163 lines
20 KiB
XML
163 lines
20 KiB
XML
<document id="077B44368FD34D71289BDD2835742C58" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D5852EFF9C5991FD775675FF01" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) cangalha Silva, Louzada & Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="480" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="479" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="4A0F2BDEBE650BDB5B53045E995CA4B1">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="13864C049C4C151F35108B3E5D5008BB">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="B96976D00EF88369EF89CA5D5669854C">Louzada, Júlio</mods:namePart>
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<mods:namePart id="8FAFE2BDBAC9625D698B1A55565098C0">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="9A02F983D9DF957FCC9B8C95A2E64276">Zootaxa</mods:title>
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<mods:date id="426402C40A9014C0A7844219666E031A">2015</mods:date>
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<mods:number id="13AB72B51F0E18A36985CE1DCEDA985C">3925</mods:number>
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<treatment id="03DE87D5852EFF9C5991FD775675FF01" ID-DOI="http://doi.org/10.5281/zenodo.6115734" ID-GBIF-Taxon="119598938" ID-Zenodo-Dep="6115734" LSID="urn:lsid:plazi:treatment:03DE87D5852EFF9C5991FD775675FF01" httpUri="http://treatment.plazi.org/id/03DE87D5852EFF9C5991FD775675FF01" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
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<subSubSection id="C36D6548852EFF9D5991FD7754FDFD1E" pageId="28" pageNumber="479" type="nomenclature">
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<paragraph id="8BC836C3852EFF9D5991FD7757B8FD7C" blockId="28.[151,759,654,714]" box="[151,759,654,680]" pageId="28" pageNumber="479">
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<heading id="D08081AF852EFF9D5991FD7757B8FD7C" bold="true" box="[151,759,654,680]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
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<emphasis id="B903EAD1852EFF9D5991FD7757B8FD7C" bold="true" box="[151,759,654,680]" pageId="28" pageNumber="479">
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<taxonomicName id="4C774D40852EFF9D5991FD77572FFD7C" ID-CoL="6CJ5T" box="[151,608,654,680]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="cangalha" status="sp. nov." subGenus="Aganhyboma">
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<emphasis id="B903EAD1852EFF9D5991FD775462FD7C" bold="true" box="[151,301,654,680]" italics="true" pageId="28" pageNumber="479">Deltochilum</emphasis>
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(
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<emphasis id="B903EAD1852EFF9D583BFD775491FD7C" bold="true" box="[317,478,654,680]" italics="true" pageId="28" pageNumber="479">Aganhyboma</emphasis>
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)
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<emphasis id="B903EAD1852EFF9D58F6FD77572FFD7C" bold="true" box="[496,608,654,680]" italics="true" pageId="28" pageNumber="479">cangalha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057AA852EFF9D5B6EFD7757B8FD7C" box="[616,759,654,680]" pageId="28" pageNumber="479" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836C3852EFF9D5991FD4854FDFD1E" blockId="28.[151,759,654,714]" box="[151,434,689,714]" pageId="28" pageNumber="479">
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(
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<figureCitation id="134C2A46852EFF9D5999FD4855BAFD1E" box="[159,245,689,714]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="28" pageNumber="479">Figs 10</figureCitation>
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, 76, 78, 92, 114)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852EFF9D5991FD005039FCE2" pageId="28" pageNumber="479" type="etymology">
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<paragraph id="8BC836C3852EFF9D5991FD005039FCE2" blockId="28.[151,1437,761,1974]" pageId="28" pageNumber="479">
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<emphasis id="B903EAD1852EFF9D5991FD005451FCC6" bold="true" box="[151,286,761,786]" pageId="28" pageNumber="479">Etymology.</emphasis>
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The new species name, "
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<taxonomicName id="4C774D40852EFF9D5B3EFD0057ECFCC6" box="[568,675,761,786]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="cangalha">cangalha</taxonomicName>
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", is a typical term of the northeastern region of
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<collectingCountry id="F3607653852EFF9D5DB1FD0051B1FCC6" box="[1207,1278,761,786]" name="Brazil" pageId="28" pageNumber="479">Brazil</collectingCountry>
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, which means "bowlegged", a characteristic usually found in individuals of
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<taxonomicName id="4C774D40852EFF9D5A43FCE45696FCE1" box="[837,985,797,821]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EAD1852EFF9D5A43FCE45696FCE1" box="[837,985,797,821]" italics="true" pageId="28" pageNumber="479">Aganhyboma</emphasis>
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</taxonomicName>
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. The epithet is a name in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852EFF9D59C1FCB854F5F9FE" pageId="28" pageNumber="479" type="description">
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<paragraph id="8BC836C3852EFF9D59C1FCB854F5F9FE" blockId="28.[151,1437,761,1974]" pageId="28" pageNumber="479">
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<emphasis id="B903EAD1852EFF9D59C1FCB854EAFC8E" bold="true" box="[199,421,833,858]" pageId="28" pageNumber="479">Description. Body.</emphasis>
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Head, pronotum, elytra, and ventral surface black, with a copper-red sheen. Length>
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<quantity id="4C8F9B26852EFF9D5C86FCBB5589FCA9" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="479" unit="mm" value="14.0">14 mm</quantity>
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.
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<emphasis id="B903EAD1852EFF9D59D4FC9C5456FCAA" bold="true" box="[210,281,869,894]" pageId="28" pageNumber="479">Head.</emphasis>
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Punctures extremely dense and touching; around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae, or with almost inconspicuous setae. Dorsal interocular distance approximately three times eye width (as in Fig 61).
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<emphasis id="B903EAD1852EFF9D5C3AFC5450D3FC12" bold="true" box="[1340,1436,941,966]" pageId="28" pageNumber="479">Thorax.</emphasis>
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Pronotum with a red metallic sheen (
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<figureCitation id="134C2A46852EFF9D5B55FC2857F6FC3E" box="[595,697,977,1002]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="28" pageNumber="479">Fig. 114</figureCitation>
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). Surface of pronotum with dense foveiform punctures, nearly touching on the sides, anterior angles and posterior portion. Discal punctures smaller and more dispersed than those of lateral portion. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by less than one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles approximately straight. Hypomera with moderate foveiform punctures on anterior and posterior portions (Fig. 78); anteromedial punctures separated by at least one diameter (Fig. 78).
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<emphasis id="B903EAD1852EFF9D5869FB50548DFB16" bold="true" box="[367,450,1193,1218]" pageId="28" pageNumber="479">Elytra.</emphasis>
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Elytra rounded, sides bulging outward (
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<figureCitation id="134C2A46852EFF9D5A85FB5056AFFB16" box="[899,992,1193,1218]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="28" pageNumber="479">Fig. 114</figureCitation>
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) and surface slightly irregular (rugose). Elytral striae narrow, carinate margins close and almost touching (as in Figs 74, 76). Interstriae flattened, with foveiform punctures surrounded by shiny areas poorly defined. Foveiform punctures of interstriae separated on elytral disc by less than one diameter. Microtubercles of interstriae dense and weakly defined on most of surface, united by glossy spots with a red-cupreous sheen (Fig. 76). Basal carina of seventh interstria short, length about twice the width of the seventh interestria. Basal carina of ninth interstria present in three-quarters of elytra, but it never reaches the anterior limit of apical carina of seventh interstria. Apical carina of third interstria somewhat more prominent than apical carinae of second and fourth interstriae. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria less than the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852EFF9D59C1F9CC51D7F96E" pageId="28" pageNumber="479" type="materials_examined">
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<paragraph id="8BC836C3852EFF9D59C1F9CC54D2F9A6" blockId="28.[151,1437,761,1974]" pageId="28" pageNumber="479">
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<emphasis id="B903EAD1852EFF9D59C1F9CC57CCF99A" bold="true" box="[199,643,1589,1614]" pageId="28" pageNumber="479">
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Material studied.
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<typeStatus id="54CC8861852EFF9D589DF9CC5743F99A" box="[411,524,1589,1614]" pageId="28" pageNumber="479" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3607653852EFF9D5B1EF9CC57CCF99A" box="[536,643,1589,1614]" name="Brazil" pageId="28" pageNumber="479">BRAZIL</collectingCountry>
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</emphasis>
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:
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<collectionCode id="ED66AE06852EFF9D5B97F9CF57A4F99A" box="[657,747,1590,1614]" pageId="28" pageNumber="479">MINAS</collectionCode>
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GERAIS: Águas Vermelhas,
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<date id="FFC91003852EFF9D5D3EF9CF51EEF99A" box="[1080,1185,1590,1614]" pageId="28" pageNumber="479" value="1998-12">XII.1998</date>
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, A. Bello & F. Z. Vazde-Mello (
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<specimenCount id="9D71FD4A852EFF9D5816F9A35477F9A5" box="[272,312,1626,1650]" pageId="28" pageNumber="479" type="generic">1 ♂</specimenCount>
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<collectionCode id="ED66AE06852EFF9D5839F9A354DDF9A6" box="[319,402,1626,1650]" pageId="28" pageNumber="479">CEMT</collectionCode>
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).
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</paragraph>
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<paragraph id="8BC836C3852EFF9D59C1F98451D7F96E" blockId="28.[151,1437,761,1974]" pageId="28" pageNumber="479">
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<emphasis id="B903EAD1852EFF9D59C1F9845465F942" bold="true" box="[199,298,1661,1686]" pageId="28" pageNumber="479">Habitat.</emphasis>
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<collectingCountry id="F3607653852EFF9D5832F9845437F942" box="[308,376,1661,1686]" name="Brazil" pageId="28" pageNumber="479">Brazil</collectingCountry>
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(Minas Gerais) (
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<figureCitation id="134C2A46852EFF9D5B46F98457D8F942" box="[576,663,1661,1686]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="28" pageNumber="479">Fig. 92</figureCitation>
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), from available data this species inhabits “Carrasco”. This habitat consists of dense shrubby xerophilous vegetation that occurs in semi-arid domain of
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<collectingCountry id="F3607653852EFF9D5D4AF95851DBF96E" box="[1100,1172,1697,1722]" name="Brazil" pageId="28" pageNumber="479">Brazil</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852EFF9C59C1F93C5675FF01" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479" type="diagnosis">
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<paragraph id="8BC836C3852EFF9D59C1F93C55A1F862" blockId="28.[151,1437,761,1974]" pageId="28" pageNumber="479">
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<emphasis id="B903EAD1852EFF9D59C1F93C54ADF90A" bold="true" box="[199,482,1733,1758]" pageId="28" pageNumber="479">Diagnosis and remarks</emphasis>
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(within the
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<taxonomicName id="4C774D40852EFF9D5B73F93C57B3F909" box="[629,764,1733,1757]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="finestriatum">
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<emphasis id="B903EAD1852EFF9D5B73F93C57B3F909" box="[629,764,1733,1757]" italics="true" pageId="28" pageNumber="479">finestriatum</emphasis>
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</taxonomicName>
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complex): pronotal margin, between lateral and posterior angles, straight; Elytra rounded, sides bulging outward (
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<figureCitation id="134C2A46852EFF9D5A15F9105639F8D6" box="[787,886,1769,1794]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="28" pageNumber="479">Fig. 114</figureCitation>
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); anteromedial punctures of hypomera separated by at least one diameter (Fig. 78); pronotum with a red metallic sheen; elytral microtubercles weakly defined, with a red-cupreous sheen (Fig. 76). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, in lateral view forming an angle of approximately 45° with phallobase; apex rounded (
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<figureCitation id="134C2A46852EFF9D5DD1F8AC5064F8BA" box="[1239,1323,1877,1902]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="28" pageNumber="479">Fig. 10</figureCitation>
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). Lamella copulatrix (LC): left lobe subtrapezoidal; right lobe subrectangular. Fronto-lateral peripheral sclerite (FLP) as in Fig. 53.
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</paragraph>
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<paragraph id="8BC836C3852FFF9C59C1FF6E5675FF01" blockId="29.[151,1436,151,213]" pageId="29" pageNumber="480">
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Although the aedeagus of
|
||
<taxonomicName id="4C774D40852FFF9C58F2FF6057CFFF64" box="[500,640,152,176]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="cangalha" status="sp. nov.">
|
||
<emphasis id="B903EAD1852FFF9C58F2FF6057CFFF64" box="[500,640,152,176]" italics="true" pageId="29" pageNumber="480">D. cangalha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B903EAD1852FFF9C5B8FFF6E565AFF64" bold="true" box="[649,789,151,176]" pageId="29" pageNumber="480">
|
||
<taxonomicNameLabel id="A23057AA852FFF9C5B8FFF6E565AFF64" box="[649,789,151,176]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C774D40852FFF9C5A49FF605691FF64" box="[847,990,152,176]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="alpercata" status="sp. nov.">
|
||
<emphasis id="B903EAD1852FFF9C5A49FF605691FF64" box="[847,990,152,176]" italics="true" pageId="29" pageNumber="480">D. alpercata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B903EAD1852FFF9C5AE0FF6E513DFF64" bold="true" box="[998,1138,151,176]" pageId="29" pageNumber="480">
|
||
<taxonomicNameLabel id="A23057AA852FFF9C5AE0FF6E513DFF64" box="[998,1138,151,176]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
do not present significant difference, the elytral punctures in these species are distinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |