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<document id="EAA67F9612028C3A6DB9A0703ECD92D5" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D5852BFF9B5991FC5655A1FD19" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) acanthus Kohlmann &amp; Solis 2012" docType="treatment" docVersion="9" lastPageNumber="477" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="476" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="AD432F6D2C228C1C8F5A76F5E5ACEADE">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="9AE90B088D6D14DDD4648E38F6080480">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="7F30E2936F87DD29D239FBA4A1CD79D5">Louzada, Júlio</mods:namePart>
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<mods:namePart id="31388F4BFA7CD5EA24AF63CDCCC045D2">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="EB16498900EEAB35E8307AB564127D30">Zootaxa</mods:title>
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<mods:date id="5EA62EBDE57DF7BE133DE01BBC1F24A5">2015</mods:date>
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<treatment id="03DE87D5852BFF9B5991FC5655A1FD19" ID-DOI="http://doi.org/10.5281/zenodo.3510634" ID-GBIF-Taxon="119598944" ID-Zenodo-Dep="3510634" LSID="urn:lsid:plazi:treatment:03DE87D5852BFF9B5991FC5655A1FD19" httpUri="http://treatment.plazi.org/id/03DE87D5852BFF9B5991FC5655A1FD19" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476">
<subSubSection id="C36D6548852BFF985991FC565413FC3F" pageId="25" pageNumber="476" type="nomenclature">
<paragraph id="8BC836C3852BFF985991FC5656C0FC1E" blockId="25.[151,911,943,1003]" box="[151,911,943,970]" pageId="25" pageNumber="476">
<heading id="D08081AF852BFF985991FC5656C0FC1E" bold="true" box="[151,911,943,970]" fontSize="11" level="1" pageId="25" pageNumber="476" reason="1">
<taxonomicName id="4C774D40852BFF985991FC5656C0FC1E" ID-CoL="6CJ62" authority="Kohlmann &amp; Solis 2012" authorityName="Kohlmann &amp; Solis" authorityYear="2012" box="[151,911,943,970]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="acanthus" subGenus="Aganhyboma">
<emphasis id="B903EAD1852BFF985991FC5656C0FC1E" bold="true" box="[151,911,943,970]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF985991FC565462FC1D" bold="true" box="[151,301,943,969]" italics="true" pageId="25" pageNumber="476">Deltochilum</emphasis>
(
<emphasis id="B903EAD1852BFF98583BFC565491FC1D" bold="true" box="[317,478,943,969]" italics="true" pageId="25" pageNumber="476">Aganhyboma</emphasis>
)
<emphasis id="B903EAD1852BFF9858F6FC565711FC1D" bold="true" box="[496,606,943,969]" italics="true" pageId="25" pageNumber="476">acanthus</emphasis>
<bibRefCitation id="EFE64B32852BFF985B60FC4956C0FC1E" author="Kohlmann" box="[614,911,944,970]" pageId="25" pageNumber="476" refString="Kohlmann, B. &amp; Solis, A. (2012) New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 3193, 28 - 52." type="journal article" year="2012">Kohlmann &amp; Solís 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC836C3852BFF985991FC2B5413FC3F" blockId="25.[151,911,943,1003]" box="[151,348,978,1003]" pageId="25" pageNumber="476">
(
<figureCitation id="134C2A46852BFF985999FC2B55BAFC3F" box="[159,245,978,1003]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="25" pageNumber="476">Figs 15</figureCitation>
,
<figureCitation id="134C2A46852BFF985807FC2A5450FC3F" box="[257,287,979,1003]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="25" pageNumber="476">91</figureCitation>
,
<figureCitation id="134C2A46852BFF98582DFC2A541BFC3F" box="[299,340,979,1003]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="25" pageNumber="476">111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852BFF985991FBE35018FBE4" box="[151,1367,1050,1072]" pageId="25" pageNumber="476" type="reference_group">
<paragraph id="8BC836C3852BFF985991FBE35018FBE4" blockId="25.[151,1367,1050,1072]" box="[151,1367,1050,1072]" pageId="25" pageNumber="476">
<treatmentCitationGroup id="AB6711ED852BFF985991FBE35018FBE4" box="[151,1367,1050,1072]" pageId="25" pageNumber="476">
<taxonomicName id="4C774D40852BFF985991FBE35674FBE4" ID-CoL="6CJ62" authority="Kohlmann &amp; Solis 2012: 35" authorityName="Kohlmann &amp; Solis" authorityPageNumber="35" authorityYear="2012" box="[151,827,1050,1072]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="acanthus" subGenus="Deltohyboma">
<emphasis id="B903EAD1852BFF985991FBE3545BFBE4" box="[151,276,1050,1072]" italics="true" pageId="25" pageNumber="476">Deltochilum</emphasis>
(
<emphasis id="B903EAD1852BFF985825FBE354E4FBE4" box="[291,427,1050,1072]" italics="true" pageId="25" pageNumber="476">Deltohyboma</emphasis>
)
<emphasis id="B903EAD1852BFF9858BCFBE3575AFBE4" box="[442,533,1050,1072]" italics="true" pageId="25" pageNumber="476">acanthus</emphasis>
<treatmentCitation id="0AD610D2852BFF985B1AFBE35674FBE4" ID-DOI="http://doi.org/10.5281/zenodo.3499664" ID-Zenodo-Dep="3499664" author="Kohlmann" box="[540,827,1050,1072]" httpUri="http://treatment.plazi.org/id/7C0D87B7FFB8FFAB8987FF568F56F447" page="35" pageId="25" pageNumber="476" year="2012">
<bibRefCitation id="EFE64B32852BFF985B1AFBE35658FBE4" author="Kohlmann" box="[540,791,1050,1072]" pageId="25" pageNumber="476" refString="Kohlmann, B. &amp; Solis, A. (2012) New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 3193, 28 - 52." type="journal article" year="2012">Kohlmann &amp; Solís 2012</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
, 36, 37, 38,
<figureCitation id="134C2A46852BFF985ABEFBE35160FBE4" box="[952,1071,1050,1072]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="25" pageNumber="476">Figures 6, 7</figureCitation>
and
<figureCitation id="134C2A46852BFF985D67FBE35135FBE4" box="[1121,1146,1050,1072]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="25" pageNumber="476">15</figureCitation>
(original description)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852BFF985991FBA456C9F8BE" pageId="25" pageNumber="476" type="description">
<paragraph id="8BC836C3852BFF985991FBA456C9F8BE" blockId="25.[151,1437,1117,2006]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF985991FBA4543CFBA2" bold="true" box="[151,371,1117,1142]" pageId="25" pageNumber="476">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface black or dark brown, with an evident sheen. Length &lt;
<quantity id="4C8F9B26852BFF9859A8FB7B544DFB4D" box="[174,258,1154,1178]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="476" unit="mm" value="13.0">13 mm</quantity>
.
<emphasis id="B903EAD1852BFF98580BFB78541BFB4E" bold="true" box="[269,340,1153,1178]" pageId="25" pageNumber="476">Head.</emphasis>
Punctures dense, nearly touching at central portion; around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae or with almost inconspicuous setae. Dorsal interocular distance approximately 2.5 times eye width.
<emphasis id="B903EAD1852BFF985DBAFB305054FB36" bold="true" box="[1212,1307,1225,1250]" pageId="25" pageNumber="476">Thorax.</emphasis>
Surface of pronotum with dense foveiform punctures, not touching each other. Punctures on the sides, anterior and posterior angles separated by half diameter or less. Discal punctures smaller and more dispersed than those of lateral portion, separated by approximately two times their diameter. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures separated by less than one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles straight, with the same length of margin between anterior and lateral angles. Hypomera with dense foveiform punctures at anterior portion and absent at posteromedial portion; anteromedial punctures usually separated by less than one diameter. Diagonal band of hypomera from posteromedial portion to middle of lateral margin lacking punctures or with one or two isolated punctures on otherwise smooth surface (as in
<figureCitation id="134C2A46852BFF985BF0F9F45606F9F2" box="[758,841,1549,1574]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="25" pageNumber="476">Fig. 83</figureCitation>
).
<emphasis id="B903EAD1852BFF985A5BF9F456FEF9F2" bold="true" box="[861,945,1549,1574]" pageId="25" pageNumber="476">Elytra.</emphasis>
Lateral margins semiparallel (
<figureCitation id="134C2A46852BFF985C01F9F4502BF9F2" box="[1287,1380,1549,1574]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="25" pageNumber="476">Fig. 111</figureCitation>
) and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated; several punctures clearly elliptical, separated by regular spacing. Interstriae flattened, with foveiform punctures surrounded by approximately four well-defined shiny microtubercles. Foveiform punctures separated on elytral disc by approximately one diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (as in Fig. 80). Apical carina or tubercle of second interstria present. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria noticeably less than that of sum of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (as in Fig 80).
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852BFF9B59C1F88C5061FEB0" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="materials_examined">
<paragraph id="8BC836C3852BFF9859C1F88C56A1F802" blockId="25.[151,1437,1117,2006]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1F88C5742F85A" bold="true" box="[199,525,1909,1934]" pageId="25" pageNumber="476">
Material studied.
<typeStatus id="54CC8861852BFF9858A6F88C5742F85A" box="[416,525,1909,1934]" pageId="25" pageNumber="476" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="ED66AE06852BFF985B18F88C5728F85A" box="[542,615,1909,1934]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="25" pageNumber="476">INBio</collectionCode>
)
<emphasis id="B903EAD1852BFF985B7EF88C56A0F85A" bold="true" box="[632,1007,1909,1934]" pageId="25" pageNumber="476">
[not examined]:
<collectingCountry id="F3607653852BFF985A46F88C56A0F85A" box="[832,1007,1909,1934]" name="Costa Rica" pageId="25" pageNumber="476">COSTA RICA</collectingCountry>
</emphasis>
: Prov. Puntarenas, Res. Biol. Carara, Est. Quebrada Bonita,
<date id="FFC91003852BFF985890F86354B3F866" box="[406,508,1946,1970]" pageId="25" pageNumber="476" value="1993-06">
VI.
<quantity id="4C8F9B26852BFF9858C7F86354B3F866" box="[449,508,1946,1970]" metricMagnitude="3" metricUnit="m" metricValue="1.9934999999999998" pageId="25" pageNumber="476" unit="m" value="1993.5">1993</quantity>
</date>
, 50 m, J.C. Saborío, L-N-194500, 469850,
<collectionCode id="ED66AE06852BFF985AE5F860515EF866" LSID="urn:lsid:biocol.org:col:14323" box="[995,1041,1945,1970]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="25" pageNumber="476">CRI</collectionCode>
001185073 (
<specimenCount id="9D71FD4A852BFF985DA7F8635186F865" box="[1185,1225,1946,1970]" pageId="25" pageNumber="476" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE06852BFF985DD6F8605059F866" box="[1232,1302,1945,1970]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="25" pageNumber="476">INBio</collectionCode>
collection). The original description is sufficient to confirm the identification of species.
</paragraph>
<paragraph id="8BC836C38528FF9B59C1FF6E57F5FF2C" blockId="26.[151,1437,151,717]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1FF6E5715FF64" bold="true" box="[199,602,151,176]" pageId="26" pageNumber="477">
Non-type material [2]:
<collectingCountry id="F36076538528FF9B58E6FF6E5715FF64" box="[480,602,151,176]" name="Panama" pageId="26" pageNumber="477">PANAMA</collectingCountry>
</emphasis>
:
<collectingCountry id="F36076538528FF9B5B6AFF6157ADFF64" box="[620,738,152,176]" name="Panama" pageId="26" pageNumber="477">PANAMA</collectingCountry>
: Gatún Lake, Barro Colorado
<collectingCountry id="F36076538528FF9B5D43FF6E51C3FF64" box="[1093,1164,151,176]" name="Iceland" pageId="26" pageNumber="477">Island</collectingCountry>
,
<date id="FFC910038528FF9B5D9FFF6E504EFF64" box="[1177,1281,151,176]" pageId="26" pageNumber="477" value="1981-05-05">5.V.1981</date>
, B. Gill (
<specimenCount id="9D71FD4A8528FF9B5C77FF6150D3FF7B" box="[1393,1436,152,176]" pageId="26" pageNumber="477" type="generic">1 ♂</specimenCount>
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).
<emphasis id="B903EAD18528FF9B59F0FF4554E5FF01" bold="true" box="[246,426,188,213]" pageId="26" pageNumber="477">
<collectingCountry id="F36076538528FF9B59F0FF4554E5FF01" box="[246,426,188,213]" name="Costa Rica" pageId="26" pageNumber="477">COSTA RICA</collectingCountry>
</emphasis>
: PUNTARENAS: Osa Peninsula. Río Piro, la Joya
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,
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,
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,
<date id="FFC910038528FF9B5991FF195457FF2C" box="[151,280,224,248]" pageId="26" pageNumber="477" value="1997-05" valueMax="1997-09-31" valueMin="1997-05-00">V–IX.1997</date>
, N. Gibson (A. Forsyth) (1 ♀
<collectionCode id="ED66AE068528FF9B5B6FFF1957E1FF2C" box="[617,686,224,248]" pageId="26" pageNumber="477">AFIC</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38528FF9B59C1FEFD5601FEC9" blockId="26.[151,1437,151,717]" box="[199,846,260,285]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1FEFD54E3FEC9" bold="true" box="[199,428,260,285]" pageId="26" pageNumber="477">Sampling methods.</emphasis>
flight interception trap [
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].
</paragraph>
<paragraph id="8BC836C38528FF9B59C1FEDE5061FEB0" blockId="26.[151,1437,151,717]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1FEDE5465FE94" bold="true" box="[199,298,295,320]" pageId="26" pageNumber="477">Habitat.</emphasis>
<collectingCountry id="F36076538528FF9B5833FED154C1FEEB" box="[309,398,296,319]" name="Panama" pageId="26" pageNumber="477">Panama</collectingCountry>
and
<collectingCountry id="F36076538528FF9B58CBFED15705FE94" box="[461,586,295,320]" name="Costa Rica" pageId="26" pageNumber="477">Costa Rica</collectingCountry>
(
<figureCitation id="134C2A468528FF9B5B58FEDE57F9FE94" box="[606,694,295,320]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="26" pageNumber="477">Fig. 91</figureCitation>
). According to
<bibRefCitation id="EFE64B328528FF9B5A77FEDE51EBFE94" author="Kohlmann" box="[881,1188,295,320]" pageId="26" pageNumber="477" refString="Kohlmann, B. &amp; Solis, A. (2012) New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 3193, 28 - 52." type="journal article" year="2012">Kohlmann &amp; Solis (2012)</bibRefCitation>
, this species inhabits Pacific tropical rain forest of
<collectingCountry id="F36076538528FF9B58E6FEB55716FEB1" box="[480,601,332,357]" name="Costa Rica" pageId="26" pageNumber="477">Costa Rica</collectingCountry>
and the Canal Zone of
<collectingCountry id="F36076538528FF9B5A67FEB456F2FEB0" box="[865,957,333,356]" name="Panama" pageId="26" pageNumber="477">Panama</collectingCountry>
, ranging from
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<collectionCode id="ED66AE068528FF9B5DE7FEB55068FEB7" box="[1249,1319,332,355]" pageId="26" pageNumber="477">AMSL</collectionCode>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488528FF9B59C1FE9655A1FD19" pageId="26" pageNumber="477" type="diagnosis">
<paragraph id="8BC836C38528FF9B59C1FE9655A1FD19" blockId="26.[151,1437,151,717]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1FE9654ADFE5C" bold="true" box="[199,482,367,392]" pageId="26" pageNumber="477">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408528FF9B5B73FE885793FE5C" box="[629,732,369,392]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD18528FF9B5B73FE885793FE5C" box="[629,732,369,392]" italics="true" pageId="26" pageNumber="477">acropyge</emphasis>
</taxonomicName>
complex): foveiform punctures of dorsal interocular surface nearly touching; foveiform punctures of pronotal disc separated by approximately twice their diameter; posteromedial punctures of pronotum usually separated by less than one diameter; pronotal margin, between lateral and posterior angles, with approximately the same length of margin between anterior and lateral angles; elytra with lateral margins semiparallel (
<figureCitation id="134C2A468528FF9B58E1FE065705FDCC" box="[487,586,511,536]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="26" pageNumber="477">Fig. 111</figureCitation>
); punctures of elytral striae often elliptical, separated by regular spacing; apical carina or tubercle of second elytral interstria present; pseudepipleural carina incomplete, effaced medially (as in Fig 80). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, length approximately equal to phallobase; in lateral view forming an angle of approximately 45° with phallobase; apex rounded (
<figureCitation id="134C2A468528FF9B5805FD765418FD7C" box="[259,343,655,680]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="26" pageNumber="477">Fig. 15</figureCitation>
). Lamella copulatrix (LC): subrectangular sclerites. Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.
</paragraph>
</subSubSection>
</treatment>
</document>
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