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<document id="7F8B843E58A807264059E7AD8931D76F" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D429204FDAFFCE8CD652999" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius pedarioides , Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="96" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="95" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F3D1EBC6CF816425242A083C1FF90EED">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="913F727805ABB7D8FB5ED24D145BC693">Telnov, Dmitry</mods:namePart>
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<mods:title id="97DEEAB2097D989FD62A6BE3E673E901">European Journal of Taxonomy</mods:title>
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<mods:date id="8FD167A9212BE668B8778D97C0F1AB75">2020</mods:date>
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<mods:number id="F130EB12918CDA35E00FA5CC3B685E0A">2020-10-19</mods:number>
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<treatment id="1A3787F69D429204FDAFFCE8CD652999" ID-DOI="http://doi.org/10.5281/zenodo.4344449" ID-GBIF-Taxon="168648154" ID-Zenodo-Dep="4344449" LSID="urn:lsid:plazi:treatment:1A3787F69D429204FDAFFCE8CD652999" httpUri="http://treatment.plazi.org/id/1A3787F69D429204FDAFFCE8CD652999" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95">
<subSubSection id="DA84656B9D429205FDAFFCE8CBAE2B13" box="[572,1016,854,881]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="922136E09D429205FDAFFCE8CBAE2B13" blockId="94.[457,1130,854,917]" box="[572,1016,854,881]" pageId="94" pageNumber="95">
<heading id="C969818C9D429205FDAFFCE8CBAE2B13" box="[572,1016,854,881]" centered="true" fontSize="11" level="2" pageId="94" pageNumber="95" reason="2">
<taxonomicName id="559E4D639D429205FDAFFCE8CBAE2B13" ID-CoL="3XFKR" authority="Arrow, 1941" authorityName=", Arrow" authorityYear="1941" box="[572,1016,854,881]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="pedarioides">
<emphasis id="A0EAEAF29D429205FDAFFCE8CB0A2B13" bold="true" box="[572,860,854,881]" italics="true" pageId="94" pageNumber="95">Maechidius pedarioides</emphasis>
<bibRefCitation id="F60F4B119D429205FCF0FCE9CBAE2B13" author="Arrow G. E." box="[867,1016,855,881]" pageId="94" pageNumber="95" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D429205FE5AFCC4CC3C2BF6" box="[457,1130,890,917]" pageId="94" pageNumber="95" type="description">
<paragraph id="922136E09D429205FE5AFCC4CC3C2BF6" blockId="94.[457,1130,854,917]" box="[457,1130,890,917]" pageId="94" pageNumber="95">
<figureCitation id="0AA52A659D429205FE5AFCC4CA752BF6" box="[457,547,890,916]" captionStart="Figs 70–73" captionStartId="93.[189,241,1896,1922]" captionTargetBox="[327,1247,269,1869]" captionTargetId="figure-67@93.[327,1254,265,1878]" captionTargetPageId="93" captionText="Figs 70–73. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 70. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 71. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 72. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 73. M. peregrinus Lansberge, 1886, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122158" httpUri="https://zenodo.org/record/4122158/files/figure.png" pageId="94" pageNumber="95">Figs 72</figureCitation>
,
<figureCitation id="0AA52A659D429205FDBCFCC4CACE2BF6" box="[559,664,890,916]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="94" pageNumber="95">163–164</figureCitation>
,
<figureCitation id="0AA52A659D429205FD37FCC4CA852BF6" box="[676,723,890,916]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="94" pageNumber="95">244</figureCitation>
,
<figureCitation id="0AA52A659D429205FD4CFCC4CB582BF6" box="[735,782,890,916]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="94" pageNumber="95">334</figureCitation>
, 407, 507–508, 556, 688–690
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D429205FF2EFC02C94B2D21" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="922136E09D429205FF2EFC02C93A2BB7" blockId="94.[189,364,955,982]" box="[189,364,955,982]" pageId="94" pageNumber="95">
<heading id="C969818C9D429205FF2EFC02C93A2BB7" bold="true" box="[189,364,955,982]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A0EAEAF29D429205FF2EFC02C93A2BB7" bold="true" box="[189,364,955,982]" pageId="94" pageNumber="95">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D429205FF79FC54C90C2C66" blockId="94.[189,1399,1002,1347]" box="[234,346,1002,1028]" pageId="94" pageNumber="95">
<emphasis id="A0EAEAF29D429205FF79FC54C90C2C66" bold="true" box="[234,346,1002,1028]" pageId="94" pageNumber="95">
<typeStatus id="4D2588429D429205FF79FC54C90C2C66" box="[234,346,1002,1028]" pageId="94" pageNumber="95" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D429205FF2EFBB0C94B2D21" blockId="94.[189,1399,1002,1347]" pageId="94" pageNumber="95">
<materialsCitation id="22F63CBD9D429205FF2EFBB0CBD42C0D" ID-GBIF-Occurrence="2898878470" collectingDate="1938-04" collectionCode="DUTCH, NEW, BMNH" country="Guinea-Bissau" elevation="762" location="Waigeu. Camp Nok." pageId="94" pageNumber="95" specimenCount="1" typeStatus="syntype">
<collectingCountry id="EA8976709D429205FF2EFBB0C90D2C4A" box="[189,347,1038,1064]" name="Indonesia" pageId="94" pageNumber="95">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D429205FEE4FBB0C9D92C4A" box="[375,399,1038,1064]" pageId="94" pageNumber="95">♂</specimenCount>
; “
<typeStatus id="4D2588429D429205FE3BFBB0CA792C4A" box="[424,559,1038,1064]" pageId="94" pageNumber="95" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] //
<typeStatus id="4D2588429D429205FC3FFBB0CBB02C4A" box="[940,998,1038,1064]" pageId="94" pageNumber="95">Type</typeStatus>
[p, label circular, red frame] //
<specimenCount id="8498FD699D429205FAF3FBB0CD202C4A" box="[1376,1398,1038,1064]" pageId="94" pageNumber="95">♂</specimenCount>
[p] // N.
<collectionCode id="F48FAE259D429205FE83FB8FC92F2C29" box="[272,377,1073,1099]" pageId="94" pageNumber="95">DUTCH</collectionCode>
<collectionCode id="F48FAE259D429205FE17FB8FC9902C29" box="[388,454,1073,1099]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="94" pageNumber="95">NEW</collectionCode>
<collectingCountry id="EA8976709D429205FE41FB8FCA112C29" box="[466,583,1073,1099]" name="Guinea-Bissau" pageId="94" pageNumber="95">GUINEA</collectingCountry>
:
<location id="9741603B9D429205FDC5FB8FCB182C29" LSID="urn:lsid:plazi:treatment:1A3787F69D429204FDAFFCE8CD652999:9741603B9D429205FDC5FB8FCB182C29" box="[598,846,1073,1099]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="94" pageNumber="95">Waigeu. Camp Nok.</location>
<quantity id="55669B059D429205FCCAFB8FCB962C29" box="[857,960,1073,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">
<elevation id="19B3D1D39D429205FCCAFB8FCB962C29" box="[857,960,1073,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">2,500 ft.</elevation>
</quantity>
<date id="E62010209D429205FC5FFB8FCC702C29" box="[972,1062,1073,1099]" pageId="94" pageNumber="95" value="1938-04">
<collectingDate id="F664E9C89D429205FC5FFB8FCC702C29" box="[972,1062,1073,1099]" pageId="94" pageNumber="95" value="1938-04">iv.1938</collectingDate>
</date>
. L.E.Cheesman. B.M.1938- 593. [p]//
<taxonomicName id="559E4D639D429205FEA1FBEACAFF2C0C" authority=", Arrow" authorityName=", Arrow" authorityYear="1941" box="[306,681,1108,1135]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="pedarioides">
<emphasis id="A0EAEAF29D429205FEA1FBEACA1B2C0C" box="[306,589,1108,1134]" italics="true" pageId="94" pageNumber="95">Maechidius pedarioides</emphasis>
, Arrow
</taxonomicName>
<typeStatus id="4D2588429D429205FD23FBEBCAB52C0D" box="[688,739,1109,1135]" pageId="94" pageNumber="95">type</typeStatus>
[h]”;
<collectionCode id="F48FAE259D429205FCB9FBEBCBD42C0D" box="[810,898,1109,1135]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="94" pageNumber="95">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D429205FC0EFBEACAC82CD7" ID-GBIF-Occurrence="2898878532" collectingDate="1938-04" collectionCode="DUTCH, NEW, BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="Waigeu. Camp Nok." pageId="94" pageNumber="95" specimenCount="1" typeStatus="syntype">
<specimenCount id="8498FD699D429205FC0EFBEACB9A2C0D" box="[925,972,1108,1135]" pageId="94" pageNumber="95" type="male">1 ♂</specimenCount>
; “
<typeStatus id="4D2588429D429205FC77FBEACC3D2C0C" box="[996,1131,1108,1134]" pageId="94" pageNumber="95" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] // N.
<collectionCode id="F48FAE259D429205FEAFFBC6C9F32CF0" box="[316,421,1144,1170]" pageId="94" pageNumber="95">DUTCH</collectionCode>
<collectionCode id="F48FAE259D429205FE3FFBC6C9B82CF0" box="[428,494,1144,1170]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="94" pageNumber="95">NEW</collectionCode>
<collectingCountry id="EA8976709D429205FE66FBC6CA3C2CF0" box="[501,618,1144,1170]" name="Guinea-Bissau" pageId="94" pageNumber="95">GUINEA</collectingCountry>
:
<location id="9741603B9D429205FDE6FBC9CB322CF3" LSID="urn:lsid:plazi:treatment:1A3787F69D429204FDAFFCE8CD652999:9741603B9D429205FDE6FBC9CB322CF3" box="[629,868,1143,1170]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="94" pageNumber="95">Waigeu. Camp Nok.</location>
<quantity id="55669B059D429205FCF8FBC9CB982CF3" box="[875,974,1143,1170]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">
<elevation id="19B3D1D39D429205FCF8FBC9CB982CF3" box="[875,974,1143,1170]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">2,500 ft.</elevation>
</quantity>
<date id="E62010209D429205FC46FBC9CC792CF3" box="[981,1071,1143,1169]" pageId="94" pageNumber="95" value="1938-04">
<collectingDate id="F664E9C89D429205FC46FBC9CC792CF3" box="[981,1071,1143,1169]" pageId="94" pageNumber="95" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D429205FBA9FBC9CCB92CF3" box="[1082,1263,1143,1169]" pageId="94" pageNumber="95">L.E.Cheesman.</collectorName>
B.M.1938- 593. [p] //
<taxonomicName id="559E4D639D429205FEA2FB25C9EA2CD7" authorityName=", Arrow" authorityYear="1941" box="[305,444,1179,1205]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="pedarioides">
<emphasis id="A0EAEAF29D429205FEA2FB25C9EA2CD7" box="[305,444,1179,1205]" italics="true" pageId="94" pageNumber="95">pedarioides</emphasis>
</taxonomicName>
? var. [h]”;
<collectionCode id="F48FAE259D429205FDD5FB25CAC82CD7" box="[582,670,1179,1205]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="94" pageNumber="95">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D429205FD25FB25CAC92C9E" ID-GBIF-Occurrence="2898878415" collectingDate="1938-04" collectionCode="DUTCH, NEW, BMNH" country="Guinea-Bissau" elevation="762" location="Waigeu. Camp Nok." pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<specimenCount id="8498FD699D429205FD25FB25CAB52CD7" box="[694,739,1179,1205]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
; “
<typeStatus id="4D2588429D429205FD68FB25CBD42CD7" box="[763,898,1179,1205]" pageId="94" pageNumber="95" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] // N.
<collectionCode id="F48FAE259D429205FA9DFB25CD212CD7" box="[1294,1399,1179,1205]" pageId="94" pageNumber="95">DUTCH</collectionCode>
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<collectingCountry id="EA8976709D429205FE96FB00C92C2CBA" box="[261,378,1214,1240]" name="Guinea-Bissau" pageId="94" pageNumber="95">GUINEA</collectingCountry>
:
<location id="9741603B9D429205FE10FB00CA392CBA" LSID="urn:lsid:plazi:treatment:1A3787F69D429204FDAFFCE8CD652999:9741603B9D429205FE10FB00CA392CBA" box="[387,623,1214,1240]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="94" pageNumber="95">Waigeu. Camp Nok.</location>
<quantity id="55669B059D429205FDE6FB00CA802CBA" box="[629,726,1214,1241]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">
<elevation id="19B3D1D39D429205FDE6FB00CA802CBA" box="[629,726,1214,1241]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">2,500 ft.</elevation>
</quantity>
<date id="E62010209D429205FD4FFB00CB602CBA" box="[732,822,1214,1240]" pageId="94" pageNumber="95" value="1938-04">
<collectingDate id="F664E9C89D429205FD4FFB00CB602CBA" box="[732,822,1214,1240]" pageId="94" pageNumber="95" value="1938-04">iv.1938</collectingDate>
</date>
. L.E.Cheesman. B.M.1938-593. [p] //
<taxonomicName id="559E4D639D429205FB7DFB00C91E2C9E" authorityName=", Arrow" authorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="pedarioides">
<emphasis id="A0EAEAF29D429205FB7DFB00C91E2C9E" italics="true" pageId="94" pageNumber="95">Maechidius pedarioides</emphasis>
</taxonomicName>
, co-type Arrow [h]”;
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</materialsCitation>
•
<materialsCitation id="22F63CBD9D429205FD2BFB5CC94F2D21" ID-GBIF-Occurrence="2898878456" collectingDate="1938-04" collectionCode="DUTCH, NEW, BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="Waigeu. Camp Nok." pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
<specimenCount id="8498FD699D429205FD2BFB5CCAB32C9E" box="[696,741,1250,1276]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
; “
<typeStatus id="4D2588429D429205FD6DFB5CCBD32C9E" box="[766,901,1250,1276]" pageId="94" pageNumber="95" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] //
<specimenCount id="8498FD699D429205FA90FB5CCD4F2C9E" box="[1283,1305,1250,1276]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
[p] // N.
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<collectionCode id="F48FAE259D429205FEB8FABBC93B2D7D" box="[299,365,1285,1311]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="94" pageNumber="95">NEW</collectionCode>
<collectingCountry id="EA8976709D429205FEE5FABBC9BD2D7D" box="[374,491,1285,1311]" name="Guinea-Bissau" pageId="94" pageNumber="95">GUINEA</collectingCountry>
:
<location id="9741603B9D429205FE64FABBCABF2D7D" LSID="urn:lsid:plazi:treatment:1A3787F69D429204FDAFFCE8CD652999:9741603B9D429205FE64FABBCABF2D7D" box="[503,745,1285,1311]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="94" pageNumber="95">Waigeu. Camp Nok.</location>
<quantity id="55669B059D429205FD62FABBCB002D7D" box="[753,854,1285,1312]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">
<elevation id="19B3D1D39D429205FD62FABBCB002D7D" box="[753,854,1285,1312]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="94" pageNumber="95" unit="ft" value="2500.0">2,500 ft.</elevation>
</quantity>
<date id="E62010209D429205FCCDFABBCBEF2D7D" box="[862,953,1285,1311]" pageId="94" pageNumber="95" value="1938-04">
<collectingDate id="F664E9C89D429205FCCDFABBCBEF2D7D" box="[862,953,1285,1311]" pageId="94" pageNumber="95" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D429205FC56FABBCC2C2D7D" box="[965,1146,1285,1311]" pageId="94" pageNumber="95">L.E.Cheesman.</collectorName>
B.M.1938-593. [p]”;
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D429204FF2EFAD4CD652999" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95" type="description">
<paragraph id="922136E09D429205FF2EFAD4C9182DE6" blockId="94.[189,334,1386,1412]" box="[189,334,1386,1412]" pageId="94" pageNumber="95">
<heading id="C969818C9D429205FF2EFAD4C9182DE6" bold="true" box="[189,334,1386,1412]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A0EAEAF29D429205FF2EFAD4C9182DE6" bold="true" box="[189,334,1386,1412]" pageId="94" pageNumber="95">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D429204FF2EFA27CA4829EE" blockId="94.[189,1399,1432,2025]" lastBlockId="95.[189,1399,265,397]" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95">
Male labroclypeus (
<figureCitation id="0AA52A659D429205FE3EFA27CA422DD1" box="[429,532,1432,1459]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="94" pageNumber="95">Fig. 164</figureCitation>
) broadly emarginate on anterior margin, female labroclypeus shallowly emarginate. Male anterolateral angles of labroclypeus rounded, strongly protruding, female one broadly obtuse, not protruding (
<figureCitation id="0AA52A659D429205FE58FA61CA662D98" box="[459,560,1503,1530]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="94" pageNumber="95">Fig. 164</figureCitation>
). Lateral margin of labroclypeus sinuous in both dorsal and lateral views. Punctures of frons circular to ovoid, variably large, shallow, those on vertex annular. Long suberect seta rises from anterior margin of each puncture. Pronotum broadly emarginate on anterior margin, anterolateral angles protruding. Basal margin of pronotum broadly rounded. Lateral margin of pronotum in dorsal view broadly rounded, all along with large deep punctures splitting margin into indistinct long and flat crenulae (
<figureCitation id="0AA52A659D429205FE1FF92EC9A52EC8" box="[396,499,1680,1706]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="94" pageNumber="95">Fig. 244</figureCitation>
). Long erect seta present in each puncture between two crenulae. Hypomeron separated from prosternum by low arched carina, with long setae on anterolateral margin opposite to compound eye. Antennal pocket shallow. Pronotal punctures large, shallow, ovoid, those along lateral sides annular. Intervening spaces glossy, smaller to much smaller than punctures. Pronotal punctures in part with inner margin with delicate membrane covered with microscopical velvety pubescence. Pronotal setae minute, seta rises from anterior margin of each puncture, not surpassing its midlength. Elytral punctures circular to ovoid, in part annular, shallow, arranged in double longitudinal rows (
<figureCitation id="0AA52A659D429205FF55F836C9642FC0" box="[198,306,1928,1954]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="94" pageNumber="95">Fig. 334</figureCitation>
). Intervening spaces glossy, generally larger than punctures. Elytral setae minute, surface looks glabrous under 50 × magnification; minute seta rises from anterior margin of each puncture, not surpassing midlength of corresponding puncture. Male pygidium flattened dorsally, with large shallow annular punctures. Intervening spaces variably large, generally smaller than punctures. Setae of pygidium sparse, long, suberect. Male and female protibia widened distally, with three distal teeth on external margin (Fig. 407). Male terminal spur of protibia short. Spiculum gastrale as in
<figureCitation id="0AA52A659D439204FA9FFEEECD252908" box="[1292,1395,335,362]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="95" pageNumber="96">Fig. 556</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D439204FEE5FECCCA4C29EE" box="[374,538,370,396]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="95" pageNumber="96">Figs 688–690</figureCitation>
.
</paragraph>
<paragraph id="922136E09D439204FF2EFE0DC9E629AF" blockId="95.[189,432,435,461]" box="[189,432,435,461]" pageId="95" pageNumber="96">
<heading id="C969818C9D439204FF2EFE0DC9E629AF" bold="true" box="[189,432,435,461]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
<emphasis id="A0EAEAF29D439204FF2EFE0DC9E629AF" bold="true" box="[189,432,435,461]" pageId="95" pageNumber="96">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D439204FF2EFE5FCD652999" blockId="95.[189,1331,481,508]" box="[189,1331,481,508]" pageId="95" pageNumber="96">Female lamellae of antennal club considerably shorter; terminal spur of protibia distinctly longer.</paragraph>
</subSubSection>
</treatment>
</document>
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