="true" box="[189,324,624,650]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A0EAEAF29D5C921BFF2EFDCEC9122AE8" bold="true" box="[189,324,624,650]" pageId="64" pageNumber="65">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5C921BFF2EFD21CAF12ABC" blockId="64.[189,1398,671,734]" pageId="64" pageNumber="65">
Patronymic. This species is named after the famous expert of Curculionoidea and respected colleague Andrei A. Legalov (
<collectingRegion id="505AF8029D5C921BFE34FD7DCA6D2ABF" box="[423,571,707,733]" country="Russia" name="Novosibirsk" pageId="64" pageNumber="65">Novosibirsk</collectingRegion>
,
<collectingCountry id="EA8976709D5C921BFDD4FD7DCACC2ABC" box="[583,666,707,734]" name="Russia" pageId="64" pageNumber="65">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5C921BFF2EFCBBCB782C0A" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="922136E09D5C921BFF2EFCBBC93A2B7D" blockId="64.[189,364,773,799]" box="[189,364,773,799]" pageId="64" pageNumber="65">
<heading id="C969818C9D5C921BFF2EFCBBC93A2B7D" bold="true" box="[189,364,773,799]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A0EAEAF29D5C921BFF2EFCBBC93A2B7D" bold="true" box="[189,364,773,799]" pageId="64" pageNumber="65">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5C921BFF79FC8AC90A2B2C" blockId="64.[189,1399,820,918]" box="[234,348,820,846]" pageId="64" pageNumber="65">
<emphasis id="A0EAEAF29D5C921BFF79FC8AC90A2B2C" bold="true" box="[234,348,820,846]" pageId="64" pageNumber="65">
<typeStatus id="4D2588429D5C921BFF79FC8AC90A2B2C" box="[234,348,820,846]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D5C921BFF2EFCE6CC272BF4" blockId="64.[189,1399,820,918]" pageId="64" pageNumber="65">
<materialsCitation id="22F63CBD9D5C921BFF2EFCE6CC272BF4" ID-GBIF-Occurrence="2898878500" collectingDate="1980-01-27" collectingDateMax="1980-04-20" collectingDateMin="1980-01-27" collectionCode="BMNH" collectorName="Nr. Morowali &amp; S. L. Sutton C. J. Rees" country="Indonesia" elevation="30" location="Vert. Series" municipality="Ranu River Area" pageId="64" pageNumber="65" specimenCount="1" specimenCount-male="1" stateProvince="SULAWESI TENGAH" typeStatus="holotype">
<collectingCountry id="EA8976709D5C921BFF2EFCE6C90D2B10" box="[189,347,856,882]" name="Indonesia" pageId="64" pageNumber="65">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D5C921BFEEDFCE7C9C02B11" box="[382,406,857,883]" pageId="64" pageNumber="65" type="male">♂</specimenCount>
; “
<collectingRegion id="505AF8029D5C921BFE21FCE6CA862B10" box="[434,720,856,882]" country="Indonesia" name="Sulawesi Tengah" pageId="64" pageNumber="65">SULAWESI TENGAH</collectingRegion>
:
<collectorName id="3F6B53369D5C921BFD4CFCE6CB2D2B10" box="[735,891,856,882]" pageId="64" pageNumber="65">Nr.Morowali</collectorName>
,
<collectingMunicipality id="7245AC9A9D5C921BFC18FCE7CC082B10" box="[907,1118,856,883]" pageId="64" pageNumber="65">Ranu River Area</collectingMunicipality>
.
<date id="E62010209D5C921BFBFDFCE6CD712B10" box="[1134,1319,856,882]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">
<collectingDate id="F664E9C89D5C921BFBFDFCE6CD712B10" box="[1134,1319,856,882]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">27.i.-20.iv.1980</collectingDate>
</date>
//
<location id="9741603B9D5C921BFAADFCE7C9532BF4" LSID="urn:lsid:plazi:treatment:1A3787F69D5C921AFDFCFEB6CA9E2D8E:9741603B9D5C921BFAADFCE7C9532BF4" country="Indonesia" municipality="Ranu River Area" name="Vert. Series" pageId="64" pageNumber="65" stateProvince="SULAWESI TENGAH">Vert. Series</location>
<quantity id="55669B059D5C921BFE9FFCC2C9172BF4" box="[268,321,892,918]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="64" pageNumber="65" unit="m" value="30.0">30m</quantity>
.actinic code:
<date id="E62010209D5C921BFE73FCC2CA682BF4" box="[480,574,892,918]" pageId="64" pageNumber="65" value="1980-03-11">
<collectingDate id="F664E9C89D5C921BFE73FCC2CA682BF4" box="[480,574,892,918]" pageId="64" pageNumber="65" value="1980-03-11">11.iii.80</collectingDate>
</date>
//
<collectorName id="3F6B53369D5C921BFDC6FCC2CB172BF4" box="[597,833,892,918]" pageId="64" pageNumber="65">S.L.Sutton C.J.Rees</collectorName>
B.M.1980-281”;
<collectionCode id="F48FAE259D5C921BFB82FCC2CC3B2BF4" box="[1041,1133,892,918]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="64" pageNumber="65">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D5C921BFF79FC00CA452BBB" blockId="64.[189,1399,958,1128]" box="[234,531,958,985]" pageId="64" pageNumber="65">
<emphasis id="A0EAEAF29D5C921BFF79FC00C9312BBA" bold="true" box="[234,359,958,984]" pageId="64" pageNumber="65">
<typeStatus id="4D2588429D5C921BFF79FC00C9312BBA" box="[234,359,958,984]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D5C921BFEEBFC00CA5D2BBB" box="[376,523,958,985]" pageId="64" pageNumber="65" type="generic">2 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D5C921BFF2EFC5CCB782C0A" blockId="64.[189,1399,958,1128]" pageId="64" pageNumber="65">
<materialsCitation id="22F63CBD9D5C921BFF2EFC5CCC282C42" ID-GBIF-Occurrence="2898878362" collectingDate="1980-01-27" collectingDateMax="1980-04-20" collectingDateMin="1980-01-27" collectionCode="BMNH" collectorName="Nr. Morowali &amp; S. L. Sutton C. J. Rees" country="Indonesia" elevation="30" location="Ranu River Area" municipality="Ranu River Area" pageId="64" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="SULAWESI TENGAH">
<collectingCountry id="EA8976709D5C921BFF2EFC5CC90D2B9E" box="[189,347,994,1020]" name="Indonesia" pageId="64" pageNumber="65">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D5C921BFEEDFC5CC9E62B9E" box="[382,432,994,1020]" pageId="64" pageNumber="65" type="female">1 ♀</specimenCount>
; “Vert.Series
<quantity id="55669B059D5C921BFDCEFC5CCAC42B9E" box="[605,658,994,1020]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="64" pageNumber="65" unit="m" value="30.0">30m</quantity>
.actinic code:
<date id="E62010209D5C921BFCA5FC5CCBC32B9F" box="[822,917,994,1021]" pageId="64" pageNumber="65" value="1980-03-15">
<collectingDate id="F664E9C89D5C921BFCA5FC5CCBC32B9F" box="[822,917,994,1021]" pageId="64" pageNumber="65" value="1980-03-15">15.iii.80</collectingDate>
</date>
//
<collectingRegion id="505AF8029D5C921BFC3FFC5CCC912B9E" box="[940,1223,994,1020]" country="Indonesia" name="Sulawesi Tengah" pageId="64" pageNumber="65">SULAWESI TENGAH</collectingRegion>
:
<collectorName id="3F6B53369D5C921BFB44FC5CCD242B9E" box="[1239,1394,994,1020]" pageId="64" pageNumber="65">Nr.Morowali</collectorName>
,
<collectingMunicipality id="7245AC9A9D5C921BFF2EFBB8C9DF2C42" box="[189,393,1030,1056]" pageId="64" pageNumber="65">Ranu River Area</collectingMunicipality>
.
<date id="E62010209D5C921BFE06FBB8CA182C42" box="[405,590,1030,1056]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">
<collectingDate id="F664E9C89D5C921BFE06FBB8CA182C42" box="[405,590,1030,1056]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">27.i.-20.iv.1980</collectingDate>
</date>
//
<collectorName id="3F6B53369D5C921BFDF5FBB8CB052C42" box="[614,851,1030,1056]" pageId="64" pageNumber="65">S.L.Sutton C.J.Rees</collectorName>
B.M.1980-281”;
<collectionCode id="F48FAE259D5C921BFBB5FBB8CC282C42" box="[1062,1150,1030,1056]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="64" pageNumber="65">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D5C921BFB0AFBB8CB7C2C0A" ID-GBIF-Occurrence="2898878329" collectingDate="1980-01-27" collectingDateMax="1980-04-20" collectingDateMin="1980-01-27" collectionCode="BMNH" collectorName="Nr. Morowali &amp; S. L. Sutton C. J. Rees" country="Indonesia" elevation="30" location="Ranu River Area" municipality="Ranu River Area" pageId="64" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="SULAWESI TENGAH">
<specimenCount id="8498FD699D5C921BFB0AFBB8CC9E2C42" box="[1177,1224,1030,1056]" pageId="64" pageNumber="65" type="female">1 ♀</specimenCount>
; “
<collectingRegion id="505AF8029D5C921BFB72FBB8C96C2C26" country="Indonesia" name="Sulawesi Tengah" pageId="64" pageNumber="65">SULAWESI TENGAH</collectingRegion>
:
<collectorName id="3F6B53369D5C921BFEC7FB94C9B92C26" box="[340,495,1066,1092]" pageId="64" pageNumber="65">Nr.Morowali</collectorName>
,
<collectingMunicipality id="7245AC9A9D5C921BFD99FB94CAA42C26" box="[522,754,1066,1092]" pageId="64" pageNumber="65">Ranu River Area</collectingMunicipality>
.
<date id="E62010209D5C921BFC98FB94CB922C26" box="[779,964,1066,1092]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">
<collectingDate id="F664E9C89D5C921BFC98FB94CB922C26" box="[779,964,1066,1092]" pageId="64" pageNumber="65" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">27.i.-20.iv.1980</collectingDate>
</date>
// Vert.Series
<quantity id="55669B059D5C921BFBE1FB94CCF12C26" box="[1138,1191,1066,1092]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="64" pageNumber="65" unit="m" value="30.0">30m</quantity>
.actinic code:
<date id="E62010209D5C921BFAC5FB94C8AC2C05" pageId="64" pageNumber="65" value="1980-03-13">
<collectingDate id="F664E9C89D5C921BFAC5FB94C8AC2C05" pageId="64" pageNumber="65" value="1980-03-13">13. iii.80</collectingDate>
</date>
//
<collectorName id="3F6B53369D5C921BFE81FBF0C9AB2C0A" box="[274,509,1102,1128]" pageId="64" pageNumber="65">S.L.Sutton C.J.Rees</collectorName>
B.M.1980-281”;
<collectionCode id="F48FAE259D5C921BFD5DFBF0CB7C2C0A" box="[718,810,1102,1128]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="64" pageNumber="65">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5C921AFF2EFB31CA8D2D6D" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
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<heading id="C969818C9D5C921BFF2EFB31C9182CCB" bold="true" box="[189,334,1167,1193]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="A0EAEAF29D5C921BFF2EFB31C9182CCB" bold="true" box="[189,334,1167,1193]" pageId="64" pageNumber="65">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5C921BFF2EFB01CC062D0B" blockId="64.[189,1399,1215,1385]" pageId="64" pageNumber="65">
MEASUREMENTS (exposed abdominal ventrites not included). Male
<typeStatus id="4D2588429D5C921BFC50FB01CC7B2CBB" box="[963,1069,1215,1241]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D5C921BFA94FB01CD242CBB" box="[1287,1394,1215,1241]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="64" pageNumber="65" unit="mm" value="7.4">7.40 mm</quantity>
. Head
<quantity id="55669B059D5C921BFE97FB5CC9382C9F" box="[260,366,1250,1277]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="64" pageNumber="65" unit="mm" value="1.5">1.50 mm</quantity>
long, across eyes
<quantity id="55669B059D5C921BFDD9FB5DCAE22C9F" box="[586,692,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="64" pageNumber="65" unit="mm" value="1.78">1.78 mm</quantity>
wide. Pronotum
<quantity id="55669B059D5C921BFC10FB5DCBBB2C9F" box="[899,1005,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="64" pageNumber="65" unit="mm" value="1.8">1.80 mm</quantity>
long, maximum width
<quantity id="55669B059D5C921BFA96FB5DCD252C9F" box="[1285,1395,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="64" pageNumber="65" unit="mm" value="2.8">2.80 mm</quantity>
. Elytral length
<quantity id="55669B059D5C921BFEFEFAB9C98B2D43" box="[365,477,1287,1313]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="64" pageNumber="65" unit="mm" value="4.1">4.10 mm</quantity>
, maximum combined width
<quantity id="55669B059D5C921BFCAFFAB8CBFD2D43" box="[828,939,1286,1313]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="64" pageNumber="65" unit="mm" value="3.5">3.50 mm</quantity>
. Selected female
<typeStatus id="4D2588429D5C921BFB16FAB9CCBB2D43" box="[1157,1261,1287,1313]" pageId="64" pageNumber="65" type="paratype">paratype</typeStatus>
, total body length
<quantity id="55669B059D5C921BFE9CFA94C92A2D27" box="[271,380,1322,1349]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="64" pageNumber="65" unit="mm" value="8.5">8.50 mm</quantity>
. Head
<quantity id="55669B059D5C921BFE5DFA94CA6E2D27" box="[462,568,1322,1349]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="64" pageNumber="65" unit="mm" value="1.5">1.50 mm</quantity>
long, across eyes
<quantity id="55669B059D5C921BFC80FA95CB2A2D27" box="[787,892,1323,1349]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="64" pageNumber="65" unit="mm" value="1.87">1.87 mm</quantity>
wide. Pronotum 2.00 mm long, maximum width
<quantity id="55669B059D5C921BFE94FAF0C9252D0B" box="[263,371,1358,1385]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="64" pageNumber="65" unit="mm" value="3.2">3.20 mm</quantity>
. Elytral length 5.00 mm, maximum combined width
<quantity id="55669B059D5C921BFC73FAF0CC1A2D0B" box="[992,1100,1358,1385]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="64" pageNumber="65" unit="mm" value="4.1">4.10 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D5C921AFF2EFA2FCB3D2C75" blockId="64.[189,1399,1424,2025]" lastBlockId="65.[189,1399,265,1048]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
Dorsum and venter uniformly black with dark castaneous labroclypeus and anterior and lateral margins of pronotum, mouthparts and legs. Head transverse, trapezoid, slightly convex on frons in lateral view, glossy dorsally and ventrally, with large not prominent compound eyes occupying nearly half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D5C921BFE6AFA42CA372E75" box="[505,609,1532,1559]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="64" pageNumber="65">Fig. 136</figureCitation>
) truncate, its lateral margins smooth, straight in both dorsal and lateral views. Anterolateral angles obtuse angulate, not protruding. Female labroclypeus generally as in male, anterolateral angles strongly obtuse. Frons punctures circular to irregularly hexagonal, smaller in anterior, somewhat larger in posterior half. Intervening spaces glossy, much smaller than punctures, in part wrinkled. Setae dirty yellowish, sparse, somewhat thickened and suberect; seta rises from anterior margin of corresponding puncture, surpassing or not its length. Longer setae present near compound eyes and on underside of labroclypeus. Labroclypeus laterally and frontally with longer and slender setae. Male and female antenna 9-segmented, club 3-lamellate. Scape large, strongly widened on upper side in distal half, provided with few long erect setae on posterior and distal margins. Antennomere 2 short cylindrical, transverse or nearly so. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum rather deeply emarginate, anterolateral angles protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum broadly rounded, slightly constricted towards base, delicately crenulate all along (
<figureCitation id="0AA52A659D5C921BFDEBF814CAB22FA7" box="[632,740,1962,1989]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="64" pageNumber="65">Fig. 224</figureCitation>
). Lateral margin of pronotum sinuous in lateral view. Punctures of pronotal disc of irregular shape (generally ovoid), deep and large, becoming flatter along lateral margins; inner margin with delicate membrane covered with microscopical velvety pubescence. Intervening spaces much smaller than punctures, in part wrinkled, glossy. Setation as on head; setae generally not surpassing length of corresponding punctures. Longer and thicker appressed seta between each of two lateral crenulae and all along basal margin. Hypomeron smooth, its anterolateral margin flange-like produced, with large sparse circular punctures, with large shallow emargination opposite to eye (to receive canthus separating compound eye), covered with long setae. Antennal pocket deep. Scutellar shield triangular, narrowly rounded apically. Elytra broadly cylindrical, maximum width across median third, glossy and slightly convex dorsally, with rounded humeri. Indistinct tracks of three flat longitudinal carinae on each elytron. Short sutural carinae indicated near apices. Rather large obtuse hump on each elytron near apex. Disc and lateral sides of elytra irregularly densely punctured, punctures irregularly ovoid (
<figureCitation id="0AA52A659D5D921AFD94FDDFCA3A2A1E" box="[519,620,609,636]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="65" pageNumber="66">Fig. 315</figureCitation>
). Periphery of punctures in part covered with microscopical velvety pubescence, completely or not encircling them. Intervening spaces generally smaller than punctures, glossy, in part microreticulate. Inconspicuous suberect seta rises from anterior margin of each puncture. Epipleuron with row or two (in broad anterior part) of suberect setae. Pygidium and abdominal ventrites glossy. Pygidium in both sexes dorsally flattened, covered with moderately large and deep irregularly ovoid punctures, each provided with moderately long suberect seta (
<figureCitation id="0AA52A659D5D921AFC78FCB3CC032B45" box="[1003,1109,781,807]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="65" pageNumber="66">Fig. 483</figureCitation>
). Legs long and slender, femora and tibiae covered with delicate whitish setae. Pro- and mesotibiae less distinct, metatibiae stronger thickened distally. Outer margins of meso- and metatibiae densely denticulate. Two teeth on external margin of protibia in both sexes (Fig. 388). Male lower meso- and metatibial terminal spur strongly curved. Distal margin of metatibia in both sexes with projection over basal metatarsomere. Male metatibial terminal spurs shorter than basal metatarsomere, female one as long as this. Male basal metatarsomere widened and dorso-ventrally flattened. Tarsal claws with pulvilli in both sexes. Spiculum gastrale as in
<figureCitation id="0AA52A659D5D921AFECEFC43C9952C7A" box="[349,451,1021,1048]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="65" pageNumber="66">Fig. 549</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D5D921AFD51FC43CB302C75" box="[706,870,1021,1047]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="65" pageNumber="66">Figs 640–642</figureCitation>
.
</paragraph>
<paragraph id="922136E09D5D921AFF2EFB80C9E62C3A" blockId="65.[189,432,1086,1112]" box="[189,432,1086,1112]" pageId="65" pageNumber="66">
<heading id="C969818C9D5D921AFF2EFB80C9E62C3A" bold="true" box="[189,432,1086,1112]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="A0EAEAF29D5D921AFF2EFB80C9E62C3A" bold="true" box="[189,432,1086,1112]" pageId="65" pageNumber="66">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5D921AFF2EFBD2CA8D2D6D" blockId="65.[189,1398,1132,1296]" pageId="65" pageNumber="66">Female is generally larger; lamellae of antennal club shorter; protibial spur slightly curved distally in male, straight in female; male metatibial terminal spur shorter than, in female as long as basal metatarsomere; apical margin of female pygidium with vague emargination, not emarginate in male; female basal metatarsomere not conspicuously widened or flattened; female lower meso- and metatibial terminal spur slender, slightly curved apically.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5D921AFF2EFA88CB512D1C" box="[189,775,1334,1406]" pageId="65" pageNumber="66" type="biology_ecology">
<paragraph id="922136E09D5D921AFF2EFA88C9762D32" blockId="65.[189,288,1334,1360]" box="[189,288,1334,1360]" pageId="65" pageNumber="66">
<heading id="C969818C9D5D921AFF2EFA88C9762D32" bold="true" box="[189,288,1334,1360]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="A0EAEAF29D5D921AFF2EFA88C9762D32" bold="true" box="[189,288,1334,1360]" pageId="65" pageNumber="66">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5D921AFF2EFADACB512D1C" blockId="65.[189,775,1380,1406]" box="[189,775,1380,1406]" pageId="65" pageNumber="66">Occurs in lowland rainforests. Possibly nocturnal.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5D921AFF2EFA1ACA9E2D8E" box="[189,712,1444,1516]" pageId="65" pageNumber="66" type="distribution">
<paragraph id="922136E09D5D921AFF2EFA1AC9002DDC" blockId="65.[189,342,1444,1470]" box="[189,342,1444,1470]" pageId="65" pageNumber="66">
<heading id="C969818C9D5D921AFF2EFA1AC9002DDC" bold="true" box="[189,342,1444,1470]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="A0EAEAF29D5D921AFF2EFA1AC9002DDC" bold="true" box="[189,342,1444,1470]" pageId="65" pageNumber="66">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5D921AFF2EFA6CCA9E2D8E" blockId="65.[189,712,1490,1516]" box="[189,712,1490,1516]" pageId="65" pageNumber="66">
Hitherto only known from
<collectingRegion id="505AF8029D5D921AFE6AFA6CCA932D8E" box="[505,709,1490,1516]" country="Indonesia" name="Sulawesi Tengah" pageId="65" pageNumber="66">Central Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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