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<document id="6E10DB4B7CD6CA736FEAE9C63719B0D9" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D209266FDEFFBA4CAE12DA3" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius konjo Telnov 2020, sp. nov." docType="treatment" docUuid="EEEEC581-5447-42AA-9CAC-144395EB5C9C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="62" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="61" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="739E51A76D6E2A0699F0D52FCE92E8AE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9F301FD8D3C5E226849466F3BCFD2383">
<mods:title id="EB364751506923E10DBE685A18A28A66">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="61CDDE9E03F3808D9C939C104613283E" type="personal">
<mods:role id="56C0793F5DA2B9F6458DB2FA9CFBFD93">
<mods:roleTerm id="D0DFA998A3DD18C84C677443BA222CEB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA7489C8EC7492FD1E3C93C7782014B4">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="5FE116D15BCCB46A0199342E7D6717E1">text</mods:typeOfResource>
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<mods:titleInfo id="46ADF6C3E7FC16542A4C3A6681F6B5B9">
<mods:title id="A5A24FBFB68026475FD7A732AF9CA3AD">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E6162761248B84D1ACCB02C7AC2F7EA5">
<mods:date id="F7FE86788134E553CBDFA852251A90BC">2020</mods:date>
<mods:detail id="BA2A74BD404593F49400EDB0773D9C4A" type="pubDate">
<mods:number id="8215CBF5E94A6BE3C13E5B5C38042221">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="AF35385A7340894115EB8C2306B4591E" type="volume">
<mods:number id="D205E9E5D758BB0B56E0AE3E7BDF8E6D">721</mods:number>
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<mods:classification id="7A8E9CCE495D57E5E10EDE4A2207A0F3">journal article</mods:classification>
<mods:identifier id="1ACF20A20DF39EA9B0084BC5854CD09E" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="BD1CE11302ADFD8430EA04F1FCFB5FE6" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="AD0AD890DA641ABDC929FD3BBAC545B8" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
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<mods:identifier id="82D2964FAB40941C415B4D5C26297C65" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D209266FDEFFBA4CAE12DA3" ID-DOI="http://doi.org/10.5281/zenodo.4344313" ID-GBIF-Taxon="168648131" ID-Zenodo-Dep="4344313" LSID="urn:lsid:zoobank.org:act:EEEEC581-5447-42AA-9CAC-144395EB5C9C" httpUri="http://treatment.plazi.org/id/1A3787F69D209266FDEFFBA4CAE12DA3" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<subSubSection id="DA84656B9D209267FDEFFBA4CBEE2C57" box="[636,952,1050,1077]" pageId="60" pageNumber="61" type="nomenclature">
<paragraph id="922136E09D209267FDEFFBA4CBEE2C57" blockId="60.[360,1227,1050,1148]" box="[636,952,1050,1077]" pageId="60" pageNumber="61">
<heading id="C969818C9D209267FDEFFBA4CBEE2C57" box="[636,952,1050,1077]" centered="true" fontSize="11" level="2" pageId="60" pageNumber="61" reason="2">
<taxonomicName id="559E4D639D209267FDEFFBA4CB032C57" ID-CoL="7X3WW" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[636,853,1050,1077]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="konjo" status="sp. nov.">
<emphasis id="A0EAEAF29D209267FDEFFBA4CB032C57" bold="true" box="[636,853,1050,1077]" italics="true" pageId="60" pageNumber="61">Maechidius konjo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D209267FCCEFBA5CBEE2C57" box="[861,952,1051,1077]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D209267FEFBFB80CC442C1E" pageId="60" pageNumber="61" type="description">
<paragraph id="922136E09D209267FEFBFB80CC9D2C3B" blockId="60.[360,1227,1050,1148]" box="[360,1227,1086,1113]" pageId="60" pageNumber="61">
<uri id="E60F3AE29D209267FEFBFB80CC9D2C3B" box="[360,1227,1086,1113]" pageId="60" pageNumber="61">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D209267FD16FB80CC9D2C3B" box="[645,1227,1086,1113]" pageId="60" pageNumber="61">EEEEC581-5447-42AA-9CAC-144395EB5C9C</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D209267FDB2FBDCCC442C1E" blockId="60.[360,1227,1050,1148]" box="[545,1042,1122,1148]" pageId="60" pageNumber="61">
<figureCitation id="0AA52A659D209267FDB2FBDCCA2D2C1E" box="[545,635,1122,1148]" captionStart="Figs 46–49" captionStartId="56.[189,241,1892,1918]" captionTargetBox="[364,1221,267,1852]" captionTargetId="figure-64@56.[361,1224,254,1856]" captionTargetPageId="56" captionText="Figs 46–49. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 46. M.interruptocarinulatusHeller, 1914,holotype, ♀ (SNSD).47. M.jobiensis Moser, 1920,lectotype, ♂ (ZMHB). 48. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 49. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122146" httpUri="https://zenodo.org/record/4122146/files/figure.png" pageId="60" pageNumber="61">Figs 49</figureCitation>
,
<figureCitation id="0AA52A659D209267FD14FBDCCAE32C1E" box="[647,693,1122,1148]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="60" pageNumber="61">131</figureCitation>
,
<figureCitation id="0AA52A659D209267FD52FBDCCAA62C1E" box="[705,752,1122,1148]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="60" pageNumber="61">222</figureCitation>
,
<figureCitation id="0AA52A659D209267FD6FFBDCCB7D2C1E" box="[764,811,1122,1148]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="60" pageNumber="61">313</figureCitation>
, 386, 480, 629–631
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D209267FF2EFB1ACB0F2D57" pageId="60" pageNumber="61" type="diagnosis">
<paragraph id="922136E09D209267FF2EFB1AC99F2CDC" blockId="60.[189,457,1188,1214]" box="[189,457,1188,1214]" pageId="60" pageNumber="61">
<heading id="C969818C9D209267FF2EFB1AC99F2CDC" bold="true" box="[189,457,1188,1214]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="A0EAEAF29D209267FF2EFB1AC99F2CDC" bold="true" box="[189,457,1188,1214]" pageId="60" pageNumber="61">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D209267FF2EFB6DCB0F2D57" blockId="60.[189,1399,1235,1333]" pageId="60" pageNumber="61">
This new species is undoubtedly very close to other
<collectingRegion id="505AF8029D209267FCBFFB6DCBC02C8F" box="[812,918,1235,1261]" country="Indonesia" name="Sulawesi Utara" pageId="60" pageNumber="61">Sulawesi</collectingRegion>
congeners, primarily in the shape of the aedeagus in combination with the V-shaped emarginate male labroclypeus and the lateral margin of the pronotum being somewhat obtuse angulate postmedially.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D209267FF2EFAE3CC452DA8" pageId="60" pageNumber="61" type="etymology">
<paragraph id="922136E09D209267FF2EFAE3C9122D15" blockId="60.[189,324,1373,1399]" box="[189,324,1373,1399]" pageId="60" pageNumber="61">
<heading id="C969818C9D209267FF2EFAE3C9122D15" bold="true" box="[189,324,1373,1399]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="A0EAEAF29D209267FF2EFAE3C9122D15" bold="true" box="[189,324,1373,1399]" pageId="60" pageNumber="61">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D209267FF2EFA32CC452DA8" blockId="60.[189,1398,1420,1483]" pageId="60" pageNumber="61">
The new species is named after Konjo, a group of
<collectingRegion id="505AF8029D209267FC99FA32CB962DC4" box="[778,960,1420,1446]" country="Indonesia" name="Sulawesi Selatan" pageId="60" pageNumber="61">South Sulawesi</collectingRegion>
native languages (Coastal Konjo and
<collectingRegion id="505AF8029D209267FF2EFA0EC97C2DA8" box="[189,298,1456,1482]" country="United Kingdom" name="Highland" pageId="60" pageNumber="61">Highland</collectingRegion>
Konjo) spoken in the area where it occurs. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D209267FF2EFA4CCB0F2F52" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="922136E09D209267FF2EFA4CC93A2E69" blockId="60.[189,364,1521,1548]" box="[189,364,1521,1548]" pageId="60" pageNumber="61">
<heading id="C969818C9D209267FF2EFA4CC93A2E69" bold="true" box="[189,364,1521,1548]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="A0EAEAF29D209267FF2EFA4CC93A2E69" bold="true" box="[189,364,1521,1548]" pageId="60" pageNumber="61">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D209267FF79F99FC9C82EA8" ID-GBIF-Occurrence="2898878546" collectingDate="2015-02-11" collectingDateMax="2015-02-13" collectingDateMin="2015-02-11" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Sumpich" country="Indonesia" county="Gowa Distr." elevation="1520" latitude="-5.2566667" location="Lembanna Base" longLatPrecision="130" longitude="119.90833" municipality="Gn. Bawakaraeng Area" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Sulawesi" typeStatus="holotype">
<paragraph id="922136E09D209267FF79F99FC90A2E59" blockId="60.[189,1399,1569,1738]" box="[234,348,1569,1595]" pageId="60" pageNumber="61">
<emphasis id="A0EAEAF29D209267FF79F99FC90A2E59" bold="true" box="[234,348,1569,1595]" pageId="60" pageNumber="61">
<typeStatus id="4D2588429D209267FF79F99FC90A2E59" box="[234,348,1569,1595]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D209267FF2EF9FBC9C82EA8" blockId="60.[189,1399,1569,1738]" pageId="60" pageNumber="61">
<collectingCountry id="EA8976709D209267FF2EF9FBC90D2E3D" box="[189,347,1605,1631]" name="Indonesia" pageId="60" pageNumber="61">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D209267FEE0F9FBC9DD2E3D" box="[371,395,1605,1631]" pageId="60" pageNumber="61" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D209267FE30F9FBCA1F2E3D" box="[419,585,1605,1631]" name="Indonesia" pageId="60" pageNumber="61">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D209267FDF8F9FBCA8C2E3D" box="[619,730,1605,1631]" country="Indonesia" name="Sulawesi Utara" pageId="60" pageNumber="61">Sulawesi</collectingRegion>
:
<collectingCounty id="7B404E6C9D209267FD77F9FBCB202E3D" box="[740,886,1605,1631]" pageId="60" pageNumber="61">Gowa Distr.</collectingCounty>
;
<locationDeviation id="83932FC29D209267FC12F9FACC022E3D" box="[897,1108,1604,1631]" location="Malino" pageId="60" pageNumber="61">
<quantity id="55669B059D209267FC12F9FACBEA2E3D" box="[897,956,1604,1631]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="60" pageNumber="61" unit="km" value="6.0">6 km</quantity>
E of Malino
</locationDeviation>
,
<collectingMunicipality id="7245AC9A9D209267FBCCF9FBCD242E3D" box="[1119,1394,1605,1631]" pageId="60" pageNumber="61">Gn. Bawakaraeng Area</collectingMunicipality>
, border gardens and mixed forest (dominant
<taxonomicName id="559E4D639D209267FD40F9D7CB402EE0" box="[723,790,1641,1666]" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="A0EAEAF29D209267FD40F9D7CB402EE0" box="[723,790,1641,1666]" italics="true" pageId="60" pageNumber="61">Pinus</emphasis>
</taxonomicName>
) nr
<location id="9741603B9D209267FCDEF9D6CC5A2EE1" LSID="urn:lsid:plazi:treatment:1A3787F69D209266FDEFFBA4CAE12DA3:9741603B9D209267FCDEF9D6CC5A2EE1" box="[845,1036,1640,1667]" country="Indonesia" county="Gowa Distr." latitude="-5.2566667" longLatPrecision="130" longitude="119.90833" municipality="Gn. Bawakaraeng Area" name="Lembanna Base" pageId="60" pageNumber="61" stateProvince="Sulawesi">Lembanna Base</location>
camp
<geoCoordinate id="F7AA50279D209267FBF2F9D6CC8C2EE1" box="[1121,1242,1640,1667]" degrees="05" direction="south" minutes="15.4" orientation="latitude" pageId="60" pageNumber="61" precision="92" value="-5.2566667">05°15.4’S</geoCoordinate>
,
<geoCoordinate id="F7AA50279D209267FB7AF9D6CD242EE1" box="[1257,1394,1640,1667]" degrees="119" direction="east" minutes="54.5" orientation="longitude" pageId="60" pageNumber="61" precision="92" value="119.90833">119°54.5’E</geoCoordinate>
,
<quantity id="55669B059D209267FF2EF932C9762EC5" box="[189,288,1676,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="60" pageNumber="61" unit="m" value="1520.0">
<elevation id="19B3D1D39D209267FF2EF932C9762EC5" box="[189,288,1676,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="60" pageNumber="61" unit="m" value="1520.0">1520 m</elevation>
</quantity>
;
<collectorName id="3F6B53369D209267FEA0F932C9DA2EC4" box="[307,396,1676,1702]" pageId="60" pageNumber="61">J.Hájek</collectorName>
&amp;
<collectorName id="3F6B53369D209267FE53F933CA6C2ECB" box="[448,570,1677,1705]" pageId="60" pageNumber="61">J.Šumpich</collectorName>
leg.
<date id="E62010209D209267FD10F932CB762EC5" box="[643,800,1676,1703]" pageId="60" pageNumber="61" value="2015-02-11" valueMax="2015-02-13" valueMin="2015-02-11">
<collectingDate id="F664E9C89D209267FD10F932CB762EC5" box="[643,800,1676,1703]" pageId="60" pageNumber="61" value="2015-02-11" valueMax="2015-02-13" valueMin="2015-02-11">11-13.ii.2015</collectingDate>
</date>
// coll. general National Museum Prague, Czech Republic”;
<collectionCode id="F48FAE259D209267FED0F90EC9CC2EA8" box="[323,410,1712,1738]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="60" pageNumber="61">NMPC</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D209267FF79F94CC90A2F6E" blockId="60.[189,857,1778,1840]" box="[234,348,1778,1804]" pageId="60" pageNumber="61">
<emphasis id="A0EAEAF29D209267FF79F94CC90A2F6E" bold="true" box="[234,348,1778,1804]" pageId="60" pageNumber="61">
<typeStatus id="4D2588429D209267FF79F94CC90A2F6E" box="[234,348,1778,1804]" pageId="60" pageNumber="61" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D209267FF2EF8A8CB0F2F52" blockId="60.[189,857,1778,1840]" box="[189,857,1814,1840]" pageId="60" pageNumber="61">
<materialsCitation id="22F63CBD9D209267FF2EF8A8CB032F52" ID-GBIF-Occurrence="2898878535" box="[189,853,1814,1840]" collectingDate="2015-02-11" collectingDateMax="2015-02-13" collectingDateMin="2015-02-11" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Sumpich" country="Indonesia" county="Gowa Distr." elevation="1520" latitude="-5.2566667" location="INDONESIA" longLatPrecision="130" longitude="119.90833" municipality="Gn. Bawakaraeng Area" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi" typeStatus="paratype">
<collectingCountry id="EA8976709D209267FF2EF8A8C90D2F52" box="[189,347,1814,1840]" name="Indonesia" pageId="60" pageNumber="61">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D209267FEE7F8A8C9F42F52" box="[372,418,1814,1840]" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D209267FD6DF8A8CB032F52" box="[766,853,1814,1840]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="60" pageNumber="61">NMPC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D209266FF2EF8E9C9FD2CB5" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="description">
<paragraph id="922136E09D209267FF2EF8E9C9182F13" blockId="60.[189,334,1879,1905]" box="[189,334,1879,1905]" pageId="60" pageNumber="61">
<heading id="C969818C9D209267FF2EF8E9C9182F13" bold="true" box="[189,334,1879,1905]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="A0EAEAF29D209267FF2EF8E9C9182F13" bold="true" box="[189,334,1879,1905]" pageId="60" pageNumber="61">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D209267FF2EF839CACE2F8A" blockId="60.[189,1399,1927,2025]" pageId="60" pageNumber="61">
MEASUREMENTS. Male
<typeStatus id="4D2588429D209267FE52F839CA7D2FC3" box="[449,555,1927,1953]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D209267FC9AF839CB202FC3" box="[777,886,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="60" pageNumber="61" unit="mm" value="7.3">7.30 mm</quantity>
. Head
<quantity id="55669B059D209267FC5AF839CC652FC3" box="[969,1075,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="60" pageNumber="61" unit="mm" value="1.4">1.40 mm</quantity>
long, across eyes
<quantity id="55669B059D209267FA9EF839CD212FC3" box="[1293,1399,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="60" pageNumber="61" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D209267FEEDF815C9B02FA7" box="[382,486,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="60" pageNumber="61" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D209267FD66F814CB092FA7" box="[757,863,1962,1989]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="60" pageNumber="61" unit="mm" value="2.55">2.55 mm</quantity>
. Elytral length
<quantity id="55669B059D209267FB83F815CC2D2FA7" box="[1040,1147,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="60" pageNumber="61" unit="mm" value="4.2">4.20 mm</quantity>
, maximum combined width
<quantity id="55669B059D209267FE94F870C9252F8B" box="[263,371,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="60" pageNumber="61" unit="mm" value="3.4">3.40 mm</quantity>
.
<typeStatus id="4D2588429D209267FEEDF871C9B32F8B" box="[382,485,1999,2025]" pageId="60" pageNumber="61" type="paratype">Paratype</typeStatus>
<quantity id="55669B059D209267FE7FF870CA032F8B" box="[492,597,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="60" pageNumber="61" unit="mm" value="6.9">6.90 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D219266FF2EFEB7C8A62C00" blockId="61.[189,1399,265,1122]" pageId="61" pageNumber="62">
Dorsum and venter uniformly black to black-brown, labroclypeus, antennae and legs castaneous brown. Male labroclypeus (
<figureCitation id="0AA52A659D219266FE3FFE91CA44292B" box="[428,530,303,329]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="61" pageNumber="62">Fig. 131</figureCitation>
) shallowly emarginate on anterior margin, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, strongly protruding. Frons convex dorsally. Compound eye moderately large, less than half head length. Head punctures ovoid to irregularly hexagonal, dense. Intervening spaces glossy, generally smaller than punctures. Head setae appressed to suberect, inconspicuous, sparse; each seta rises from anterior margin of corresponding puncture, surpassing its length. Pronotum broadly emarginate on anterior margin, basal margin broadly rounded. Lateral margin of pronotum broadly rounded, with very obtuse postmedian angulation, shallowly emarginate prebasally, crenulate all along (
<figureCitation id="0AA52A659D219266FCA0FD89CBCA2A33" box="[819,924,567,593]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="61" pageNumber="62">Fig. 222</figureCitation>
). Long erect seta present between every two crenulae. Pronotal punctures elongate ovoid, deep and dense. Intervening spaces glossy, smaller than punctures, in part wrinkled. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Setae generally as on head, but with numerous minute ones not surpassing midlength of corresponding punctures. Elytron with tracks of two flat glabrous longitudinal carinae, sutural carina not indicated. Elytral punctures irregularly ovoid, deep and dense. Intervening spaces glossy, generally larger than punctures (
<figureCitation id="0AA52A659D219266FCFAFCA7CB982B56" box="[873,974,793,820]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="61" pageNumber="62">Fig. 313</figureCitation>
). Elytral setae appressed to suberect. Minute setae rise from anterior margin of corresponding punctures, not surpassing their midlength. Longer suberect setae arranged in 5–6 irregular longitudinal rows on each elytron. Male pygidium with moderately large sparse annular punctures, intervening spaces glossy, variably large (
<figureCitation id="0AA52A659D219266FB0DFC35CD532BC7" box="[1182,1285,907,933]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="61" pageNumber="62">Fig. 480</figureCitation>
). Setae of pygidium rise from anterior margin of corresponding punctures, generally suberect to erect and longer than corresponding punctures. Several minute setae not surpassing length of corresponding punctures located along anterior margin of pygidium. Male protibia widened distally, with two obtuse distal teeth on external margin (Fig. 386). Male protibial terminal spur strongly curved. Aedeagus as in
<figureCitation id="0AA52A659D219266FA92FB9CC8BA2C00" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="61" pageNumber="62">Figs 629– 631</figureCitation>
.
</paragraph>
<paragraph id="922136E09D219266FF2EFB35C9E62CC7" blockId="61.[189,432,1163,1189]" box="[189,432,1163,1189]" pageId="61" pageNumber="62">
<heading id="C969818C9D219266FF2EFB35C9E62CC7" bold="true" box="[189,432,1163,1189]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="A0EAEAF29D219266FF2EFB35C9E62CC7" bold="true" box="[189,432,1163,1189]" pageId="61" pageNumber="62">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D219266FF2EFB03C9FD2CB5" blockId="61.[189,427,1213,1239]" box="[189,427,1213,1239]" pageId="61" pageNumber="62">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D219266FF2EFABFCD1F2D2E" box="[189,1353,1281,1357]" pageId="61" pageNumber="62" type="biology_ecology">
<paragraph id="922136E09D219266FF2EFABFC9762D79" blockId="61.[189,288,1281,1307]" box="[189,288,1281,1307]" pageId="61" pageNumber="62">
<heading id="C969818C9D219266FF2EFABFC9762D79" bold="true" box="[189,288,1281,1307]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="A0EAEAF29D219266FF2EFABFC9762D79" bold="true" box="[189,288,1281,1307]" pageId="61" pageNumber="62">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D219266FF2EFA8CCD1F2D2E" blockId="61.[189,1353,1330,1357]" box="[189,1353,1330,1357]" pageId="61" pageNumber="62">
Occurs in lower to mid-montane mixed rainforests (altitude ~
<quantity id="55669B059D219266FC1EFA8CCBB12D2E" box="[909,999,1330,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="62" unit="m" value="1500.0">1500 m</quantity>
a.s.l.) dominated by
<taxonomicName id="559E4D639D219266FB4DFA8DCD772D2E" box="[1246,1313,1331,1356]" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="genus">
<emphasis id="A0EAEAF29D219266FB4DFA8DCD772D2E" box="[1246,1313,1331,1356]" italics="true" pageId="61" pageNumber="62">Pinus</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D219266FF2EFAC8CAE12DA3" box="[189,695,1398,1473]" pageId="61" pageNumber="62" type="distribution">
<paragraph id="922136E09D219266FF2EFAC8C9002DF2" blockId="61.[189,342,1398,1424]" box="[189,342,1398,1424]" pageId="61" pageNumber="62">
<heading id="C969818C9D219266FF2EFAC8C9002DF2" bold="true" box="[189,342,1398,1424]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="A0EAEAF29D219266FF2EFAC8C9002DF2" bold="true" box="[189,342,1398,1424]" pageId="61" pageNumber="62">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D219266FF2EFA19CAE12DA3" blockId="61.[189,695,1447,1473]" box="[189,695,1447,1473]" pageId="61" pageNumber="62">
Hitherto only known from
<collectingRegion id="505AF8029D219266FE6AFA19CAE52DA3" box="[505,691,1447,1473]" country="Indonesia" name="Sulawesi Selatan" pageId="61" pageNumber="62">South Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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