223 lines
32 KiB
XML
223 lines
32 KiB
XML
<document id="19A5F0A7914AC3F3599EEAE075507E98" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFEEE312FF78FC43FD9283D0" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Harpes prescheri Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="51" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="50" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="C0500C76FD96F151004DFBFAEE480842">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="157089E3402382F92EA030EADE6C2BC7">Johnson, Robert G.</mods:namePart>
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<mods:typeOfResource id="44E1328B91AD04EC495638FA2D663015">text</mods:typeOfResource>
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<mods:title id="A34F71B5FABD14F8F4BD63335E91128E">Zootaxa</mods:title>
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<mods:part id="7C21E79168CE8E3184FA14DDE3F1FD19">
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<mods:date id="6B78B3A3B7B3634B39EE3B62572E9BC9">2024</mods:date>
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<mods:number id="EA44B9022CFFB120377DC83D9164B755">2024-05-15</mods:number>
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<mods:number id="ED42A3AFD0107F3CCB448941CE7D7A5D">5450</mods:number>
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<mods:url id="1247CC367B667B3870578D722829838B">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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<mods:classification id="3BA39A77C2B9337F08335B27910D6D98">journal article</mods:classification>
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<mods:identifier id="58ADD8270C009CCFEA254257EADEB01D" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="456B845AA3E89B7E2B2EC3C10A2AAA8E" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03FD8227FFEEE312FF78FC43FD9283D0" ID-DOI="http://doi.org/10.5281/zenodo.11232551" ID-Zenodo-Dep="11232551" LSID="urn:lsid:plazi:treatment:03FD8227FFEEE312FF78FC43FD9283D0" httpUri="http://treatment.plazi.org/id/03FD8227FFEEE312FF78FC43FD9283D0" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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<subSubSection id="C34E60BAFFEEE311FF78FC43FE048328" box="[151,433,1086,1113]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="8BEB3331FFEEE311FF78FC43FE048328" blockId="49.[151,433,1086,1149]" box="[151,433,1086,1113]" pageId="49" pageNumber="50">
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<heading id="D0A3845DFFEEE311FF78FC43FE048328" bold="true" box="[151,433,1086,1113]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
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<emphasis id="B920EF23FFEEE311FF78FC43FE048328" bold="true" box="[151,433,1086,1113]" pageId="49" pageNumber="50">
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<taxonomicName id="4C5448B2FFEEE311FF78FC43FED28328" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,359,1086,1113]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="prescheri" status="sp. nov.">
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<emphasis id="B920EF23FFEEE311FF78FC43FED28328" bold="true" box="[151,359,1086,1113]" italics="true" pageId="49" pageNumber="50">Harpes prescheri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FFEEE311FE81FC43FE048328" box="[366,433,1087,1113]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEEE311FF78FC1FFEA5830C" box="[151,272,1123,1149]" pageId="49" pageNumber="50" type="description">
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<paragraph id="8BEB3331FFEEE311FF78FC1FFEA5830C" blockId="49.[151,433,1086,1149]" box="[151,272,1123,1149]" pageId="49" pageNumber="50">
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<figureCitation id="136F2FB4FFEEE311FF78FC1FFEA5830C" box="[151,272,1123,1149]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="49" pageNumber="50">Plate 9A–I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEEE311FF78FCD6FDCD8224" pageId="49" pageNumber="50" type="diagnosis">
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<paragraph id="8BEB3331FFEEE311FF78FCD6FDCD8224" blockId="49.[151,1437,1194,1977]" pageId="49" pageNumber="50">
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<emphasis id="B920EF23FFEEE311FF78FCD6FEA783B5" bold="true" box="[151,274,1194,1220]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
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Cephalon widest across eyes and glabella is narrow, cone-shaped, rounded anteriorly with suppressed tubercles straddling its crest. Brim width ratio low at around 0.50, brim slightly concave (sag.), becoming gently convex close to posterior border, alar furrows effaced distally, perforations on brim very fine. Prolongations shorter (exsag.) than cephalon (sag.). Pleurae shortening (tr.) posteriorly over last fifteen thoracic segments to become very short, giving pointed appearance to thorax.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEEE311FF28FD23FAB48208" box="[199,1281,1375,1401]" pageId="49" pageNumber="50" type="etymology">
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<paragraph id="8BEB3331FFEEE311FF28FD23FAB48208" blockId="49.[151,1437,1194,1977]" box="[199,1281,1375,1401]" pageId="49" pageNumber="50">
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<emphasis id="B920EF23FFEEE311FF28FD23FEFB8208" bold="true" box="[199,334,1375,1401]" pageId="49" pageNumber="50">Etymology.</emphasis>
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Named in remembrance of Harald Prescher, a trilobite researcher and a good friend.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEEE311FF28FDFEFA878120" pageId="49" pageNumber="50" type="materials_examined">
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<paragraph id="8BEB3331FFEEE311FF28FDFEFD5C8178" blockId="49.[151,1437,1194,1977]" pageId="49" pageNumber="50">
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<emphasis id="B920EF23FFEEE311FF28FDFEFE4D82EC" bold="true" box="[199,504,1410,1437]" pageId="49" pageNumber="50">Material and occurrence.</emphasis>
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<materialsCitation id="3B3C396CFFEEE311FDEFFDFFFDC88294" collectionCode="NHMUK" elevation="100" pageId="49" pageNumber="50" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FFEEE311FDEFFDFFFDD882EC" box="[512,621,1411,1437]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFEEE311FD97FDFFFD5782EC" box="[632,738,1411,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="49" pageNumber="50" type="Museum">NHMUK</collectionCode>
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It 29244,
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<figureCitation id="136F2FB4FFEEE311FCB7FDFFFC7782EC" box="[856,962,1411,1437]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="49" pageNumber="50">Pl. 9A–E</figureCitation>
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, from a horizon around
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<quantity id="4CAC9ED4FFEEE311FB39FDFFFAAB82EC" box="[1238,1310,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="m" value="100.0">
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||
<elevation id="0079D402FFEEE311FB39FDFFFAAB82EC" box="[1238,1310,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="m" value="100.0">100 m</elevation>
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</quantity>
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upslope of “Couche Rouge” close to the “
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<taxonomicName id="4C5448B2FFEEE311FE1FFDDBFD6D82B0" authority="Couche" authorityName="Couche" box="[496,728,1447,1473]" class="Trilobita" family="Acastidae" genus="Psychopyge" kingdom="Animalia" order="Phacopida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EF23FFEEE311FE1FFDDBFDC382B0" box="[496,630,1447,1473]" italics="true" pageId="49" pageNumber="50">Psychopyge</emphasis>
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Couche
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</taxonomicName>
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”, Er Remlia Formation, Emsian, Jbel Issoumour (
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<figureCitation id="136F2FB4FFEEE311FAE9FDDBFAE882B0" box="[1286,1373,1447,1473]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="49" pageNumber="50">Fig. 2B</figureCitation>
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, Map 7, site 9), disarticulated dorsal exoskeleton
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</materialsCitation>
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||
.
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<materialsCitation id="3B3C396CFFEEE311FD66FDB7FD508178" collectionCode="NHMUK" pageId="49" pageNumber="50" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EF8D93FFEEE311FD66FDB7FD5A8294" box="[649,751,1483,1509]" pageId="49" pageNumber="50" type="paratype">Paratype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFEEE311FD15FDB7FCD18294" box="[762,868,1483,1509]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="49" pageNumber="50" type="Museum">NHMUK</collectionCode>
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It 29243,
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<figureCitation id="136F2FB4FFEEE311FC35FDB7FB828294" box="[986,1079,1483,1509]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="49" pageNumber="50">Pl. 9F–I</figureCitation>
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, from type horizon and locality (
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<figureCitation id="136F2FB4FFEEE311FF70FD93FF428178" box="[159,247,1519,1545]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="49" pageNumber="50">Fig. 2B</figureCitation>
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, Map 7, site 8), cephalic dorsal exoskeleton
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEB3331FFEEE311FF28FE6FFA878120" blockId="49.[151,1437,1194,1977]" pageId="49" pageNumber="50">
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Other material:
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<taxonomicName id="4C5448B2FFEEE311FE99FE6FFD05815C" authority=", NHMUK It" authorityName="NHMUK It" box="[374,688,1555,1581]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="prescheri">
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<emphasis id="B920EF23FFEEE311FE99FE6FFE3F815D" box="[374,394,1555,1580]" italics="true" pageId="49" pageNumber="50">H</emphasis>
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. cf.
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<emphasis id="B920EF23FFEEE311FE55FE6FFD9F815C" box="[442,554,1555,1581]" italics="true" pageId="49" pageNumber="50">prescheri,</emphasis>
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NHMUK It
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</taxonomicName>
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29245,
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<figureCitation id="136F2FB4FFEEE311FCE6FE6FFCDC815C" box="[777,873,1555,1581]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="49" pageNumber="50">Pl. 9J–N</figureCitation>
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from the “
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<taxonomicName id="4C5448B2FFEEE311FC30FE6FFB1A815C" authority="Couche" authorityName="Couche" box="[991,1199,1555,1581]" class="Diplopoda" family="Pyrgodesmidae" genus="Lobopyge" kingdom="Animalia" order="Polydesmida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EF23FFEEE311FC30FE6FFBF8815C" box="[991,1101,1555,1581]" italics="true" pageId="49" pageNumber="50">Lobopyge</emphasis>
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Couche
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</taxonomicName>
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”, El Otfal Formation, Eifelian, Issoumour (see Issoumour section,
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<bibRefCitation id="EFC54EC0FFEEE311FD65FE4BFC648120" author="McKellar, R. C. & Chatterton, B. D. E." box="[650,977,1591,1617]" pageId="49" pageNumber="50" pagination="1 - 109" refId="ref89135" refString="McKellar, R. C. & Chatterton, B. D. E. (2009) Early and Middle Devonian Phacopidae (trilobite) of southern Morocco. Palaeontographica Canadiana, 28, 1 - 109." type="journal article" year="2009">McKellar & Chatterton 2009</bibRefCitation>
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, p. 12) (
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<figureCitation id="136F2FB4FFEEE311FBC2FE4BFB318120" box="[1069,1156,1591,1617]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="49" pageNumber="50">Fig. 2B</figureCitation>
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, Map 7, site 2).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E60BAFFEEE312FF28FE26FD5C8464" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
|
||
<paragraph id="8BEB3331FFEEE311FF28FE26FC9B80C8" blockId="49.[151,1437,1194,1977]" pageId="49" pageNumber="50">
|
||
<emphasis id="B920EF23FFEEE311FF28FE26FEEC8105" bold="true" box="[199,345,1626,1652]" pageId="49" pageNumber="50">Description.</emphasis>
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Cephalon and genicranium narrow, length (sag.) 86% and 76% of width (tr.), respectively. Widest point of cephalon and genicranium of
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<typeStatus id="54EF8D93FFEEE311FDBCFE03FD0081E8" box="[595,693,1663,1689]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
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at midpoint between eye lobes and alae and at posterior border, respectively. Glabella narrow, carinate, and with coarse, barely-inflated tubercles on crest. Width (tr.) of glabella at S1 around 65% of length (sag.) of glabella anterior to S0. S1 sloping posteriorly and curving slightly anteriorly, S2 and S3 very shallow, barely rising out of axial furrow. Axial furrows very gently convex and converging anteriorly at more than
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<superScript id="7C219E79FFEEE311FEC8FF72FEFE806D" attach="left" box="[295,331,1806,1833]" fontSize="6" pageId="49" pageNumber="50">150</superScript>
|
||
to sagittal line. Preglabellar furrow shallow and straight (tr.) in anterior view. Occipital furrow deep and straight (tr.). Occipital ring medially broadened and in lateral view, lower than glabella anterior to S0. Occipital node small and barely inflated. L1 not inflated although alae are. Alae large (tr.) and narrow and reaching over 60% of way across genal area. Alar furrow very shallow and distally effaced. No interalar furrows or ridges. Posterior border short (tr.), under 30% of width of occipital ring (tr.).
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEDE312FF28F8EBFE4B86DC" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
Genal area narrow, width (tr.) around 50% of length (exsag.) and lacking pits and caeca. Preglabellar genal area narrow (sag.), length (sag.) ≈20% of width (tr.). Eye lobes very large, strongly inflated, cone-shaped and extending to axial furrow. Eye location ratio around 0.83. Two fairly small oval eye lenses (
|
||
<figureCitation id="136F2FB4FFEDE312FBB3F8A3FB198788" box="[1116,1196,223,249]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="50" pageNumber="51">Pl. 9D</figureCitation>
|
||
). Genal ridge barely indicated, becoming a little more visible posterolaterally on genal roll. Anterior boss barely inflated and girder weakly kinked. Genal roll gently sloping anteriorly (≈ 45
|
||
<superScript id="7C219E79FFEDE312FCF8F95BFCAA8644" attach="none" box="[791,799,295,309]" fontSize="6" pageId="50" pageNumber="51">o</superScript>
|
||
) and moderately steeply laterally (≈
|
||
<superScript id="7C219E79FFEDE312FB5BF95BFB6C8645" attach="none" box="[1204,1241,294,321]" fontSize="6" pageId="50" pageNumber="51">600</superScript>
|
||
). Perforations on genal roll about same size as those on brim, becoming slightly smaller away from girder. Row of large perforations immediately above girder, and at inner margin of fringe, anterior to glabella, row of perforations a little larger than those on main part of genal roll.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEDE312FF28F9CBFE7A85BC" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
Brim wide, about one third of cephalic length (sag.), sloping moderately (≈
|
||
<superScript id="7C219E79FFEDE312FBD9F9CBFBEC86B5" attach="none" box="[1078,1113,438,465]" fontSize="6" pageId="50" pageNumber="51">170</superScript>
|
||
), gently concave anteriorly, straight laterally and becoming gently convex close to posterior border. Brim width ratio just over 0.50. Brim perforations becoming smaller away from girder to become very fine (standardised diameter of ≈80 μm). Row of larger perforations at proximal and distal edges, no caeca. Rim stout and lacking tubercles on dorsal surface. Marginal band near vertical with dorsal surface overhanging. Prolongations moderately long (exsag.) at 85% of cephalic length (sag.). In dorsal view, internal rim curving smoothly adaxially and concave posteriorly in lateral view. Internal rim wide (tr.), wider than external rim (sag.) and with tubercles on dorsal edge. Genal spine short set slightly obliquely adaxially.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEDE312FF28FAABFD5C8464" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
Thorax little known but inner portion of pleurae shortening (tr.) strongly posteriorly, much more than axial rings. (
|
||
<figureCitation id="136F2FB4FFEDE312FF09FA87FE848464" box="[230,305,763,789]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="50" pageNumber="51">Pl. 9A</figureCitation>
|
||
). Pygidium and hypostome not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFEDE312FF28FB63FD9283D0" pageId="50" pageNumber="51" type="discussion">
|
||
<paragraph id="8BEB3331FFEDE312FF28FB63FD5D842C" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
<emphasis id="B920EF23FFEDE312FF28FB63FE8E8448" bold="true" box="[199,315,799,825]" pageId="50" pageNumber="51">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FFEDE312FEABFB63FDBC8448" box="[324,521,799,825]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FEABFB63FDBC8448" box="[324,521,799,825]" italics="true" pageId="50" pageNumber="51">Harpes prescheri</emphasis>
|
||
</taxonomicName>
|
||
lacks only one of the diagnostic characters for the genus shown in
|
||
<tableCitation id="C6D6068AFFEDE312FAFAFB63FACD8448" box="[1301,1400,798,825]" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="50" pageNumber="51">Table 5a</tableCitation>
|
||
, in having a weak girder kink rather than no girder kink.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEDE312FF28FB1BFC8F8344" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
<taxonomicName id="4C5448B2FFEDE312FF28FB1BFE3F84F0" box="[199,394,871,897]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FF28FB1BFE3F84F0" box="[199,394,871,897]" italics="true" pageId="50" pageNumber="51">Harpes prescheri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5448B2FFEDE312FE2FFB1BFD8184F0" box="[448,564,871,897]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFEDE312FE2FFB1BFD8184F0" box="[448,564,871,897]" italics="true" pageId="50" pageNumber="51">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
are similar but have well defined differences. The cephalon, genicranium and glabella of
|
||
<taxonomicName id="4C5448B2FFEDE312FEF7FBF7FE1184D4" box="[280,420,907,933]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FEF7FBF7FE9984D5" box="[280,300,907,932]" italics="true" pageId="50" pageNumber="51">H</emphasis>
|
||
.
|
||
<emphasis id="B920EF23FFEDE312FED4FBF7FE1184D4" box="[315,420,907,933]" italics="true" pageId="50" pageNumber="51">prescheri</emphasis>
|
||
</taxonomicName>
|
||
are much narrower, the genal roll is more gently-sloping anteriorly and the brim is wider (sag.) and has a brim width ratio of just over 0.50, whereas that of
|
||
<taxonomicName id="4C5448B2FFEDE312FC93FBD3FC4584B8" box="[892,1008,943,969]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFEDE312FC93FBD3FC4584B8" box="[892,1008,943,969]" italics="true" pageId="50" pageNumber="51">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
is 0.76. Also, the dorsal surface of the rim lacks tubercles, and the length (exsag.) of the prolongations expressed as a percentage of cephalic length (sag.) is 82% and 112% for
|
||
<taxonomicName id="4C5448B2FFEDE312FE64FB8BFDA28360" box="[395,535,1015,1041]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FE64FB8BFDA28360" box="[395,535,1015,1041]" italics="true" pageId="50" pageNumber="51">H. prescheri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B920EF23FFEDE312FDA1FB8BFD7C8360" box="[590,713,1015,1041]" italics="true" pageId="50" pageNumber="51">
|
||
<taxonomicName id="4C5448B2FFEDE312FDA1FB8BFD708360" box="[590,709,1015,1041]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lahceni">H. lahceni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
respectively. In lateral view, the profile of the internal rim of
|
||
<taxonomicName id="4C5448B2FFEDE312FA6EFB8BFEB58344" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FA6EFB8BFEB58344" italics="true" pageId="50" pageNumber="51">H. prescheri</emphasis>
|
||
</taxonomicName>
|
||
is concave and not wave-like as it is in
|
||
<taxonomicName id="4C5448B2FFEDE312FD51FC67FC878344" box="[702,818,1051,1077]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFEDE312FD51FC67FC878344" box="[702,818,1051,1077]" italics="true" pageId="50" pageNumber="51">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEDE312FF28FC43FD9283D0" blockId="50.[151,1437,150,1185]" pageId="50" pageNumber="51">
|
||
NHMUK It 29245, designated as
|
||
<taxonomicName id="4C5448B2FFEDE312FDA9FC43FC9A8328" box="[582,815,1087,1113]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FDA9FC43FD2D8329" box="[582,664,1087,1112]" italics="true" pageId="50" pageNumber="51">Harpes</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FFEDE312FD29FC43FC9A8328" box="[710,815,1087,1113]" italics="true" pageId="50" pageNumber="51">prescheri</emphasis>
|
||
</taxonomicName>
|
||
, (
|
||
<figureCitation id="136F2FB4FFEDE312FCA9FC43FC1F8328" box="[838,938,1087,1113]" captionStart="PLATE 9" captionStartId="115.[151,234,1675,1699]" captionTargetBox="[236,1351,181,1651]" captionTargetId="figure-15@115.[236,1351,181,1651]" captionTargetPageId="115" captionText="PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232561" httpUri="https://zenodo.org/record/11232561/files/figure.png" pageId="50" pageNumber="51">Pl.9 J–N</figureCitation>
|
||
) is similar to
|
||
<taxonomicName id="4C5448B2FFEDE312FBA7FC43FB618328" box="[1096,1236,1087,1113]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFEDE312FBA7FC43FB618328" box="[1096,1236,1087,1113]" italics="true" pageId="50" pageNumber="51">H. prescheri</emphasis>
|
||
</taxonomicName>
|
||
in having a steep genal roll anteriorly and a short (sag.) preglabellar field, but differs in having a narrower brim (sag.) and a brim width ratio of 0.73 rather than 0.50.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |