247 lines
39 KiB
XML
247 lines
39 KiB
XML
<document id="0CC509DC47E348DA847B72CC495CC580" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D2D9269FDF9FE35CC382DD5" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius hamatus Telnov 2020, sp. nov." docType="treatment" docUuid="DCA112DF-06B6-4767-8991-20D54ED810E7" docUuidSource="ZooBank" docVersion="10" lastPageNumber="51" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="50" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C1D296550B60D5A99A88B86398FEDC30">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:roleTerm id="AE5FD3B0061CCAE20E879FF76AE4050A">Author</mods:roleTerm>
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<mods:namePart id="6C0BB204B355BE73CD396C5BAE7889DB">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="3A94ECD4AF66221980A1F6E1563C3206">text</mods:typeOfResource>
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<mods:title id="3786C6289DC163BB15310A9AF14248EA">European Journal of Taxonomy</mods:title>
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<mods:part id="198C8B5F14BDA7FB1999C1A948B32C5D">
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<mods:date id="8676CFE3514D48A8044C4591D717786B">2020</mods:date>
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<mods:number id="99F44A726B12E08CA9B7E7212FDFD055">2020-10-19</mods:number>
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<mods:number id="E39B49B3C840B95FEA47AAF85EEF5014">721</mods:number>
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<mods:classification id="06446807BFBE107FE55331D27ACCF8D0">journal article</mods:classification>
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<mods:identifier id="5A9D2A845F0FF328BA00CF8E5DA4D25A" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D2D9269FDF9FE35CC382DD5" ID-DOI="http://doi.org/10.5281/zenodo.4344347" ID-GBIF-Taxon="168648138" ID-Zenodo-Dep="4344347" LSID="urn:lsid:zoobank.org:act:DCA112DF-06B6-4767-8991-20D54ED810E7" httpUri="http://treatment.plazi.org/id/1A3787F69D2D9269FDF9FE35CC382DD5" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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<subSubSection id="DA84656B9D2D926AFDF9FE35CB9C29C4" box="[618,970,395,422]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="922136E09D2D926AFDF9FE35CB9C29C4" blockId="49.[368,1220,395,490]" box="[618,970,395,422]" pageId="49" pageNumber="50">
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<heading id="C969818C9D2D926AFDF9FE35CB9C29C4" box="[618,970,395,422]" centered="true" fontSize="11" level="2" pageId="49" pageNumber="50" reason="2">
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<taxonomicName id="559E4D639D2D926AFDF9FE35CB3129C4" ID-CoL="7X3WR" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[618,871,395,422]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
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<emphasis id="A0EAEAF29D2D926AFDF9FE35CB3129C4" bold="true" box="[618,871,395,422]" italics="true" pageId="49" pageNumber="50">Maechidius hamatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D2D926AFCFCFE32CB9C29C4" box="[879,970,396,422]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D2D926AFEE3FE13CCDE298B" pageId="49" pageNumber="50" type="description">
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<paragraph id="922136E09D2D926AFEE3FE13CC9229AA" blockId="49.[368,1220,395,490]" box="[368,1220,429,456]" pageId="49" pageNumber="50">
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<uri id="E60F3AE29D2D926AFEE3FE13CC9229AA" box="[368,1220,429,456]" pageId="49" pageNumber="50">
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urn:lsid:zoobank.org:act:
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<uuid id="E6380C359D2D926AFD1DFE13CC9229AA" box="[654,1220,429,456]" pageId="49" pageNumber="50">DCA112DF-06B6-4767-8991-20D54ED810E7</uuid>
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</uri>
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</paragraph>
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<paragraph id="922136E09D2D926AFE38FE71CCDE298B" blockId="49.[368,1220,395,490]" box="[427,1160,463,490]" pageId="49" pageNumber="50">
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<figureCitation id="0AA52A659D2D926AFE38FE71CA53298B" box="[427,517,463,489]" captionStart="Figs 38–41" captionStartId="47.[189,241,1893,1919]" captionTargetBox="[360,1224,277,1874]" captionTargetId="figure-68@47.[360,1227,265,1875]" captionTargetPageId="47" captionText="Figs 38–41. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 38. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 39. M. hamatus sp. nov., holotype, ♂ (IECA). 40. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (BPBM) (photograph courtesy BPBM). 41. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122142" httpUri="https://zenodo.org/record/4122142/files/figure.png" pageId="49" pageNumber="50">Figs 39</figureCitation>
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,
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<figureCitation id="0AA52A659D2D926AFD82FE71CA69298B" box="[529,575,463,489]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="49" pageNumber="50">121</figureCitation>
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,
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<figureCitation id="0AA52A659D2D926AFDDFFE71CA2D2988" box="[588,635,463,490]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="49" pageNumber="50">215</figureCitation>
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,
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<figureCitation id="0AA52A659D2D926AFD14FE71CAE02988" box="[647,694,463,490]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="49" pageNumber="50">305</figureCitation>
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, 378–379, 431–432, 472, 544, 617–619
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DA84656B9D2D926AFF2EFDB0C93A2A14" pageId="49" pageNumber="50" type="diagnosis">
|
||
<paragraph id="922136E09D2D926AFF2EFDB0C99F2A4A" blockId="49.[189,457,526,552]" box="[189,457,526,552]" pageId="49" pageNumber="50">
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||
<heading id="C969818C9D2D926AFF2EFDB0C99F2A4A" bold="true" box="[189,457,526,552]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
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<emphasis id="A0EAEAF29D2D926AFF2EFDB0C99F2A4A" bold="true" box="[189,457,526,552]" pageId="49" pageNumber="50">Differential diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="922136E09D2D926AFF2EFD85C93A2A14" blockId="49.[189,1398,571,631]" pageId="49" pageNumber="50">
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This is the second species of the
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<taxonomicName id="559E4D639D2D926AFDC0FD85CAD12A37" authorityName="Telnov" authorityYear="2020" box="[595,647,571,597]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dani">
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<emphasis id="A0EAEAF29D2D926AFDC0FD85CAD12A37" box="[595,647,571,597]" italics="true" pageId="49" pageNumber="50">dani</emphasis>
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</taxonomicName>
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informal species-group, see the diagnosis of
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<taxonomicName id="559E4D639D2D926AFB3CFD85CD202A37" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1199,1398,571,597]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dani" status="sp. nov.">
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<emphasis id="A0EAEAF29D2D926AFB3CFD85CD202A37" box="[1199,1398,571,597]" italics="true" pageId="49" pageNumber="50">Maechidius dani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D2D926AFF2EFDE3C94E2A15" box="[189,280,605,631]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
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above.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D2D926AFF2EFD22CA5E2B66" pageId="49" pageNumber="50" type="etymology">
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<paragraph id="922136E09D2D926AFF2EFD22C9122AD4" blockId="49.[189,324,668,694]" box="[189,324,668,694]" pageId="49" pageNumber="50">
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<heading id="C969818C9D2D926AFF2EFD22C9122AD4" bold="true" box="[189,324,668,694]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
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<emphasis id="A0EAEAF29D2D926AFF2EFD22C9122AD4" bold="true" box="[189,324,668,694]" pageId="49" pageNumber="50">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D2D926AFF2EFD77CA5E2B66" blockId="49.[189,1398,713,773]" pageId="49" pageNumber="50">
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The specific name is derived from the Latin ‘
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<taxonomicName id="559E4D639D2D926AFD4AFD77CB6A2A81" authorityName="Telnov" authorityYear="2020" box="[729,828,713,739]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hamatus">
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<emphasis id="A0EAEAF29D2D926AFD4AFD77CB6A2A81" box="[729,828,713,739]" italics="true" pageId="49" pageNumber="50">hamatus</emphasis>
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</taxonomicName>
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’ (‘hooked’), referring to the curved denticles of male protarsomeres 1 and 2.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="DA84656B9D2D926AFF2EFC94CD1B2C54" pageId="49" pageNumber="50" type="materials_examined">
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<paragraph id="922136E09D2D926AFF2EFC94C93A2B26" blockId="49.[189,364,810,836]" box="[189,364,810,836]" pageId="49" pageNumber="50">
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||
<heading id="C969818C9D2D926AFF2EFC94C93A2B26" bold="true" box="[189,364,810,836]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
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||
<emphasis id="A0EAEAF29D2D926AFF2EFC94C93A2B26" bold="true" box="[189,364,810,836]" pageId="49" pageNumber="50">Type material</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="22F63CBD9D2D926AFF79FCE9CC532BB7" ID-GBIF-Occurrence="2898878602" collectingDate="2015-02-02" collectingDateMax="2015-02-03" collectingDateMin="2015-02-02" collectionCode="NMPC" collectorName="J. Hajek" country="Indonesia" elevation="2050" latitude="-4.06" location="Baliem valley" longLatPrecision="130" longitude="139.03166" municipality="Jayawijaya Distr." pageId="49" pageNumber="50" specimenCount="1" stateProvince="Papua" typeStatus="holotype">
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||
<paragraph id="922136E09D2D926AFF79FCE9C90A2B13" blockId="49.[189,1399,855,982]" box="[234,348,855,881]" pageId="49" pageNumber="50">
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||
<emphasis id="A0EAEAF29D2D926AFF79FCE9C90A2B13" bold="true" box="[234,348,855,881]" pageId="49" pageNumber="50">
|
||
<typeStatus id="4D2588429D2D926AFF79FCE9C90A2B13" box="[234,348,855,881]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
</paragraph>
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||
<paragraph id="922136E09D2D926AFF2EFCC6CC532BB7" blockId="49.[189,1399,855,982]" pageId="49" pageNumber="50">
|
||
<collectingCountry id="EA8976709D2D926AFF2EFCC6C90D2BF0" box="[189,347,888,914]" name="Indonesia" pageId="49" pageNumber="50">INDONESIA</collectingCountry>
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||
•
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||
<specimenCount id="8498FD699D2D926AFEEDFCC7C9C02BF1" box="[382,406,889,915]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
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||
; “
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||
<collectingCountry id="EA8976709D2D926AFE21FCC6CA0E2BF0" box="[434,600,888,914]" name="Indonesia" pageId="49" pageNumber="50">INDONESIA</collectingCountry>
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||
,
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<collectingRegion id="505AF8029D2D926AFDFAFCC7CAE32BF1" box="[617,693,889,915]" country="Indonesia" name="Papua Barat" pageId="49" pageNumber="50">Papua</collectingRegion>
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||
:
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<collectingMunicipality id="7245AC9A9D2D926AFD56FCC6CBCF2BF0" box="[709,921,888,914]" pageId="49" pageNumber="50">Jayawijaya Distr.</collectingMunicipality>
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||
:
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<location id="9741603B9D2D926AFC3BFCC6CC042BF0" LSID="urn:lsid:plazi:treatment:1A3787F69D2D9269FDF9FE35CC382DD5:9741603B9D2D926AFC3BFCC6CC042BF0" box="[936,1106,888,914]" country="Indonesia" latitude="-4.06" longLatPrecision="130" longitude="139.03166" municipality="Jayawijaya Distr." name="Baliem valley" pageId="49" pageNumber="50" stateProvince="Papua">Baliem valley</location>
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||
,
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<locationDeviation id="83932FC29D2D926AFBF0FCC6CD242BF1" box="[1123,1394,888,915]" location="Wamena" pageId="49" pageNumber="50">
|
||
<quantity id="55669B059D2D926AFBF0FCC6CCF02BF0" box="[1123,1190,888,914]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="km" value="10.0">10km</quantity>
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||
NE of Wamena
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</locationDeviation>
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, forest above “Baliem valley resort”,
|
||
<geoCoordinate id="F7AA50279D2D926AFDE2FC27CABC2BD1" box="[625,746,921,947]" degrees="04" direction="south" minutes="03.6" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="-4.06">04°03.6’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F7AA50279D2D926AFD64FC24CBD72BD6" box="[759,897,922,948]" degrees="139" direction="east" minutes="01.9" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="139.03166">139°01.9’E</geoCoordinate>
|
||
,
|
||
<quantity id="55669B059D2D926AFC1DFC27CBBD2BD6" box="[910,1003,921,948]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="49" pageNumber="50" unit="m" value="2050.0">
|
||
<elevation id="19B3D1D39D2D926AFC1DFC27CBBD2BD6" box="[910,1003,921,948]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="49" pageNumber="50" unit="m" value="2050.0">2050 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="E62010209D2D926AFC6AFC27CC2C2BD6" box="[1017,1146,921,948]" pageId="49" pageNumber="50" value="2015-02-02" valueMax="2015-02-03" valueMin="2015-02-02">
|
||
<collectingDate id="F664E9C89D2D926AFC6AFC27CC2C2BD6" box="[1017,1146,921,948]" pageId="49" pageNumber="50" value="2015-02-02" valueMax="2015-02-03" valueMin="2015-02-02">2-3.ii.2015</collectingDate>
|
||
</date>
|
||
<collectorName id="3F6B53369D2D926AFB11FC24CC8C2BD6" box="[1154,1242,922,948]" pageId="49" pageNumber="50">J.Hájek</collectorName>
|
||
& JŠumpich leg. // coll. general National Museum Prague, Czech Republic”;
|
||
<collectionCode id="F48FAE259D2D926AFC39FC05CC572BB7" box="[938,1025,955,981]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="49" pageNumber="50">NMPC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="922136E09D2D926AFF79FC45CA452C74" blockId="49.[189,1357,1019,1079]" box="[234,531,1019,1046]" pageId="49" pageNumber="50">
|
||
<emphasis id="A0EAEAF29D2D926AFF79FC45C9312C77" bold="true" box="[234,359,1019,1045]" pageId="49" pageNumber="50">
|
||
<typeStatus id="4D2588429D2D926AFF79FC45C9312C77" box="[234,359,1019,1045]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="8498FD699D2D926AFEEBFC45CA5D2C74" box="[376,523,1019,1046]" pageId="49" pageNumber="50" type="generic">4 specimens</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="922136E09D2D926AFF2EFBA2CD1B2C54" blockId="49.[189,1357,1019,1079]" box="[189,1357,1052,1079]" pageId="49" pageNumber="50">
|
||
<materialsCitation id="22F63CBD9D2D926AFF2EFBA2CB312C54" ID-GBIF-Occurrence="2898878409" box="[189,871,1052,1079]" collectingDate="2015-02-02" collectingDateMax="2015-02-03" collectingDateMin="2015-02-02" collectionCode="NMPC" collectorName="J. Hajek" country="Indonesia" elevation="2050" latitude="-4.06" location="INDONESIA" longLatPrecision="130" longitude="139.03166" municipality="Jayawijaya Distr." pageId="49" pageNumber="50" specimenCount="3" specimenCount-male="3" stateProvince="Papua" typeStatus="holotype">
|
||
<collectingCountry id="EA8976709D2D926AFF2EFBA2C90D2C54" box="[189,347,1052,1078]" name="Indonesia" pageId="49" pageNumber="50">INDONESIA</collectingCountry>
|
||
•
|
||
<specimenCount id="8498FD699D2D926AFEE7FBA2C9EE2C55" box="[372,440,1052,1079]" pageId="49" pageNumber="50" type="male">3 ♂♂</specimenCount>
|
||
; same labels as for holotype;
|
||
<collectionCode id="F48FAE259D2D926AFC87FBA2CB312C54" box="[788,871,1052,1078]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="49" pageNumber="50">NMPC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="22F63CBD9D2D926AFC13FBA2CD1F2C54" ID-GBIF-Occurrence="2898878461" box="[896,1353,1052,1079]" collectingDate="2015-02-02" collectingDateMax="2015-02-03" collectingDateMin="2015-02-02" collectionCode="DTC" collectorName="J. Hajek" country="Indonesia" elevation="2050" latitude="-4.06" location="Baliem valley" longLatPrecision="130" longitude="139.03166" municipality="Jayawijaya Distr." pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Papua" typeStatus="holotype">
|
||
<specimenCount id="8498FD699D2D926AFC13FBA2CBF82C55" box="[896,942,1052,1079]" pageId="49" pageNumber="50" type="male">1 ♂</specimenCount>
|
||
; same labels as for holotype;
|
||
<collectionCode id="F48FAE259D2D926AFA98FBA2CD1F2C54" box="[1291,1353,1052,1078]" collectionName="Dmitrijs Telnovs" pageId="49" pageNumber="50">DTC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D2D9269FF2EFBE5C9FD2CB6" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
|
||
<paragraph id="922136E09D2D926AFF2EFBE5C9182C17" blockId="49.[189,334,1115,1141]" box="[189,334,1115,1141]" pageId="49" pageNumber="50">
|
||
<heading id="C969818C9D2D926AFF2EFBE5C9182C17" bold="true" box="[189,334,1115,1141]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
|
||
<emphasis id="A0EAEAF29D2D926AFF2EFBE5C9182C17" bold="true" box="[189,334,1115,1141]" pageId="49" pageNumber="50">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D2D926AFF2EFB37CACD2C87" blockId="49.[189,1399,1160,1254]" pageId="49" pageNumber="50">
|
||
MEASUREMENTS.
|
||
<typeStatus id="4D2588429D2D926AFEEFFB36C9BA2CC0" box="[380,492,1160,1186]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
, total body length
|
||
<quantity id="55669B059D2D926AFD54FB37CB652CC1" box="[711,819,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="49" pageNumber="50" unit="mm" value="7.1">7.10 mm</quantity>
|
||
. Head
|
||
<quantity id="55669B059D2D926AFC16FB37CBB82CC1" box="[901,1006,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="49" pageNumber="50" unit="mm" value="1.3">1.30 mm</quantity>
|
||
long, across eyes
|
||
<quantity id="55669B059D2D926AFB55FB36CD792CC1" box="[1222,1327,1160,1187]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="49" pageNumber="50" unit="mm" value="1.63">1.63 mm</quantity>
|
||
wide. Pronotum
|
||
<quantity id="55669B059D2D926AFEA4FB14C9C92CA6" box="[311,415,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="mm" value="1.6">1.60 mm</quantity>
|
||
long, maximum width
|
||
<quantity id="55669B059D2D926AFD3EFB14CB412CA6" box="[685,791,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" pageId="49" pageNumber="50" unit="mm" value="2.67">2.67 mm</quantity>
|
||
. Elytral length
|
||
<quantity id="55669B059D2D926AFC54FB14CC642CA6" box="[967,1074,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="49" pageNumber="50" unit="mm" value="4.2">4.20 mm</quantity>
|
||
, maximum combined width
|
||
<quantity id="55669B059D2D926AFF2EFB75C97F2C84" box="[189,297,1227,1254]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="49" pageNumber="50" unit="mm" value="3.3">3.30 mm</quantity>
|
||
.
|
||
<typeStatus id="4D2588429D2D926AFEA7FB72C9F02C84" box="[308,422,1228,1254]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
|
||
<quantity id="55669B059D2D926AFE3EFB75CA0E2C84" box="[429,600,1227,1254]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="7.9" metricValueMin="7.3" pageId="49" pageNumber="50" unit="mm" value="7.6" valueMax="7.9" valueMin="7.3">7.30–7.90 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="922136E09D2D9269FF2EFAB3CA402C00" blockId="49.[189,1399,1293,2025]" lastBlockId="50.[189,1399,265,1123]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
|
||
Dorsum and venter uniformly black-brown with reddish brown labroclypeus, mouthparts, antennae, legs and most of venter. Head transverse, glossy dorsally and ventrally, somewhat convex between eyes and vague impressed on labroclypeus anterior to each compound eye. Compound eye large, occupying about half head side. Male labroclypeus (
|
||
<figureCitation id="0AA52A659D2D926AFDF0FAC9CA9F2DF3" box="[611,713,1399,1425]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="49" pageNumber="50">Fig. 121</figureCitation>
|
||
) very shallowly emarginate anteriorly, its lateral margins curved in dorsal and sinuous in lateral view, anterolateral angles weakly protruding, obtuse and rounded in dorsal view, bent up at ~80–90° to frons in lateral view. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Dorsal punctures circular to ovoid, small and shallow on labroclypeus, larger and moderately deep on frons. Intervening spaces glossy and glabrous, generally larger than punctures. Inconspicuous very short appressed seta rises from anterior margin of each puncture, surpassing or not length of corresponding puncture. Few much longer suberect setae along interior surface of either eye. Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 trapezoid, transverse. Pronotum glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles protruding anteriad and mesal portion nearly straight. Basal margin of pronotum slightly sinuous. Lateral margin of pronotum broadly rounded (
|
||
<figureCitation id="0AA52A659D2D926AFBF6F8A0CC862F5B" box="[1125,1232,1822,1849]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="49" pageNumber="50">Fig. 215</figureCitation>
|
||
). Crenulae of lateral margin poorly defined and rather short; short suberect and posteriad directed seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Punctures of pronotal disc ovoid, moderately deep, variably large; inner margin with delicate membrane in part covered by microscopical velvety pubescence and partly to completely encircles puncture. Intervening spaces glossy and glabrous, variably large but generally as large as to larger than punctures. Setation as on head but setae not surpassing length of corresponding punctures. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Hypomeron slightly curved and very long setose on anterior margin which is flange-like protruding. Antennal pocket deep. Median anterior process of prosternum long brushy setose, broad, strongly raised. Scutellar shield rounded apically. Elytra slightly widened in posterior half, maximum width across midlength, glossy dorsally, with distinct humeri. Vague tracks of two inconspicuous, very flat glabrous longitudinal carinae on disc of each elytron. Punctures of elytral disc ovoid, somewhat denser than those on pronotum, irregular but becoming arranged in irregular rows in apical third (
|
||
<figureCitation id="0AA52A659D2E9269FCCBFDAFCBE92A4E" box="[856,959,529,556]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="50" pageNumber="51">Fig. 305</figureCitation>
|
||
). Inner margin of each puncture with delicate membrane, in some punctures in part covered by microscopical velvety pubescence and partly encircles the puncture. Setation inconspicuous and short, appressed; each seta rises from anterior margin of corresponding puncture, not or hardly surpassing length of puncture. Epipleuron all along with very short appressed setae. Male pygidium flattened dorsally, sparsely circular to oblongo-punctate, opaque microreticulate on intervening spaces, with sparse moderately long suberect clavate setae in distal part and short appressed ones not surpassing length of corresponding punctures in anterior part of pygidium (
|
||
<figureCitation id="0AA52A659D2E9269FF55FCA4C9672B56" box="[198,305,794,820]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="50" pageNumber="51">Fig. 472</figureCitation>
|
||
). Venter covered with sparse small shallow punctures, each provided with very short seta. Third visible abdominal sternite with small median triangular hump on posterior margin (
|
||
<figureCitation id="0AA52A659D2E9269FB56FCFECD3C2B38" box="[1221,1386,832,858]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="50" pageNumber="51">Figs 431–432</figureCitation>
|
||
). Legs long and slender. Male protibia nearly straight on external margin (slightly curved), delicately crenulate in basal two-thirds, with almost complete obtuse dorsal carina. Two obtuse distal teeth on male protibia, distal more acute than inconspicuous broad basal one (Fig. 378). Protibial terminal spur nearly straight, acute. Male metatibial terminal spurs paired, long, acute, lower (longer) one slightly curved. Underside of male protarsomeres 1 and 2 each with large acute and apically slightly hooked distal denticle (Fig. 379). Male tarsal claws with large pulvilli. Spiculum gastrale as in Fig. 544. Male aedeagus as in
|
||
<figureCitation id="0AA52A659D2E9269FEFDFBF6CA442C00" box="[366,530,1096,1122]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="50" pageNumber="51">Figs 617–619</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="922136E09D2E9269FF2EFB34C9E62CC6" blockId="50.[189,432,1162,1188]" box="[189,432,1162,1188]" pageId="50" pageNumber="51">
|
||
<heading id="C969818C9D2E9269FF2EFB34C9E62CC6" bold="true" box="[189,432,1162,1188]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="A0EAEAF29D2E9269FF2EFB34C9E62CC6" bold="true" box="[189,432,1162,1188]" pageId="50" pageNumber="51">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D2E9269FF2EFB04C9FD2CB6" blockId="50.[189,427,1210,1236]" box="[189,427,1210,1236]" pageId="50" pageNumber="51">Female is unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D2E9269FF2EFB42CBD72D27" box="[189,897,1276,1350]" pageId="50" pageNumber="51" type="biology_ecology">
|
||
<paragraph id="922136E09D2E9269FF2EFB42C9762D74" blockId="50.[189,288,1276,1302]" box="[189,288,1276,1302]" pageId="50" pageNumber="51">
|
||
<heading id="C969818C9D2E9269FF2EFB42C9762D74" bold="true" box="[189,288,1276,1302]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="A0EAEAF29D2E9269FF2EFB42C9762D74" bold="true" box="[189,288,1276,1302]" pageId="50" pageNumber="51">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D2E9269FF2EFA95CBD72D27" blockId="50.[189,897,1323,1350]" box="[189,897,1323,1350]" pageId="50" pageNumber="51">
|
||
Occurs in mid-montane rainforests at about
|
||
<quantity id="55669B059D2E9269FD52FA95CB4C2D24" box="[705,794,1323,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="50" pageNumber="51" unit="m" value="2050.0">2050 m</quantity>
|
||
altitude.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D2E9269FF2EFAD3CC382DD5" box="[189,1134,1389,1463]" pageId="50" pageNumber="51" type="distribution">
|
||
<paragraph id="922136E09D2E9269FF2EFAD3C9002DE5" blockId="50.[189,342,1389,1415]" box="[189,342,1389,1415]" pageId="50" pageNumber="51">
|
||
<heading id="C969818C9D2E9269FF2EFAD3C9002DE5" bold="true" box="[189,342,1389,1415]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="A0EAEAF29D2E9269FF2EFAD3C9002DE5" bold="true" box="[189,342,1389,1415]" pageId="50" pageNumber="51">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D2E9269FF2EFA23CC382DD5" blockId="50.[189,1134,1437,1463]" box="[189,1134,1437,1463]" pageId="50" pageNumber="51">
|
||
Hitherto only known from the Baliem Valley, Central Cordillera of New
|
||
<collectingCountry id="EA8976709D2E9269FB81FA23CC3D2DD5" box="[1042,1131,1437,1463]" name="Guinea-Bissau" pageId="50" pageNumber="51">Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |