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<document id="C68E4FBD32F04F044C1A831BEA071FF2" ID-CLB-Dataset="56065" ID-DOI="10.5852/ejt.2022.809.1719" ID-GBIF-Dataset="f76de735-01e3-454c-a7f1-bbafd26686f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6419425" ID-ZooBank="A754493E-5466-4479-B515-AABEDDE09D93" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1649273451180" checkinUser="felipe" docAuthor="Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. &amp; Abakumov, Evgeny V." docDate="2022" docId="03FC86665572FF9F500EBA96FC8EF8CC" docLanguage="en" docName="ejt-809_1-71.pdf" docOrigin="European Journal of Taxonomy 809" docSource="http://dx.doi.org/10.5852/ejt.2022.809.1719" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dietomorpha Reymond 1938" docType="treatment" docVersion="12" lastPageNumber="26" masterDocId="FFC5FE1E556AFF865230BB19FFBAFF8B" masterDocTitle="A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes" masterLastPageNumber="71" masterPageNumber="1" pageNumber="25" updateTime="1733334329041" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="B0F0A6CC933E16E0B19138A1E060A447">
<mods:title id="44B01B1B4EA85AF18FDB3FBE61C7FEAF">A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes</mods:title>
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<mods:roleTerm id="3C5594E4EDF77ABA8CBCF05A5A3D6A7B">Author</mods:roleTerm>
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<mods:namePart id="C9385D98DD9B5F2D6FC12E4684364A67">Chigray, Svetlana N.</mods:namePart>
<mods:nameIdentifier id="41154041B1A8080E491C558759BF9C91" type="LSID">5097C803-AA1A-4856-95D7-F11DB747EB74</mods:nameIdentifier>
<mods:affiliation id="24B434E975238AB67FDC5EE574327E98">St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia.</mods:affiliation>
<mods:nameIdentifier id="5CDBE852C830496007B35081C804D7BD" type="email">s.chigray@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="5153A00A525EABBBA62FA3A30999764F" type="personal">
<mods:role id="1B55E872DE57BD8EA79C206FAA89562F">
<mods:roleTerm id="62BB8B24C854AAA5FD7ED4171443EAF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5474DA9379E2E7AD53CDF87CAA50643E">Nabozhenko, Maxim V.</mods:namePart>
<mods:nameIdentifier id="A80175CBA6E077C3655DDE4C6BB54780" type="LSID">A339C181-A65D-4F68-AA3E-01D07DFF25AD</mods:nameIdentifier>
<mods:affiliation id="D91145AFE0CC26CFBB1D2D14C01965E4">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000 Russia. &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia. &amp; Severtsov Institute of ecology and evolution of the Russian Academy of sciences, Leninsky Prospect, 33, Moscow 119071 Russia.</mods:affiliation>
<mods:nameIdentifier id="91809B1FDEB2C1060D1716D41AC82A4B" type="email">nalassus@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="BF3B03DA895D4808AF536F0AA2B076C3" type="personal">
<mods:role id="F8E6A21274AAE693D014EF41FBE2CD87">
<mods:roleTerm id="A33990F9F008EBBBD016559EA4504DAB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C06E26CD418A670FA8A873F812BD1FF8">Chigray, Ivan A.</mods:namePart>
<mods:nameIdentifier id="7244B319F028172F26E3B869EC1A4C9C" type="LSID">4CDA2EFB-5C20-4E11-870E-BDFD7DBB78F7</mods:nameIdentifier>
<mods:affiliation id="FE57AE3D3AF73CB1BE59327330CC96A0">Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, St. Petersburg 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="93F32D6FFC8C8B5DD8E99A273899E65F" type="email">chigray93@bk.ru</mods:nameIdentifier>
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<mods:namePart id="48EF6DCB94B772379AD47C55C5DD4931">Abakumov, Evgeny V.</mods:namePart>
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<mods:affiliation id="15E81FE939E99C9497DB5BC412D37ECE">St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia.</mods:affiliation>
<mods:nameIdentifier id="08E16EE8E222AC6C76C57735336B647E" type="email">e.abakumov@spb.ru</mods:nameIdentifier>
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<mods:title id="206D5B22DAE100CDFB15C0B6EA91E2D7">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="70AF533C03DD67D7358E9CD77761732B">
<mods:date id="0FE491C64A67AB8C3282458BFD1B8EB3">2022</mods:date>
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<mods:number id="888242359B173324CB9F2C1CF7C886CC">2022-04-05</mods:number>
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<mods:number id="B030781652332B8924185BBD40B239FC">809</mods:number>
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<mods:start id="FE7AD50A9905BD1AC783C4B4693209E4">1</mods:start>
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<treatment id="03FC86665572FF9F500EBA96FC8EF8CC" ID-DOI="http://doi.org/10.5281/zenodo.6449866" ID-GBIF-Taxon="194566013" ID-Zenodo-Dep="6449866" LSID="urn:lsid:plazi:treatment:03FC86665572FF9F500EBA96FC8EF8CC" httpUri="http://treatment.plazi.org/id/03FC86665572FF9F500EBA96FC8EF8CC" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" scope_class="Insecta" scope_family="Tenebrionidae" scope_order="Coleoptera" scope_subFamily="Pimeliinae" scope_subOrder="Polyphaga" scope_subTribe="Habrobatina" scope_tribe="Pimeliini">
<subSubSection id="C34F64FB5572FF9E500EBA96FC4FFE22" box="[574,1013,399,426]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BEA37705572FF9E500EBA96FC4FFE22" blockId="24.[574,1013,399,460]" box="[574,1013,399,426]" pageId="24" pageNumber="25">
<heading id="D0A2801C5572FF9E500EBA96FC4FFE22" box="[574,1013,399,426]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
Genus
<taxonomicName id="4C554CF35572FF9E50A1BA96FC4FFE22" authority="Reymond, 1938" authorityName="Reymond" authorityYear="1938" box="[657,1013,399,426]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625572FF9E50A1BA96FC88FE21" bold="true" box="[657,818,399,426]" italics="true" pageId="24" pageNumber="25">Dietomorpha</emphasis>
Reymond, 1938
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5572FF9E5090BAABFC29FE47" box="[672,915,434,460]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BEA37705572FF9E5090BAABFC29FE47" blockId="24.[574,1013,399,460]" box="[672,915,434,460]" pageId="24" pageNumber="25">
<figureCitation id="136E2BF55572FF9E5090BAABFD50FE47" box="[672,746,434,460]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[249,1340,656,1847]" captionTargetId="figure-122@8.[241,1346,643,1855]" captionTargetPageId="8" captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray &amp; Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales." figureDoi="http://doi.org/10.5281/zenodo.6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="24" pageNumber="25">Figs 3</figureCitation>
,
<figureCitation id="136E2BF55572FF9E50C7BAABFCACFE47" box="[759,790,434,460]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="24" pageNumber="25">17</figureCitation>
,
<figureCitation id="136E2BF55572FF9E5113BAABFCEDFE47" box="[803,855,434,460]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="24" pageNumber="25">23C</figureCitation>
,
<figureCitation id="136E2BF55572FF9E5153BAABFC29FE47" box="[867,915,434,460]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="24" pageNumber="25">26C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5572FF9E528DBAEDFB0EFDF8" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="8BEA37705572FF9E528DBAEDFDDBFD85" blockId="24.[189,609,500,526]" box="[189,609,500,526]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB45105E5572FF9E528DBAEDFDDBFD85" box="[189,609,500,526]" pageId="24" pageNumber="25">
<treatmentCitation id="0AF411615572FF9E528DBAEDFDDBFD85" author="Reymond" box="[189,609,500,526]" page="143" pageId="24" pageNumber="25" year="1938">
<taxonomicName id="4C554CF35572FF9E528DBAEDFDE6FD85" authority="Reymond, 1938: 143" authorityName="Reymond" authorityPageNumber="143" authorityYear="1938" box="[189,604,500,526]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625572FF9E528DBAEDFEE1FD85" box="[189,347,500,526]" italics="true" pageId="24" pageNumber="25">Dietomorpha</emphasis>
Reymond, 1938: 143
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA37705572FF9E528DB92CFD22FDC4" blockId="24.[189,1204,565,627]" box="[189,664,565,592]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB45105E5572FF9E528DB92CFD22FDC4" box="[189,664,565,592]" pageId="24" pageNumber="25">
<emphasis id="B921EB625572FF9E528DB92CFEC8FDC4" box="[189,370,565,591]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C554CF35572FF9E528DB92CFEE1FDC4" authorityName="Reymond" authorityYear="1938" box="[189,347,565,591]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Dietomorpha</taxonomicName>
–
</emphasis>
<treatmentCitation id="0AF411615572FF9E5349B92CFDF3FDC4" author="Kuhnelt W." box="[377,585,565,592]" page="86" pageId="24" pageNumber="25" year="1957">
<bibRefCitation id="EFC44A815572FF9E5349B92CFDF3FDC4" author="Kuhnelt W." box="[377,585,565,592]" pageId="24" pageNumber="25" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt 1957: 86</bibRefCitation>
</treatmentCitation>
, fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA37705572FF9E528DB940FB0EFDF8" blockId="24.[189,1204,565,627]" box="[189,1204,600,627]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB45105E5572FF9E528DB940FB0EFDF8" box="[189,1204,600,627]" pageId="24" pageNumber="25">
<taxonomicName id="4C554CF35572FF9E528DB940FEA4FDF9" authorityName="Gabrieli Schuster" authorityYear="1935" box="[189,286,601,626]" class="Insecta" family="Tenebrionidae" genus="Kawiria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625572FF9E528DB940FEA4FDF9" box="[189,286,601,626]" italics="true" pageId="24" pageNumber="25">Kawiria</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AF411615572FF9E530CB941FDF0FDF8" author="Kaszab Z." box="[316,586,600,627]" page="295" pageId="24" pageNumber="25" year="1957">
<bibRefCitation id="EFC44A815572FF9E530CB941FDF0FDF8" author="Kaszab Z." box="[316,586,600,627]" pageId="24" pageNumber="25" pagination="289 - 299" refId="ref31014" refString="Kaszab Z. 1957. Zehn neue Tenebrioniden aus Asien (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici. Series nova 8: 289 - 299." type="journal article" year="1957">Kaszab 1957: 295–296</bibRefCitation>
</treatmentCitation>
(erroneous interpretation of
<taxonomicName id="4C554CF35572FF9E51ABB941FB19FDF9" authorityName="Kuhnelt" authorityYear="1957" box="[923,1187,600,626]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E51ABB941FB19FDF9" box="[923,1187,600,626]" italics="true" pageId="24" pageNumber="25">Dietomorpha pardalis</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5572FF9E528DB983FC82FD68" pageId="24" pageNumber="25" type="type_taxon">
<paragraph id="8BEA37705572FF9E528DB983FEE3FD3F" blockId="24.[189,345,666,692]" box="[189,345,666,692]" pageId="24" pageNumber="25">
<heading id="D0A2801C5572FF9E528DB983FEE3FD3F" bold="true" box="[189,345,666,692]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B921EB625572FF9E528DB983FEE3FD3F" bold="true" box="[189,345,666,692]" pageId="24" pageNumber="25">
<typeStatus id="54EE89D25572FF9E528DB983FF41FD3F" box="[189,251,666,692]" pageId="24" pageNumber="25">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705572FF9E528DB9D1FC82FD68" blockId="24.[189,824,712,739]" box="[189,824,712,739]" pageId="24" pageNumber="25">
<taxonomicName id="4C554CF35572FF9E528DB9D1FD30FD69" authority="Reymond, 1938" authorityName="Reymond" authorityYear="1938" box="[189,650,712,738]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E528DB9D1FE7FFD69" box="[189,453,712,738]" italics="true" pageId="24" pageNumber="25">Dietomorpha pardalis</emphasis>
Reymond, 1938
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5572FF9E528DB810FAF6FCD8" pageId="24" pageNumber="25" type="taxon_list">
<paragraph id="8BEA37705572FF9E528DB810FE34FCAF" blockId="24.[189,398,777,804]" box="[189,398,777,804]" pageId="24" pageNumber="25">
<heading id="D0A2801C5572FF9E528DB810FE34FCAF" bold="true" box="[189,398,777,804]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B921EB625572FF9E528DB810FE34FCAF" bold="true" box="[189,398,777,804]" pageId="24" pageNumber="25">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705572FF9E528DB821FAF6FCD8" blockId="24.[189,1356,824,851]" box="[189,1356,824,851]" pageId="24" pageNumber="25">
<taxonomicName id="4C554CF35572FF9E528DB821FD33FCD9" authority="Reymond, 1938" authorityName="Reymond" authorityYear="1938" box="[189,649,824,850]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E528DB821FE7FFCD9" box="[189,453,824,850]" italics="true" pageId="24" pageNumber="25">Dietomorpha pardalis</emphasis>
Reymond, 1938
</taxonomicName>
,
<taxonomicName id="4C554CF35572FF9E50A6B821FB50FCD9" authority="S. Chigray &amp; Nabozhenko" authorityName="S. Chigray &amp; Nabozhenko" authorityYear="2022" box="[662,1258,824,850]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gonzalesi" status="sp. nov.">
<emphasis id="B921EB625572FF9E50A6B821FC11FCD9" box="[662,939,824,850]" italics="true" pageId="24" pageNumber="25">Dietomorpha gonzalesi</emphasis>
S. Chigray &amp; Nabozhenko
</taxonomicName>
<taxonomicNameLabel id="A21256195572FF9E56C1B820FAF6FCD8" box="[1265,1356,825,851]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5572FF9E528DB860FEC4F83B" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BEA37705572FF9E528DB860FE8DFC18" blockId="24.[189,311,889,915]" box="[189,311,889,915]" pageId="24" pageNumber="25">
<heading id="D0A2801C5572FF9E528DB860FE8DFC18" bold="true" box="[189,311,889,915]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B921EB625572FF9E528DB860FE8DFC18" bold="true" box="[189,311,889,915]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705572FF9E528DB8B1FADCFBDB" blockId="24.[189,1400,936,1104]" pageId="24" pageNumber="25">
Body robust (
<figureCitation id="136E2BF55572FF9E5351B8B1FE10FC49" box="[353,426,936,962]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[249,1340,656,1847]" captionTargetId="figure-122@8.[241,1346,643,1855]" captionTargetPageId="8" captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray &amp; Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales." figureDoi="http://doi.org/10.5281/zenodo.6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="24" pageNumber="25">Fig. 3</figureCitation>
), completely covered with lanceolate longitudinally striated, creamy scales (each scale with median longitudinal ridge at middle) (
<figureCitation id="136E2BF55572FF9E50CAB8D2FCCDFC6D" box="[762,887,971,998]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,457,1853]" captionTargetId="figure-12@15.[189,1398,457,1853]" captionTargetPageId="15" captionText="Fig. 9. Habrobates Semenov, 1903 and Dietomorpha Kühnelt, 1957, details of structure. A. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), left elytron. B. Ditto, elytral scales and rows of setae. C. D. pardalis Kühnelt, 1957, ♂ (ZIN), pronotum. D. Ditto, scales, setae and granules in middle of pronotum. E. Ditto, base of left elytron. F. Ditto, scales, spine-like tubercle and seta on elytron." figureDoi="http://doi.org/10.5281/zenodo.6419453" httpUri="https://zenodo.org/record/6419453/files/figure.png" pageId="24" pageNumber="25">Fig. 9D, F</figureCitation>
), with spotted (
<figureCitation id="136E2BF55572FF9E561DB8D2FB33FC6D" box="[1069,1161,971,998]" captionStart="Fig" captionStartId="7.[190,233,1892,1918]" captionTargetBox="[189,1398,684,1851]" captionTargetId="figure-135@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 2. Habrobates gabrieli (Schuster, 1935) comb. nov., type specimens, habitus, labels (NHB). A. Lectotype, dorsal view. B. Ditto, ventral view. C. Ditto, lateral view. D. Paralectotype, dorsal view. E. Ditto, lateral view. F. Labels of the lectotype. G. Labels of the paralectotype. Photos courtesy of © Naturhistorisches Museum Basel (Basel, Switzerland)." figureDoi="http://doi.org/10.5281/zenodo.6419431" httpUri="https://zenodo.org/record/6419431/files/figure.png" pageId="24" pageNumber="25">Fig. 2A</figureCitation>
) or striped (
<figureCitation id="136E2BF55572FF9E5728B8D2FF4EFB82" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[249,1340,656,1847]" captionTargetId="figure-122@8.[241,1346,643,1855]" captionTargetPageId="8" captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray &amp; Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales." figureDoi="http://doi.org/10.5281/zenodo.6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="24" pageNumber="25">Fig. 3C, E–G</figureCitation>
) pattern of dark scales on elytra and dark spots on pronotum. Body under scales black or darkbrown. Body length
<quantity id="4CAD9A955572FF9E5381BF0BFD9FFBA7" box="[433,549,1042,1068]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="24" pageNumber="25" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">9–14 mm</quantity>
. Epistoma and frons covered with sparse, erected yellowish setae. Eyes large, circular, convex in dorsal view; surface behind eyes concealed by anterior margin of pronotum.
</paragraph>
<paragraph id="8BEA37705572FF9E528DBF6EFD04FA0C" blockId="24.[187,1400,1142,1416]" pageId="24" pageNumber="25">
Pronotum slightly transverse (1.4–1.7× as wide as long); lateral margins and partly prothoracic hypomera basally with bare and smooth black spot. Disc of pronotum (Fig. 52) convex, with two triangular deep impressions at base. Pronotum covered with sparse, erect, conical tubercles becoming larger along lateral margins, each tubercle with spine at apex. Prosternum without transverse triangular depression along anterior margin, longitudinal length 2.2× as short as longitudinal length of one procoxa. Prosternal process elongated and narrow (length 3 × width), raised or not raised between procoxae, strongly (
<taxonomicName id="4C554CF35572FF9E531FBE52FE0FFAEF" authorityName="Reymond" authorityYear="1938" box="[303,437,1354,1380]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E531FBE52FE0FFAEF" box="[303,437,1354,1380]" italics="true" pageId="24" pageNumber="25">D. pardalis</emphasis>
</taxonomicName>
) (
<figureCitation id="136E2BF55572FF9E53FFBE53FD93FAEF" box="[463,553,1354,1380]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[249,1340,656,1847]" captionTargetId="figure-122@8.[241,1346,643,1855]" captionTargetPageId="8" captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray &amp; Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales." figureDoi="http://doi.org/10.5281/zenodo.6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="24" pageNumber="25">Fig. 3B</figureCitation>
) or weakly (
<taxonomicName id="4C554CF35572FF9E50F0BE52FB2FFAEF" authority="S. Chigray &amp; Nabozhenko" authorityName="S. Chigray &amp; Nabozhenko" authorityYear="2022" box="[704,1173,1354,1380]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gonzalesi" status="sp. nov.">
<emphasis id="B921EB625572FF9E50F0BE52FCEFFAEF" box="[704,853,1354,1380]" italics="true" pageId="24" pageNumber="25">D. gonzalesi</emphasis>
S. Chigray &amp; Nabozhenko
</taxonomicName>
<taxonomicNameLabel id="A21256195572FF9E56ADBE53FB43FAEF" box="[1181,1273,1354,1380]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
) (
<figureCitation id="136E2BF55572FF9E5722BE53FAD7FAEF" box="[1298,1389,1354,1380]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[249,1340,656,1847]" captionTargetId="figure-122@8.[241,1346,643,1855]" captionTargetPageId="8" captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray &amp; Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales." figureDoi="http://doi.org/10.5281/zenodo.6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="24" pageNumber="25">Fig. 3D</figureCitation>
) protruding, not touching with mesoventrite.
</paragraph>
<paragraph id="8BEA37705572FF9E528DBEB7FC7DF9F2" blockId="24.[189,1399,1454,1657]" pageId="24" pageNumber="25">
Elytra oval or widely oval, with one (
<taxonomicName id="4C554CF35572FF9E50B8BEB6FBD2FA43" authority="S. Chigray &amp; Nabozhenko" authorityName="S. Chigray &amp; Nabozhenko" authorityYear="2022" box="[648,1128,1454,1481]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gonzalesi" status="sp. nov.">
<emphasis id="B921EB625572FF9E50B8BEB6FCA5FA43" box="[648,799,1454,1480]" italics="true" pageId="24" pageNumber="25">D. gonzalesi</emphasis>
S. Chigray &amp; Nabozhenko
</taxonomicName>
<taxonomicNameLabel id="A21256195572FF9E5643BEB6FB6BFA42" box="[1139,1233,1455,1481]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
) or double in middle (
<taxonomicName id="4C554CF35572FF9E5314BECBFE15FA67" authorityName="Reymond" authorityYear="1938" box="[292,431,1490,1516]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E5314BECBFE15FA67" box="[292,431,1490,1516]" italics="true" pageId="24" pageNumber="25">D. pardalis</emphasis>
</taxonomicName>
) humeral rib; with strong, acute spinose tubercles (with spine in base of each tubercle) (
<figureCitation id="136E2BF55572FF9E5308BEECFE09F984" box="[312,435,1525,1551]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,457,1853]" captionTargetId="figure-12@15.[189,1398,457,1853]" captionTargetPageId="15" captionText="Fig. 9. Habrobates Semenov, 1903 and Dietomorpha Kühnelt, 1957, details of structure. A. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), left elytron. B. Ditto, elytral scales and rows of setae. C. D. pardalis Kühnelt, 1957, ♂ (ZIN), pronotum. D. Ditto, scales, setae and granules in middle of pronotum. E. Ditto, base of left elytron. F. Ditto, scales, spine-like tubercle and seta on elytron." figureDoi="http://doi.org/10.5281/zenodo.6419453" httpUri="https://zenodo.org/record/6419453/files/figure.png" pageId="24" pageNumber="25">Fig. 9E–F</figureCitation>
) on dark spots (
<taxonomicName id="4C554CF35572FF9E504ABEEFFCB9F984" authorityName="Reymond" authorityYear="1938" box="[634,771,1525,1551]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625572FF9E504ABEEFFCB9F984" box="[634,771,1525,1551]" italics="true" pageId="24" pageNumber="25">D. pardalis</emphasis>
</taxonomicName>
) or with five longitudinal rows of spinose tubercles (
<taxonomicName id="4C554CF35572FF9E52F7BD00FD0AF9B9" authority="S. Chigray &amp; Nabozhenko" authorityName="S. Chigray &amp; Nabozhenko" authorityYear="2022" box="[199,688,1560,1586]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gonzalesi" status="sp. nov.">
<emphasis id="B921EB625572FF9E52F7BD00FEDAF9B9" box="[199,352,1560,1586]" italics="true" pageId="24" pageNumber="25">D. gonzalesi</emphasis>
S. Chigray &amp; Nabozhenko
</taxonomicName>
<taxonomicNameLabel id="A21256195572FF9E508DBD00FC9AF9B8" box="[701,800,1561,1587]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
). Scutellar shield completely or partly concealed by base of pronotum; surface around scutellar shield with triangular depression. Transverse length of metacoxae subequal to intercoxal process of abdominal ventrite 1.
</paragraph>
<paragraph id="8BEA37705572FF9E528DBDB9FEC4F83B" blockId="24.[186,1399,1696,1968]" pageId="24" pageNumber="25">
Trochanters with elongate, sparse setae. Meso- and metafemora curved outward. Protibiae with projecting process at apex of outer margin, armed with tuft of spines (
<figureCitation id="136E2BF55572FF9E5140BDDAFC66F955" box="[880,988,1731,1758]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="24" pageNumber="25">Fig. 26B</figureCitation>
). Lateral margins of protibiae with long, fine setae and short spines. Protibial terminal spurs elongated, extending to base of protarsomere 4; mesotibial terminal spurs extending to apex of mesotarsomere 2; metatibial terminal spurs extending to midlength of metatarsomere 1. Outer and inner protibial terminal spurs subequal in length, inner meso- and metatibial terminal spurs longer than outer ones, spurs flattened from sides. All tarsi flattened from sides, covered with long setae dorsally and short setae ventrally. Tarsal claws elongated, thin, weakly curved outward.
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5573FF9F52DABA10FACDFAA1" pageId="25" pageNumber="26" type="description">
<paragraph id="8BEA37705573FF9F52DABA10FDFCFEA8" blockId="25.[189,1399,264,362]" box="[234,582,264,291]" pageId="25" pageNumber="26">
<emphasis id="B921EB625573FF9F52DABA10FE25FEA9" bold="true" box="[234,415,264,291]" pageId="25" pageNumber="26">Male genitalia</emphasis>
(
<figureCitation id="136E2BF55573FF9F539FBA11FD84FEA8" box="[431,574,264,291]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17A–H</figureCitation>
)
</paragraph>
<paragraph id="8BEA37705573FF9F528DBA35FABFFEE1" blockId="25.[189,1399,264,362]" pageId="25" pageNumber="26">
Inner sternite VIII (
<figureCitation id="136E2BF55573FF9F5397BA35FDA8FECD" box="[423,530,300,326]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17H</figureCitation>
) weakly sclerotized on margins, densely covered with long, dense setae; anterior margin deeply emarginated in middle; gland is absent (or at least, was not observed).
</paragraph>
<paragraph id="8BEA37705573FF9F528DBA89FD09FD9F" blockId="25.[189,1399,400,532]" pageId="25" pageNumber="26">
Rods of spiculum gastrale (
<figureCitation id="136E2BF55573FF9F53CEBA89FDD3FE21" box="[510,617,400,426]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17G</figureCitation>
) widely spaced, thickened, arcuately connected at apex, with wide membrane along this connection and near derivatives of inner sternite IX; these derivatives elongated, ladle-shaped, evenly sclerotized in middle and slightly more sclerotized terminally; apical margin of derivatives covered with short dense setae.
</paragraph>
<paragraph id="8BEA37705573FF9F528DB925FE49FC8D" blockId="25.[187,1399,571,774]" pageId="25" pageNumber="26">
Tegmen of aedeagus thickened, obtuse (
<figureCitation id="136E2BF55573FF9F50B9B922FCA2FDDE" box="[649,792,571,597]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17A–C</figureCitation>
). Basal piece of tegmen much shorter and wider than apical piece, with tubercle-shaped apex (
<figureCitation id="136E2BF55573FF9F50ACB947FC97FDF2" box="[668,813,606,633]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17A–B</figureCitation>
). Apical piece strongly curved, narrowly rounded at apex, covered with short setae in apical half; ventral apophyses not expressed, dorsal apophyses moderately long, triangular. Median lobe (
<figureCitation id="136E2BF55573FF9F509AB9BCFC88FD34" box="[682,818,677,703]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17D, F</figureCitation>
) widened in basal ⅔ and narrowed in apical third, with weakly separated apex, strongly curved; basal part with additional four sclerotized armatures; baculi not merged at apex.
</paragraph>
<paragraph id="8BEA37705573FF9F52DAB834FE01FCCD" blockId="25.[189,1398,812,909]" box="[234,443,812,839]" pageId="25" pageNumber="26">
<emphasis id="B921EB625573FF9F52DAB834FE01FCCD" bold="true" box="[234,443,812,839]" pageId="25" pageNumber="26">Female genitalia</emphasis>
</paragraph>
<paragraph id="8BEA37705573FF9F528DB849FE2DFC06" blockId="25.[189,1398,812,909]" pageId="25" pageNumber="26">
Spiculum ventrale (
<figureCitation id="136E2BF55573FF9F5395B849FDBFFCE1" box="[421,517,848,874]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17I</figureCitation>
) very short and strongly widened, sternite VIII with long acutely angulate curved apophyses.
</paragraph>
<paragraph id="8BEA37705573FF9F528DB8ACFC40FBF4" blockId="25.[189,1400,949,1151]" pageId="25" pageNumber="26">
Ovipositor (
<figureCitation id="136E2BF55573FF9F537CB8ACFE63FC44" box="[332,473,949,975]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[189,1398,335,1814]" captionTargetId="figure-17@38.[189,1398,327,1814]" captionTargetPageId="38" captionText="Fig. 17. Dietomorpha pardalis Kühnelt, 1957, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419477" httpUri="https://zenodo.org/record/6419477/files/figure.png" pageId="25" pageNumber="26">Fig. 17G–L</figureCitation>
) short, weakly sclerotized. Coxite with four distinct not merged lobes. Baculi of coxite lobe I short, shorter than paraproct baculi; lobe II small, sclerotized; lobe III larger, conical, sclerotized; lobe IV membranous (only at apex slightly sclerotized), conical, with dense pubescence of long setae. Paraproct V-shaped on each side ventrally, baculi sharply widened in base and narrow in apical third. Vulva with narrow sclerotization in the middle of ventral side. Apical margin of proctiger with deep V-shape emargination, baculi of proctiger strongly widened.
</paragraph>
<paragraph id="8BEA37705573FF9F52DABFBFFACDFAA1" blockId="25.[189,1400,1190,1322]" pageId="25" pageNumber="26">
<emphasis id="B921EB625573FF9F52DABFBFFE54FB4B" bold="true" box="[234,494,1190,1216]" pageId="25" pageNumber="26">Female genital ducts</emphasis>
(
<figureCitation id="136E2BF55573FF9F53CFBFBFFDD2FB4B" box="[511,616,1190,1216]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="25" pageNumber="26">Fig. 23C</figureCitation>
) Vagina moderately widened, sacciform, narrower before oviduct and after spermatheca, apical part of vagina not elastically curved. Spermatheca very short, single-tube. Accessory gland long, with two tubelike constrictions, short in base and long in middle. Basal part between two constrictions not sacciform.
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5573FF9F528DBE48FC40FA11" box="[189,1018,1361,1435]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BEA37705573FF9F528DBE48FEE2FAE0" blockId="25.[189,344,1361,1387]" box="[189,344,1361,1387]" pageId="25" pageNumber="26">
<heading id="D0A2801C5573FF9F528DBE48FEE2FAE0" bold="true" box="[189,344,1361,1387]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B921EB625573FF9F528DBE48FEE2FAE0" bold="true" box="[189,344,1361,1387]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705573FF9F528DBE99FC40FA11" blockId="25.[189,1018,1408,1435]" box="[189,1018,1408,1435]" pageId="25" pageNumber="26">
<collectingCountry id="F34277E05573FF9F528DBE99FF56FA11" box="[189,236,1408,1434]" name="Iran" pageId="25" pageNumber="26">Iran</collectingCountry>
and
<collectingCountry id="F34277E05573FF9F5315BE99FE32FA11" box="[293,392,1408,1434]" name="Pakistan" pageId="25" pageNumber="26">Pakistan</collectingCountry>
(
<collectingRegion id="4991F9925573FF9F53A8BE99FDC1FA10" box="[408,635,1408,1435]" country="Pakistan" name="Balochistan" pageId="25" pageNumber="26">Balochistan region</collectingRegion>
),
<collectingCountry id="F34277E05573FF9F50BDBE99FCA6FA11" box="[653,796,1408,1434]" name="Afghanistan" pageId="25" pageNumber="26">Afghanistan</collectingCountry>
(Registan Desert).
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5573FF9F528DBED8FC8EF8CC" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BEA37705573FF9F528DBED8FEBEFA50" blockId="25.[189,260,1473,1499]" box="[189,260,1473,1499]" pageId="25" pageNumber="26">
<heading id="D0A2801C5573FF9F528DBED8FEBEFA50" bold="true" box="[189,260,1473,1499]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B921EB625573FF9F528DBED8FEBEFA50" bold="true" box="[189,260,1473,1499]" pageId="25" pageNumber="26">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705573FF9F528DBEE9FC8EF8CC" blockId="25.[188,1399,1519,1863]" pageId="25" pageNumber="26">
<bibRefCitation id="EFC44A815573FF9F528DBEE9FEE6F981" author="Koch C." box="[189,348,1519,1546]" pageId="25" pageNumber="26" pagination="741 - 742" refId="ref31101" refString="Koch C. 1964. Zur Synonymie von Dietomorpha pardalis Reymond (Col., Tenebrionidae). Entomologische Arbeiten aus dem Museum G. Frey 15: 741 - 742." type="journal article" year="1964">Koch (1964)</bibRefCitation>
synonymized
<taxonomicName id="4C554CF35573FF9F502CBEE9FC94F981" authorityName="Reymond" authorityYear="1938" box="[540,814,1520,1546]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625573FF9F502CBEE9FC94F981" box="[540,814,1520,1546]" italics="true" pageId="25" pageNumber="26">Dietomorpha pardalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554CF35573FF9F514BBEE9FAAFF981" authority="Kaszab, 1957" authorityName="Kaszab" authorityYear="1957" box="[891,1301,1519,1546]" class="Insecta" family="Tenebrionidae" genus="Kawiria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="B921EB625573FF9F514BBEE9FBE3F981" box="[891,1113,1520,1546]" italics="true" pageId="25" pageNumber="26">Kawiria szekessyi</emphasis>
<bibRefCitation id="EFC44A815573FF9F565BBEE9FAAFF981" author="Kaszab Z." box="[1131,1301,1519,1546]" pageId="25" pageNumber="26" pagination="289 - 299" refId="ref31014" refString="Kaszab Z. 1957. Zehn neue Tenebrioniden aus Asien (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici. Series nova 8: 289 - 299." type="journal article" year="1957">Kaszab, 1957</bibRefCitation>
</taxonomicName>
(junior synonym). Koch also noted that Reymond (1938) should be considered as the author of the genus
<taxonomicName id="4C554CF35573FF9F528DBD2FFEE1F9DB" authorityName="Reymond" authorityYear="1938" box="[189,347,1590,1616]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625573FF9F528DBD2FFEE1F9DB" box="[189,347,1590,1616]" italics="true" pageId="25" pageNumber="26">Dietomorpha</emphasis>
</taxonomicName>
and the species
<taxonomicName id="4C554CF35573FF9F501EBD2FFC80F9DB" authorityName="Reymond" authorityYear="1938" box="[558,826,1590,1616]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625573FF9F501EBD2FFC80F9DB" box="[558,826,1590,1616]" italics="true" pageId="25" pageNumber="26">Dietomorpha pardalis</emphasis>
</taxonomicName>
, because he presented figures for this species.
<bibRefCitation id="EFC44A815573FF9F528DBD40FED1F9FF" author="Kaszab Z." box="[189,363,1625,1652]" pageId="25" pageNumber="26" pagination="289 - 299" refId="ref31014" refString="Kaszab Z. 1957. Zehn neue Tenebrioniden aus Asien (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici. Series nova 8: 289 - 299." type="journal article" year="1957">Kaszab (1957)</bibRefCitation>
noted that Reymond (1938) did not give a description or a differential diagnosis of the genus, and the author of
<taxonomicName id="4C554CF35573FF9F53C3BD65FD45F91D" authorityName="Reymond" authorityYear="1938" box="[499,767,1660,1686]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625573FF9F53C3BD65FD45F91D" box="[499,767,1660,1686]" italics="true" pageId="25" pageNumber="26">Dietomorpha pardalis</emphasis>
</taxonomicName>
should therefore be
<bibRefCitation id="EFC44A815573FF9F5630BD65FB01F91C" author="Kuhnelt W." box="[1024,1211,1660,1687]" pageId="25" pageNumber="26" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt (1957)</bibRefCitation>
, who distinctly described, compared and figured this taxon. Later
<bibRefCitation id="EFC44A815573FF9F512BBDB9FC62F931" author="Kwieton E." box="[795,984,1696,1722]" pageId="25" pageNumber="26" pagination="21 - 26" refId="ref31381" refString="Kwieton E. 1982. Notes sur les Tenebrionidae Pimeliinae du Pakistan. Bulletin de la Societe entomologique de Mulhouse 26: 21 - 26." type="journal article" year="1982">Kwieton (1982)</bibRefCitation>
cited Koch’s work but incorrectly established the following synonymy:
<taxonomicName id="4C554CF35573FF9F504ABDDDFBB6F956" authority="Reymond, 1938" authorityName="Reymond" authorityYear="1938" box="[634,1036,1731,1757]" class="Insecta" family="Tenebrionidae" genus="Kawiria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pardalis" status="comb. nov.">
<emphasis id="B921EB625573FF9F504ABDDDFCFCF956" box="[634,838,1731,1757]" italics="true" pageId="25" pageNumber="26">Kawiria pardalis</emphasis>
Reymond, 1938
</taxonomicName>
,
<taxonomicNameLabel id="A21256195573FF9F5629BDDAFB3AF956" box="[1049,1152,1731,1757]" pageId="25" pageNumber="26" rank="species">comb. n.</taxonomicNameLabel>
=
<taxonomicName id="4C554CF35573FF9F5693BDDDFACDF956" authorityName="Kaszab" authorityYear="1957" box="[1187,1399,1731,1757]" class="Insecta" family="Tenebrionidae" genus="Kawiria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="B921EB625573FF9F5693BDDDFACDF956" box="[1187,1399,1731,1757]" italics="true" pageId="25" pageNumber="26">Kawiria szekessyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554CF35573FF9F52DFBDFFFDEAF88B" authority="Reymond, 1938" authorityName="Reymond" authorityYear="1938" box="[239,592,1766,1792]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625573FF9F52DFBDFFFE37F88B" box="[239,397,1766,1792]" italics="true" pageId="25" pageNumber="26">Dietomorpha</emphasis>
Reymond, 1938
</taxonomicName>
=
<taxonomicName id="4C554CF35573FF9F505EBDFEFC3DF88A" authority="Schuster, 1957" authorityName="Schuster" authorityYear="1957" box="[622,903,1766,1793]" class="Insecta" family="Tenebrionidae" genus="Kawiria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625573FF9F505EBDFEFD75F88B" box="[622,719,1767,1792]" italics="true" pageId="25" pageNumber="26">Kawiria</emphasis>
Schuster, 1957
</taxonomicName>
.
<bibRefCitation id="EFC44A815573FF9F51A3BDFFFBDEF88A" author="Medvedev G. S." box="[915,1124,1766,1793]" pageId="25" pageNumber="26" pagination="299 - 314" refId="ref32159" refString="Medvedev G. S. 2005. On connections of the sand desert faunas of Tenebrionidae (Coleoptera) of Middle Asia, Iran, and Afghanistan. In: Konstantinov A. S., Tishechkin A. K. &amp; Penev L. (eds) Contribution to Systematics and Biology of Beetles. Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin: 299 - 314. Pensoft, Moscow." type="book chapter" year="2005">Medvedev (2005)</bibRefCitation>
was not aware of these works and re-established the synonymy previously proposed by Koch, and he also recognised the author of the binomen
<taxonomicName id="4C554CF35573FF9F5346BC35FC8AF8CC" authority="Kuhnelt, 1957" authorityName="Kuhnelt" authorityYear="1957" box="[374,816,1836,1863]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625573FF9F5346BC35FDC7F8CD" box="[374,637,1836,1862]" italics="true" pageId="25" pageNumber="26">Dietomorpha pardalis</emphasis>
<bibRefCitation id="EFC44A815573FF9F50B5BC35FC8AF8CC" author="Kuhnelt W." box="[645,816,1836,1863]" pageId="25" pageNumber="26" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt, 1957</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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