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<document id="D49637A22D5F5AC2DA818A4EE59F179F" ID-CLB-Dataset="38993" ID-DOI="10.11646/zootaxa.4114.5.2" ID-GBIF-Dataset="6e1b04bb-6365-4ce1-85f3-59c0f92854ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="265300" ID-ZooBank="8F1A8843-F034-4C7B-A1F9-F39548E3F2C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464242988099" checkinUser="plazi" docAuthor="Kuhara, Naotoshi" docDate="2016" docId="03F39E16FFA7F25D0D97F8B6FB08FF33" docLanguage="en" docName="zootaxa.4114.5.2.pdf.imf" docOrigin="Zootaxa 4114 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ecnomus hokkaidensis Kuhara, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="564" masterDocId="FFCAE66EFFA7F25E0D00FFC4FFBBFFCB" masterDocTitle="Revision of Japanese species of the genus Ecnomus McLachlan (Trichoptera: Ecnomidae), with descriptions of two new species" masterLastPageNumber="571" masterPageNumber="561" pageNumber="561" updateTime="1698662624330" updateUser="ExternalLinkService">
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<mods:title id="A9BC687316BDF186F02ECED7B49CC0E3">Revision of Japanese species of the genus Ecnomus McLachlan (Trichoptera: Ecnomidae), with descriptions of two new species</mods:title>
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<treatment id="03F39E16FFA7F25D0D97F8B6FB08FF33" ID-DOI="http://doi.org/10.5281/zenodo.6069232" ID-GBIF-Taxon="120731464" ID-Zenodo-Dep="6069232" LSID="urn:lsid:plazi:treatment:03F39E16FFA7F25D0D97F8B6FB08FF33" httpUri="http://treatment.plazi.org/id/03F39E16FFA7F25D0D97F8B6FB08FF33" lastPageId="3" lastPageNumber="564" pageId="0" pageNumber="561">
<subSubSection id="C3407C8BFFA7F25E0D97F8B6FE84F865" pageId="0" pageNumber="561" type="nomenclature">
<paragraph id="8BE52F00FFA7F25E0D97F8B6FDA8F846" blockId="0.[151,531,1906,1966]" box="[151,531,1906,1933]" pageId="0" pageNumber="561">
<heading id="D0AD986CFFA7F25E0D97F8B6FDA8F846" bold="true" box="[151,531,1906,1933]" fontSize="11" level="1" pageId="0" pageNumber="561" reason="1">
<emphasis id="B92EF312FFA7F25E0D97F8B6FDA8F846" bold="true" box="[151,531,1906,1933]" pageId="0" pageNumber="561">
<taxonomicName id="4C5A5483FFA7F25E0D97F8B6FE15F847" box="[151,430,1906,1932]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="0" pageNumber="561" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="B92EF312FFA7F25E0D97F8B6FE15F847" bold="true" box="[151,430,1906,1932]" italics="true" pageId="0" pageNumber="561">Ecnomus hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4E69FFA7F25E0CB5F8B7FDA8F846" box="[437,531,1907,1933]" pageId="0" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE52F00FFA7F25E0D97F851FE84F865" blockId="0.[151,531,1906,1966]" box="[151,319,1941,1966]" pageId="0" pageNumber="561">
(
<figureCitation id="13613385FFA7F25E0D9FF851FF5CF865" box="[159,231,1941,1966]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="0" pageNumber="561">Figs. 1</figureCitation>
AJ, 6)
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA7F25E0D97F81AFDCAF838" box="[151,625,2013,2035]" pageId="0" pageNumber="561" type="description">
<paragraph id="8BE52F00FFA7F25E0D97F81AFDCAF838" blockId="0.[151,625,2013,2035]" box="[151,625,2013,2035]" pageId="0" pageNumber="561">
<taxonomicName id="4C5A5483FFA7F25E0D97F81AFF4FF838" box="[151,244,2014,2035]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="0" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B92EF312FFA7F25E0D97F81AFF4FF838" box="[151,244,2014,2035]" italics="true" pageId="0" pageNumber="561">Ecnomus</emphasis>
</taxonomicName>
sp.:
<bibRefCitation id="EFCB52F1FFA7F25E0C24F819FE58F838" author="Kuhara" box="[292,483,2013,2035]" pageId="0" pageNumber="561" refString="Kuhara, N., Nagayasu, Y. &amp; Ito, T. (2007) Shikotsu-ko to Shikotsu-ko ryunyu kasen no tobikera mokuroku (A list of caddisflies collected in Shikotsu-ko and streams flowing into the lake). In: The Editing Committee of Nature and Lives in Lake Shikotsu-ko (Ed.), Nature and Lives in Lake Shikotsu-ko. Study Group of the Chip and the Water of Lake Shikotsu-ko, Sapporo, pp. 65 - 74 (appendix). [in Japanese]" type="book chapter" year="2007">
Kuhara
<emphasis id="B92EF312FFA7F25E0C75F81AFE1AF838" box="[373,417,2013,2035]" italics="true" pageId="0" pageNumber="561">et al</emphasis>
. 2007
</bibRefCitation>
, appendix 69.
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA6F25F0D97FF53FDF6FEAE" pageId="1" pageNumber="562" type="diagnosis">
<paragraph id="8BE52F00FFA6F25F0D97FF53FDF6FEAE" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0D97FF53FEB1FF7B" bold="true" box="[151,266,151,176]" pageId="1" pageNumber="562">Diagnosis</emphasis>
. The male of this species is very similar to that of
<taxonomicName id="4C5A5483FFA6F25F0E57FF5DFC74FF7B" box="[855,975,152,176]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="B92EF312FFA6F25F0E57FF5DFC74FF7B" box="[855,975,152,176]" italics="true" pageId="1" pageNumber="562">E. tenellus</emphasis>
</taxonomicName>
, but can be distinguished from it by the shape and structure of the inferior appendages. In
<taxonomicName id="4C5A5483FFA6F25F0FEFFF79FC18FF1F" box="[751,931,188,212]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="hokkaidensis">
<emphasis id="B92EF312FFA6F25F0FEFFF79FCBBFF1F" box="[751,768,189,212]" italics="true" pageId="1" pageNumber="562">E</emphasis>
.
<emphasis id="B92EF312FFA6F25F0E13FF78FC18FF1F" box="[787,931,188,212]" italics="true" pageId="1" pageNumber="562">hokkaidensis</emphasis>
</taxonomicName>
, each inferior appendage bears a spatulate projection basodorsally, whereas
<taxonomicName id="4C5A5483FFA6F25F0F12FF25FD30FF33" box="[530,651,224,248]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="B92EF312FFA6F25F0F12FF25FD98FF33" box="[530,547,225,248]" italics="true" pageId="1" pageNumber="562">E</emphasis>
.
<emphasis id="B92EF312FFA6F25F0F34FF24FD30FF33" box="[564,651,224,248]" italics="true" pageId="1" pageNumber="562">tenellus</emphasis>
</taxonomicName>
has an ear-like dorsal projection at midlength. In ventral aspect, the inferior appendages are relatively longer and more slender and are angled abruptly inward at midlength in
<taxonomicName id="4C5A5483FFA6F25F0884FEC1FE93FE8B" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="hokkaidensis">
<emphasis id="B92EF312FFA6F25F0884FEC1FA2EFED7" box="[1412,1429,261,284]" italics="true" pageId="1" pageNumber="562">E</emphasis>
.
<emphasis id="B92EF312FFA6F25F0D97FEECFE93FE8B" box="[151,296,296,320]" italics="true" pageId="1" pageNumber="562">hokkaidensis</emphasis>
</taxonomicName>
, but they are stouter and smoothly curved in
<taxonomicName id="4C5A5483FFA6F25F0E1EFEEDFC2FFE8B" box="[798,916,296,320]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="B92EF312FFA6F25F0E1EFEEDFC94FE8B" box="[798,815,297,320]" italics="true" pageId="1" pageNumber="562">E</emphasis>
.
<emphasis id="B92EF312FFA6F25F0E3DFEECFC2FFE8B" box="[829,916,296,320]" italics="true" pageId="1" pageNumber="562">tenellus</emphasis>
</taxonomicName>
. The female of this species is characterized by sternum VII with 2 ventromesal plates.
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA6F25F0DC7FEABFC47FAC4" pageId="1" pageNumber="562" type="description">
<paragraph id="8BE52F00FFA6F25F0DC7FEABFAF1FDA8" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7FEABFE1EFE43" bold="true" box="[199,421,367,392]" pageId="1" pageNumber="562">Description. Adult</emphasis>
(
<figureCitation id="13613385FFA6F25F0CB5FEABFE43FE43" box="[437,504,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
). Length of each forewing of male
<quantity id="4CA282E5FFA6F25F0E90FEABFBA2FE4C" box="[912,1049,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.7" metricValueMin="4.5" pageId="1" pageNumber="562" unit="mm" value="5.1" valueMax="5.7" valueMin="4.5">4.55.7 mm</quantity>
(mean = 5.0 mm, n = 17), female
<quantity id="4CA282E5FFA6F25F0D97FE50FE9BFE67" box="[151,288,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.7" pageId="1" pageNumber="562" unit="mm" value="5.1" valueMax="5.5" valueMin="4.7">4.75.5 mm</quantity>
(mean = 5.0 mm, n = 14). Head frons with single frontal wart, pair of ellipsoidal dorsal hypomedial warts, and indistinct pair of small ventral hypomedial warts; vertex with pairs of oval antennal, preocellar, ocellar, and occipital warts (
<figureCitation id="13613385FFA6F25F0C7EFE18FE73FE3F" box="[382,456,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
A, 1B). Pronotum with 2 pairs of warts; lateral warts indistinct laterally; medial warts round; mesonotum with pair of round scutal warts and pair of semicircular scutellar warts (
<figureCitation id="13613385FFA6F25F09B7FE3BFB42FDD3" box="[1207,1273,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
A). Legs with tibial spur formula 3, 4, 4. Forewings each with fork of R1 and forks I, II, III, IV and V, discoidal cell, median cell and thyridial cell; crossvein
<emphasis id="B92EF312FFA6F25F0CD4FD8FFDB7FDA9" box="[468,524,587,610]" italics="true" pageId="1" pageNumber="562">m-cu</emphasis>
near bifurcation of medial vein; hind wings with forks II and V (
<figureCitation id="13613385FFA6F25F09E5FD8EFA9FFDA8" box="[1253,1316,586,611]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="8BE52F00FFA6F25F0DC7FDAAFCD3FBB4" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7FDAAFEC0FD4C" bold="true" box="[199,379,622,647]" pageId="1" pageNumber="562">Male genitalia</emphasis>
(
<figureCitation id="13613385FFA6F25F0C93FDAAFE5EFD4C" box="[403,485,622,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
DH). Tergum IX long, reaching posterior margin of sternum IX (
<figureCitation id="13613385FFA6F25F082BFDAAFAC8FD4C" box="[1323,1395,622,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
D); anterodorsal margin deeply and widely excised (
<figureCitation id="13613385FFA6F25F0FB8FD56FD4CFD60" box="[696,759,658,683]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
E); sternum IX drop-shaped in lateral aspect (
<figureCitation id="13613385FFA6F25F09FCFD56FA87FD60" box="[1276,1340,658,683]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
D), with longitudinal median line in ventral aspect (
<figureCitation id="13613385FFA6F25F0F89FD72FD7FFD04" box="[649,708,694,719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig.1</figureCitation>
F); anteroventral margin deeply notched; posteroventral margin shallowly and broadly excised (
<figureCitation id="13613385FFA6F25F0F05FD1EFDFCFD38" box="[517,583,730,755]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
F). Segment X with pair of finger-shaped posteroventral projections, each weakly bent inward at midlength and with 2 setae apically; anterolateral arms curved outward anteriorly (
<figureCitation id="13613385FFA6F25F0838FD3AFAC3FCDC" box="[1336,1400,766,791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
E). Superior appendages elongate-triangular, tapering to blunt apices in lateral aspect (
<figureCitation id="13613385FFA6F25F0976FCE6FB07FCF0" box="[1142,1212,802,827]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
D); mesal surfaces concave, each with short triangular projection near base bearing 3 apical setae (
<figureCitation id="13613385FFA6F25F0923FC82FBDFFC94" box="[1059,1124,838,863]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
E); stout setae arising from apical and dorsal margins in distal 2/5 (
<figureCitation id="13613385FFA6F25F0F76FCAEFD7DFC48" box="[630,710,874,899]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
D, 1E). Inferior appendages subequal in length with superior appendages; each elongate, slightly upcurved, thick basally, constricted medially in lateral aspect (
<figureCitation id="13613385FFA6F25F09F9FC4AFA82FC6C" box="[1273,1337,910,935]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
D); with outer margin abruptly angled 45° inward at midlength, tapered to blunt apex in ventral aspect (
<figureCitation id="13613385FFA6F25F09F8FC76FA87FC00" box="[1272,1340,946,971]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
F); with mesally-directed spatulate projection basodorsally (
<figureCitation id="13613385FFA6F25F0FE0FC12FC99FC24" box="[736,802,982,1007]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
G). Basal plate of inferior appendages sclerotized, with pair of small triangular projections laterally 1/3rd distance from anterior end (
<figureCitation id="13613385FFA6F25F091BFC3EFBD3FBD8" box="[1051,1128,1018,1043]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Figs. 1</figureCitation>
F, 1G); with pair of small, longitudinal, posterodorsal projections directed anteromesad and bearing setae along margins (
<figureCitation id="13613385FFA6F25F09D6FBDAFAA3FBFC" box="[1238,1304,1054,1079]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
H). Phallus with strongly sclerotized phallobase, tubular, curved downward (
<figureCitation id="13613385FFA6F25F0E7AFB86FC00FB90" box="[890,955,1090,1115]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
H); distal 1/3 flattened dorsoventrally and spoon-shaped (
<figureCitation id="13613385FFA6F25F0C40FBA2FE3BFBB4" box="[320,384,1126,1151]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
G); parameres and basodorsal lobes absent.
</paragraph>
<paragraph id="8BE52F00FFA6F25F0DC7FB4EFC47FAC4" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7FB4EFE34FB68" bold="true" box="[199,399,1162,1187]" pageId="1" pageNumber="562">Female genitalia</emphasis>
(
<figureCitation id="13613385FFA6F25F0CA1FB4EFE54FB68" box="[417,495,1162,1187]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">
Figs.
<date id="FFE409C0FFA6F25F0CE3FB4FFE54FB68" box="[483,495,1163,1187]" pageId="1" pageNumber="562">1</date>
</figureCitation>
I, 1J). Sternum VII with 2 distinctly pigmented ventromesal plates; anterior plate oval; posterior plate wide, concave anteriorly with posterolateral extensions in ventral aspect (
<figureCitation id="13613385FFA6F25F09E0FB6AFA9FFB0C" box="[1248,1316,1198,1223]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
J). Ventral plates of sternum VIII suboval, each with 3 long setae along apical margin (
<figureCitation id="13613385FFA6F25F0943FB16FB31FB20" box="[1091,1162,1234,1259]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">Fig. 1</figureCitation>
J). Segment XI setose, unpigmented with narrow, oblique, sclerotized bands anterolaterally (
<figureCitation id="13613385FFA6F25F0EA0FB32FC5BFAC4" box="[928,992,1270,1295]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[162,1413,197,1880]" captionTargetId="figure@2.[162,1413,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Ecnomus hokkaidensis sp. nov. (type locality; 1 D 1 F, holotype). 1 A, head, prothorax and mesothorax, dorsal; 1 B, head, frontal; 1 C, right wings, venation, dorsal; 1 D 1 F, male genitalia: 1 D, left lateral; 1 E, dorsal; 1 F, ventral; 1 G 1 H, male phallus and inferior appendages: 1 G, dorsal; 1 H, left lateral; 1 I 1 J, female genitalia: 1 I, left lateral; 1 J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein m-cu; f R 1 = fork of R 1; F 1 F 5 = fork 1 fork 5." httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="1" pageNumber="562">
Fig.
<date id="FFE409C0FFA6F25F0ED4FB33FC5BFAC4" box="[980,992,1271,1295]" pageId="1" pageNumber="562">1</date>
</figureCitation>
I).
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA6F25F0DC7FADEFAF2F884" pageId="1" pageNumber="562" type="materials_examined">
<paragraph id="8BE52F00FFA6F25F0DC7FADEFE04FA9C" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<typeStatus id="54E191A2FFA6F25F0DC7FADEFE8FFAF8" box="[199,308,1306,1331]" pageId="1" pageNumber="562" type="holotype">
<emphasis id="B92EF312FFA6F25F0DC7FADEFE8FFAF8" bold="true" box="[199,308,1306,1331]" pageId="1" pageNumber="562">Holotype</emphasis>
</typeStatus>
. male (pinned),
<collectingCountry id="F34D6F90FFA6F25F0CF0FADFFD88FAF9" box="[496,563,1307,1330]" name="Japan" pageId="1" pageNumber="562">Japan</collectingCountry>
, Hokkaidô, Ishikari, Chitose-shi, Shikotsu-ko, Poropinai,
<geoCoordinate id="EE6E49C7FFA6F25F09C3FADFFA99FAF8" box="[1219,1314,1307,1331]" direction="north" orientation="latitude" pageId="1" pageNumber="562" precision="555" value="42.8">42.80ºN</geoCoordinate>
,
<geoCoordinate id="EE6E49C7FFA6F25F082EFADFFA23FAF8" box="[1326,1432,1307,1331]" direction="east" orientation="longitude" pageId="1" pageNumber="562" precision="555" value="141.34">141.34ºE</geoCoordinate>
,
<date id="FFE409C0FFA6F25F0D97FAFAFE9BFA9C" box="[151,288,1342,1367]" pageId="1" pageNumber="562" value="2006-08-14">14.viii.2006</date>
, NK (
<collectionCode id="ED4BB7C5FFA6F25F0C64FAFBFE08FA9C" box="[356,435,1343,1367]" pageId="1" pageNumber="562">SEHU</collectionCode>
).
</paragraph>
<paragraph id="8BE52F00FFA6F25F0DC7FAA6FC6AFA54" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<typeStatus id="54E191A2FFA6F25F0DC7FAA6FE84FAB0" box="[199,319,1378,1403]" pageId="1" pageNumber="562" type="paratype">
<emphasis id="B92EF312FFA6F25F0DC7FAA6FE84FAB0" bold="true" box="[199,319,1378,1403]" pageId="1" pageNumber="562">Paratypes</emphasis>
</typeStatus>
.
<specimenCount id="9D5CE489FFA6F25F0C51FAA7FE1BFAB0" box="[337,416,1378,1403]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
(pinned),
<typeStatus id="54E191A2FFA6F25F0F1DFAA7FDF6FAB0" box="[541,589,1379,1403]" pageId="1" pageNumber="562">type</typeStatus>
locality,
<date id="FFE409C0FFA6F25F0FBEFAA6FCF3FAB0" box="[702,840,1378,1403]" pageId="1" pageNumber="562" value="2005-08-12">12.viii.2005</date>
, NK (
<collectionCode id="ED4BB7C5FFA6F25F0E93FAA7FC59FAB0" box="[915,994,1379,1403]" pageId="1" pageNumber="562">SEHU</collectionCode>
);
<specimenCount id="9D5CE489FFA6F25F0EFAFAA7FBD8FAB0" box="[1018,1123,1378,1403]" pageId="1" pageNumber="562" type="male">14 males</specimenCount>
(in alcohol),
<typeStatus id="54E191A2FFA6F25F0807FAA7FA8CFAB0" box="[1287,1335,1379,1403]" pageId="1" pageNumber="562">type</typeStatus>
locality,
<date id="FFE409C0FFA6F25F0D97FA42FE9AFA54" box="[151,289,1414,1439]" pageId="1" pageNumber="562" value="2005-08-10">10.viii.2005</date>
, NK (
<collectionCode id="ED4BB7C5FFA6F25F0C64FA43FE13FA54" box="[356,424,1415,1439]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13737" name="Natural History Museum and Institute" pageId="1" pageNumber="562">CBM</collectionCode>
);
<specimenCount id="9D5CE489FFA6F25F0CBBFA43FDAFFA54" box="[443,532,1414,1439]" pageId="1" pageNumber="562" type="male">4 males</specimenCount>
,
<typeStatus id="54E191A2FFA6F25F0F1FFA4CFDF4FA6B" box="[543,591,1416,1440]" pageId="1" pageNumber="562">type</typeStatus>
locality,
<date id="FFE409C0FFA6F25F0FB7FA42FC88FA54" box="[695,819,1414,1439]" pageId="1" pageNumber="562" value="2005-08-05">5.viii.2005</date>
, NK (
<collectionCode id="ED4BB7C5FFA6F25F0E76FA43FC7EFA54" box="[886,965,1415,1439]" pageId="1" pageNumber="562">SEHU</collectionCode>
).
</paragraph>
<paragraph id="8BE52F00FFA6F25F0DC7FA6EFAF2F884" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7FA6EFE2FFA08" bold="true" box="[199,404,1450,1475]" pageId="1" pageNumber="562">Other specimens</emphasis>
. [Hokkaidô] Kushiro:
<specimenCount id="9D5CE489FFA6F25F0FA2FA6FFCBBFA08" box="[674,768,1450,1475]" pageId="1" pageNumber="562" type="male">2 males</specimenCount>
, Kushiro-chô, Hokuchiku-ura,
<date id="FFE409C0FFA6F25F096DFA6EFB4DFA08" box="[1133,1270,1450,1475]" pageId="1" pageNumber="562" value="2007-08-15">15.viii.2007</date>
, T. Kosugi;
<specimenCount id="9D5CE489FFA6F25F088EFA6EFF60FA2C" pageId="1" pageNumber="562" type="male">6 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0DE9FA0BFEDCFA2C" box="[233,359,1486,1511]" pageId="1" pageNumber="562" type="female">11 females</specimenCount>
, Kushiro-chô, Kushiro-gawa,
<date id="FFE409C0FFA6F25F0FC2FA0AFCF0FA2C" box="[706,843,1486,1511]" pageId="1" pageNumber="562" value="2007-08-15">15.viii.2007</date>
, TI &amp; T. Kosugi;
<specimenCount id="9D5CE489FFA6F25F0920FA0BFBC7FA2C" box="[1056,1148,1486,1511]" pageId="1" pageNumber="562" type="male">2 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F098AFA0BFB40FA2C" box="[1162,1275,1486,1511]" pageId="1" pageNumber="562" type="female">2 females</specimenCount>
, Kushiro-chô, Takkobu-numa,
<date id="FFE409C0FFA6F25F0C4EFA36FE71F9C0" box="[334,458,1522,1547]" pageId="1" pageNumber="562" value="2006-08-08">8.viii.2006</date>
, TI;
<specimenCount id="9D5CE489FFA6F25F0CFEFA37FDECF9C0" box="[510,599,1522,1547]" pageId="1" pageNumber="562" type="male">2 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0F62FA37FD7DF9C0" box="[610,710,1522,1547]" pageId="1" pageNumber="562" type="female">1 female</specimenCount>
, Kushiro-shi, Iwabokki,
<date id="FFE409C0FFA6F25F0EDBFA36FBF4F9C0" box="[987,1103,1522,1547]" pageId="1" pageNumber="562" value="1987-07-03">3.vii.1987</date>
, M. Itou;
<specimenCount id="9D5CE489FFA6F25F09BDFA37FA9AF9C0" box="[1213,1313,1522,1547]" pageId="1" pageNumber="562" type="female">1 female</specimenCount>
, Shibechachô, Shirarutoro-ko,
<date id="FFE409C0FFA6F25F0C81F9D2FE4DF9E4" box="[385,502,1558,1583]" pageId="1" pageNumber="562" value="2006-07-07">7.vii.2006</date>
, TI. Sôya:
<specimenCount id="9D5CE489FFA6F25F0F72F9D3FD77F9E4" box="[626,716,1558,1583]" pageId="1" pageNumber="562" type="male">2 males</specimenCount>
, Sarufutsu-mura, Narita-gawa, Kaede-bashi,
<date id="FFE409C0FFA6F25F09C8F9D2FAF0F9E4" box="[1224,1355,1558,1583]" pageId="1" pageNumber="562" value="2007-07-31">31.vii.2007</date>
, NK &amp; TI;
<specimenCount id="9D5CE489FFA6F25F0DC2F9FFFE9CF998" box="[194,295,1594,1619]" pageId="1" pageNumber="562" type="female">1 female</specimenCount>
, Sarufutsu-mura, Asajino, outlet of Kamuito-numa,
<date id="FFE409C0FFA6F25F0E7AF9FEFC46F998" box="[890,1021,1594,1619]" pageId="1" pageNumber="562" value="2007-07-31">31.vii.2007</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0942F9FFFB26F998" box="[1090,1181,1594,1619]" pageId="1" pageNumber="562" type="male">3 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F09AAF9FFFAA0F998" box="[1194,1307,1594,1619]" pageId="1" pageNumber="562" type="female">4 females</specimenCount>
, Sarufutsumura, pond nr. Sarufutsu-gawa,
<date id="FFE409C0FFA6F25F0F07F99AFD31F9BC" box="[519,650,1630,1655]" pageId="1" pageNumber="562" value="2007-07-31">31.vii.2007</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0FD0F99BFC82F9BC" box="[720,825,1630,1655]" pageId="1" pageNumber="562" type="male">10 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0E47F99BFC03F9BC" box="[839,952,1630,1655]" pageId="1" pageNumber="562" type="female">2 females</specimenCount>
, Sarufutsu-mura, Sarufutsu-2-gôsen-gawa, Shôbu-bashi,
<date id="FFE409C0FFA6F25F0C31F946FE0FF950" box="[305,436,1666,1691]" pageId="1" pageNumber="562" value="2007-07-31">31.vii.2007</date>
, NK &amp; TI. Rumoi:
<specimenCount id="9D5CE489FFA6F25F0F91F946FD50F950" box="[657,747,1666,1691]" pageId="1" pageNumber="562" type="male">6 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0FF7F946FCCFF950" box="[759,884,1666,1691]" pageId="1" pageNumber="562" type="female">15 females</specimenCount>
, Rumoi-shi, Ôwada, el.
<quantity id="4CA282E5FFA6F25F0986F947FB62F951" box="[1158,1241,1667,1691]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="1" pageNumber="562" unit="m" value="9.5" valueMax="10.0" valueMin="9.0">9-10 m</quantity>
,
<date id="FFE409C0FFA6F25F09E5F946FADBF950" box="[1253,1376,1666,1691]" pageId="1" pageNumber="562" value="2015-08-02">2.viii.2015</date>
, NK. Ishikari:
<specimenCount id="9D5CE489FFA6F25F0DFEF963FEE2F974" box="[254,345,1702,1727]" pageId="1" pageNumber="562" type="male">2 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0C67F963FE63F974" box="[359,472,1702,1727]" pageId="1" pageNumber="562" type="female">4 females</specimenCount>
,
<typeStatus id="54E191A2FFA6F25F0CE6F96CFDADF90B" box="[486,534,1704,1728]" pageId="1" pageNumber="562">type</typeStatus>
locality,
<date id="FFE409C0FFA6F25F0F83F962FD44F974" box="[643,767,1702,1727]" pageId="1" pageNumber="562" value="2005-08-05">5.viii.2005</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0E46F963FC2CF974" box="[838,919,1702,1727]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0EA5F962FBADF974" box="[933,1046,1702,1727]" pageId="1" pageNumber="562" type="female">6 females</specimenCount>
, ibid,
<date id="FFE409C0FFA6F25F0960F962FB51F974" box="[1120,1258,1702,1727]" pageId="1" pageNumber="562" value="2005-08-12">12.viii.2005</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0832F963FA23F974" box="[1330,1432,1702,1727]" pageId="1" pageNumber="562" type="female">1 female</specimenCount>
, ibid,
<date id="FFE409C0FFA6F25F0DD6F90EFEDBF928" box="[214,352,1738,1763]" pageId="1" pageNumber="562" value="2005-08-17">17.viii.2005</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0CAEF90FFDB9F928" box="[430,514,1738,1763]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0F13F90FFD33F928" box="[531,648,1738,1763]" pageId="1" pageNumber="562" type="female">2 females</specimenCount>
, ibid,
<date id="FFE409C0FFA6F25F0FD8F90EFCDAF928" box="[728,865,1738,1763]" pageId="1" pageNumber="562" value="2006-08-14">14.viii.2006</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0EB0F90FFBB8F928" box="[944,1027,1738,1763]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
, Chitose-shi, Shikotsu-ko, Okotan,
<date id="FFE409C0FFA6F25F0D97F92AFEA2F8CC" box="[151,281,1774,1799]" pageId="1" pageNumber="562" value="1990-07-29">29.vii.1990</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0C66F92BFE6BF8CC" box="[358,464,1774,1799]" pageId="1" pageNumber="562" type="female">1 female</specimenCount>
, Chitose-shi, Shikotsu-ko, Shikotsuko-onsen,
<date id="FFE409C0FFA6F25F0EEBF92AFBDDF8CC" box="[1003,1126,1774,1799]" pageId="1" pageNumber="562" value="2006-08-09">9.viii.2006</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F09B4F92AFAA8F8CC" box="[1204,1299,1774,1799]" pageId="1" pageNumber="562" type="male">6 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0824F92BFA23F8CC" box="[1316,1432,1774,1799]" pageId="1" pageNumber="562" type="female">2 females</specimenCount>
, Chitose-shi, Shikotsu-ko, Shishamonai,
<date id="FFE409C0FFA6F25F0F67F8D6FD58F8E0" box="[615,739,1810,1835]" pageId="1" pageNumber="562" value="2005-08-05">5.viii.2005</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0E30F8D7FC35F8E0" box="[816,910,1810,1835]" pageId="1" pageNumber="562" type="male">8 males</specimenCount>
,
<specimenCount id="9D5CE489FFA6F25F0E9EF8D7FB9BF8E0" box="[926,1056,1810,1835]" pageId="1" pageNumber="562" type="female">12 females</specimenCount>
, ibid,
<date id="FFE409C0FFA6F25F096EF8D6FB43F8E0" box="[1134,1272,1810,1835]" pageId="1" pageNumber="562" value="2006-08-10">10.viii.2006</date>
, NK;
<specimenCount id="9D5CE489FFA6F25F0845F8D7FA23F8E0" box="[1349,1432,1810,1835]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
, Eniwa-shi, Izari-gawa, el.
<quantity id="4CA282E5FFA6F25F0CBEF8F3FDBCF884" box="[446,519,1847,1871]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="562" unit="m" value="130.0">130 m</quantity>
,
<date id="FFE409C0FFA6F25F0F13F8F2FD35F884" box="[531,654,1846,1871]" pageId="1" pageNumber="562" value="2015-08-02">2.viii.2015</date>
, TI;
<specimenCount id="9D5CE489FFA6F25F0FC4F8F3FCA9F884" box="[708,786,1846,1871]" pageId="1" pageNumber="562" type="male">1 male</specimenCount>
, Eniwa-shi, Izari-gawa, el.
<quantity id="4CA282E5FFA6F25F0945F8F3FB35F884" box="[1093,1166,1847,1871]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="562" unit="m" value="170.0">170 m</quantity>
,
<date id="FFE409C0FFA6F25F0999F8F2FAA7F884" box="[1177,1308,1846,1871]" pageId="1" pageNumber="562" value="2015-07-17">17.vii.2015</date>
, TI.
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA6F25F0DC7F89EFCABF8B8" box="[199,784,1882,1907]" pageId="1" pageNumber="562" type="etymology">
<paragraph id="8BE52F00FFA6F25F0DC7F89EFCABF8B8" blockId="1.[151,1437,151,2015]" box="[199,784,1882,1907]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7F89EFEFDF8B8" bold="true" box="[199,326,1882,1907]" pageId="1" pageNumber="562">Etymology</emphasis>
. Named for its distribution in Hokkaidô.
</paragraph>
</subSubSection>
<subSubSection id="C3407C8BFFA6F25F0DC7F8BAFC27F814" pageId="1" pageNumber="562" type="distribution">
<paragraph id="8BE52F00FFA6F25F0DC7F8BAFD97F85C" blockId="1.[151,1437,151,2015]" box="[199,556,1918,1943]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7F8BAFEE2F85C" bold="true" box="[199,345,1918,1943]" pageId="1" pageNumber="562">Distribution</emphasis>
.
<collectingCountry id="F34D6F90FFA6F25F0C67F844FE10F85C" box="[359,427,1920,1943]" name="Japan" pageId="1" pageNumber="562">Japan</collectingCountry>
: Hokkaidô.
</paragraph>
<paragraph id="8BE52F00FFA6F25F0DC7F866FC27F814" blockId="1.[151,1437,151,2015]" pageId="1" pageNumber="562">
<emphasis id="B92EF312FFA6F25F0DC7F866FE98F870" bold="true" box="[199,291,1954,1979]" pageId="1" pageNumber="562">Habitat</emphasis>
. Adults of this species have been collected beside slow flowing streams or rivers and shorelines of a pond in the Kushiro Marsh, Kushiro, and Lake Shikotsu-ko, Ishikari.
</paragraph>
</subSubSection>
<caption id="DF257F88FFA5F25C0D97F8BAFA32F83B" httpUri="https://zenodo.org/record/265301/files/figure.png" pageId="2" pageNumber="563" targetBox="[162,1413,197,1880]" targetPageId="2">
<paragraph id="8BE52F00FFA5F25C0D97F8BAFA32F83B" blockId="2.[151,1437,1918,2035]" pageId="2" pageNumber="563">
<emphasis id="B92EF312FFA5F25C0D97F8BAFEA8F858" bold="true" box="[151,275,1918,1940]" pageId="2" pageNumber="563">FIGURE 1.</emphasis>
<taxonomicName id="4C5A5483FFA5F25C0C19F8BBFE44F85F" box="[281,511,1918,1940]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="B92EF312FFA5F25C0C19F8BBFE44F85F" box="[281,511,1918,1940]" italics="true" pageId="2" pageNumber="563">Ecnomus hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4E69FFA5F25C0F05F8BAFDEEF858" box="[517,597,1918,1939]" pageId="2" pageNumber="563" rank="species">
<emphasis id="B92EF312FFA5F25C0F05F8BAFDEEF858" bold="true" box="[517,597,1918,1939]" pageId="2" pageNumber="563">sp. nov.</emphasis>
</taxonomicNameLabel>
(type locality; 1D1F, holotype). 1A, head, prothorax and mesothorax, dorsal; 1B, head, frontal; 1C, right wings, venation, dorsal; 1D1F, male genitalia: 1D, left lateral; 1E, dorsal; 1F, ventral; 1G1H, male phallus and inferior appendages: 1G, dorsal; 1H, left lateral; 1I1J, female genitalia: 1I, left lateral; 1J, ventral. Abbreviations: DC = discoidal cell; MC = median cell; TC = thyridial cell; m-cu = crossvein
<emphasis id="B92EF312FFA5F25C0EACF81FFC64F83B" box="[940,991,2011,2032]" italics="true" pageId="2" pageNumber="563">m-cu</emphasis>
; f R
<emphasis id="B92EF312FFA5F25C090BF823FBA9F838" bold="true" box="[1035,1042,2023,2035]" pageId="2" pageNumber="563">1</emphasis>
= fork of R
<emphasis id="B92EF312FFA5F25C098AF823FB2AF838" bold="true" box="[1162,1169,2023,2035]" pageId="2" pageNumber="563">1</emphasis>
; F1F5 = fork 1fork 5.
</paragraph>
</caption>
<subSubSection id="C3407C8BFFA4F25D0DC7FF53FB08FF33" pageId="3" pageNumber="564" type="discussion">
<paragraph id="8BE52F00FFA4F25D0DC7FF53FB08FF33" blockId="3.[151,1436,151,248]" pageId="3" pageNumber="564">
<emphasis id="B92EF312FFA4F25D0DC7FF53FE8FFF7B" bold="true" box="[199,308,151,176]" pageId="3" pageNumber="564">Remarks</emphasis>
. The downward curved phallus with spoon-shaped apex indicates this species belongs to the
<taxonomicName id="4C5A5483FFA4F25D0884FF5DFF55FF1F" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="B92EF312FFA4F25D0884FF5DFF55FF1F" italics="true" pageId="3" pageNumber="564">E. tenellus</emphasis>
</taxonomicName>
Group of
<bibRefCitation id="EFCB52F1FFA4F25D0C69FF78FDF3FF1F" author="Li" box="[361,584,188,213]" pageId="3" pageNumber="564" refString="Li, Y. J. &amp; Morse, J. C. (1997) Species of the genus Ecnomus (Trichoptera: Ecnomidae) from the People's Republic of China. Transactions of the American Entomological Society, 123, 85 - 134." type="journal article" year="1997">Li &amp; Morse (1997)</bibRefCitation>
. This is the fourth species in the Group distributed in the East Palaearctic region, as a result of
<taxonomicName id="4C5A5483FFA4F25D0C82FF25FE5EFF33" box="[386,485,224,248]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="tsudai">
<emphasis id="B92EF312FFA4F25D0C82FF25FE5EFF33" box="[386,485,224,248]" italics="true" pageId="3" pageNumber="564">E. tsudai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A5483FFA4F25D0F1CFF25FD02FF33" box="[540,697,224,248]" class="Insecta" family="Ecnomidae" genus="Ecnomus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="kososiensis">
<emphasis id="B92EF312FFA4F25D0F1CFF25FD02FF33" box="[540,697,224,248]" italics="true" pageId="3" pageNumber="564">E. kososiensis</emphasis>
</taxonomicName>
becoming invalid names as discussed below.
</paragraph>
</subSubSection>
</treatment>
</document>