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<document id="1CBEF1B571489DEBC7CC6BB2E8AB2309" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58529FF9D5991FE255601FE5C" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) larseni Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="478" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="478" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="458813C632783908E62FD395A943E96B">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="066F2E44C1A0741AFB6C5D0A11DF1560">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="06E08F384151BB862D26B03228811D15">Louzada, Júlio</mods:namePart>
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<mods:namePart id="395CAC94DF054C16AD9853C253C60314">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:date id="AEFEDF64A03367D553C7133B6C7A92CF">2015</mods:date>
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<treatment id="03DE87D58529FF9D5991FE255601FE5C" ID-DOI="http://doi.org/10.5281/zenodo.6115730" ID-GBIF-Taxon="119598946" ID-Zenodo-Dep="6115730" LSID="urn:lsid:plazi:treatment:03DE87D58529FF9D5991FE255601FE5C" httpUri="http://treatment.plazi.org/id/03DE87D58529FF9D5991FE255601FE5C" lastPageId="28" lastPageNumber="478" pageId="27" pageNumber="478">
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<paragraph id="8BC836C38529FF9A5991FE255795FE22" blockId="27.[151,730,476,535]" box="[151,730,476,502]" pageId="27" pageNumber="478">
<heading id="D08081AF8529FF9A5991FE255795FE22" bold="true" box="[151,730,476,502]" fontSize="11" level="1" pageId="27" pageNumber="478" reason="1">
<emphasis id="B903EAD18529FF9A5991FE255795FE22" bold="true" box="[151,730,476,502]" pageId="27" pageNumber="478">
<taxonomicName id="4C774D408529FF9A5991FE25570CFE22" ID-CoL="6CJ5J" box="[151,579,476,502]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="larseni" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18529FF9A5991FE255462FE22" bold="true" box="[151,301,476,502]" italics="true" pageId="27" pageNumber="478">Deltochilum</emphasis>
(
<emphasis id="B903EAD18529FF9A583BFE255491FE22" bold="true" box="[317,478,476,502]" italics="true" pageId="27" pageNumber="478">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18529FF9A58F6FE25570CFE22" bold="true" box="[496,579,476,502]" italics="true" pageId="27" pageNumber="478">larseni</emphasis>
</taxonomicName>
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</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38529FF9A5991FE075412FDC3" blockId="27.[151,730,476,535]" box="[151,349,510,535]" pageId="27" pageNumber="478">
(
<figureCitation id="134C2A468529FF9A5999FE0755BAFDC3" box="[159,245,510,535]" captionStart="FIGURES 12 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="27" pageNumber="478">Figs 14</figureCitation>
,
<figureCitation id="134C2A468529FF9A5807FE065450FDC3" box="[257,287,511,535]" captionStart="FIGURES 89 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="27" pageNumber="478">91</figureCitation>
,
<figureCitation id="134C2A468529FF9A582DFE06541AFDC3" box="[299,341,511,535]" captionStart="FIGURES 111 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="27" pageNumber="478">113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488529FF9A5991FDBF5404FD57" pageId="27" pageNumber="478" type="etymology">
<paragraph id="8BC836C38529FF9A5991FDBF5404FD57" blockId="27.[151,1437,582,2011]" pageId="27" pageNumber="478">
<emphasis id="B903EAD18529FF9A5991FDBF5451FD8B" bold="true" box="[151,286,582,607]" pageId="27" pageNumber="478">Etymology.</emphasis>
A patronym honoring Trond Larsen, great and enthousiatic scarab ecologist who collected most of the
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specimens.
</paragraph>
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<subSubSection id="C36D65488529FF9A59C1FD775020FAA3" pageId="27" pageNumber="478" type="description">
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<emphasis id="B903EAD18529FF9A59C1FD7754EAFD73" bold="true" box="[199,421,654,679]" pageId="27" pageNumber="478">Description. Body.</emphasis>
Black or dark brown coloration, with obvious sheen (
<figureCitation id="134C2A468529FF9A5D08FD77513EFD73" box="[1038,1137,654,679]" captionStart="FIGURES 111 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="27" pageNumber="478">Fig. 113</figureCitation>
). Length &lt;
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.
<emphasis id="B903EAD18529FF9A5C53FD7750D3FD73" bold="true" box="[1365,1436,654,679]" pageId="27" pageNumber="478">Head.</emphasis>
Punctures dense, nearly touching; around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae or with almost inconspicuous setae. Dorsal interocular distance approximately 2.5 times eye width.
<emphasis id="B903EAD18529FF9A5A5CFD0356FCFCC7" bold="true" box="[858,947,762,787]" pageId="27" pageNumber="478">Thorax</emphasis>
. Surface of pronotum with dense foveiform punctures. Punctures on the sides, anterior and posterior angles nearly touching. Discal punctures smaller and more dispersed than those of lateral portion, separated approximately by one diameter. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by at least one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles with length somewhat shorter than that of margin between anterior and lateral angles. Hypomera with dense foveiform punctures on anterior portion and dispersed on posteromedial portion; anteromedial punctures almost touching. Diagonal band of hypomera from posteromedial portion to middle of lateral margin with dispersed punctures on otherwise smooth surface.
<emphasis id="B903EAD18529FF9A5B6FFBE357F3FBE7" bold="true" box="[617,700,1050,1075]" pageId="27" pageNumber="478">Elytra.</emphasis>
Lateral margins slightly curved outward (
<figureCitation id="134C2A468529FF9A5DA3FBE35048FBE7" box="[1189,1287,1050,1075]" captionStart="FIGURES 111 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="27" pageNumber="478">Fig. 113</figureCitation>
) and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated; strial punctures separated by regular spacing; most of punctures round intermingled with some slightly elliptical. Interstriae flattened, with foveiform punctures surrounded by approximately five obvious shiny microtubercles. Foveiform punctures separated on elytral disc by approximately one diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (as in Fig. 80). Apical carina or tubercle of second elytral interstria absent. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria longer than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (as in Fig. 80).
</paragraph>
</subSubSection>
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Material studied.
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:
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</emphasis>
: SUCUMBIOS: R.P.F. Cuyabeno, Trocha Zábalo-Güepí, km 10,
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.VIII.2010, Colección Manual nocturna, Bosque de tierra firme colinado, Pablo Araujo,
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557 (
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).
</paragraph>
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[15]:
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: SUCUMBIOS: R.P.F. Cuyabeno, Trocha Zábalo-Güepí, km 10,
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.VIII.2010, Colección Manual nocturna, Bosque de tierra firme colinado, Pablo Araujo,
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); PASTAZA: B.P. Oglán, T4, Bsvpm. 554 msnm,
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E 9853376N,
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, Col. W. Chamorro &amp; V. Guasumba (1 ♀
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).
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<collectingCountry id="F36076538529FF9A59FAF9CF5403F99B" box="[252,332,1590,1615]" name="Peru" pageId="27" pageNumber="478">PERU</collectingCountry>
:
</emphasis>
valle cerca Abancay,
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, C.S. Carbonell (
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); Upper Rio Marañón,
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(
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);
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: Río Tambopata,
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, Ccolpa de Guacamayos,
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,
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, A Forsyth,
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(
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); Rio Madre de Dios, Rio Los Amigos, base camp, 1° terra firme,
<quantity id="4C8F9B268529FF9A5DFBF9865005F942" box="[1277,1354,1663,1687]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="27" pageNumber="478" unit="m" value="290.0">290 m</quantity>
, pitfall crushed but live spirobolid millipede,
<geoCoordinate id="EE4350048529FF9A5B46F95A5798F96F" box="[576,727,1699,1723]" direction="south" orientation="latitude" pageId="27" pageNumber="478" precision="1" value="-12.569445">12°34'10.0&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048529FF9A5BE5F95B56CDF96F" box="[739,898,1698,1723]" direction="west" orientation="longitude" pageId="27" pageNumber="478" precision="1" value="-70.10039">70°06'01.4&quot;W</geoCoordinate>
, T. Larsen,
<date id="FFC910038529FF9A5D02F95A5112F96F" box="[1028,1117,1699,1723]" pageId="27" pageNumber="478" value="2000-04">IV.2000</date>
(
<specimenCount id="9D71FD4A8529FF9A5D69F95A51D8F96E" box="[1135,1175,1699,1723]" pageId="27" pageNumber="478" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068529FF9A5D98F95A51A2F96F" box="[1182,1261,1699,1723]" pageId="27" pageNumber="478">CEMT</collectionCode>
,
<specimenCount id="9D71FD4A8529FF9A5DFEF95A5055F96F" box="[1272,1306,1699,1723]" pageId="27" pageNumber="478" type="generic">1♂</specimenCount>
1♀
<collectionCode id="ED66AE068529FF9A5C4DF95A50DFF96F" box="[1355,1424,1699,1723]" pageId="27" pageNumber="478">AFIC</collectionCode>
); same but
<date id="FFC910038529FF9A5802F93E54E5F90B" box="[260,426,1735,1759]" pageId="27" pageNumber="478" value="2000-04-12" valueMax="2000-04-13" valueMin="2000-04-12">1213.IV.2000</date>
(
<specimenCount id="9D71FD4A8529FF9A58BAF93E5490F90B" box="[444,479,1735,1759]" pageId="27" pageNumber="478" type="generic">1♂</specimenCount>
<collectionCode id="ED66AE068529FF9A58E1F93E5775F90B" box="[487,570,1735,1759]" pageId="27" pageNumber="478">CEMT</collectionCode>
); same but Rio Amiguillos, small rever flood plain,
<quantity id="4C8F9B268529FF9A5D8FF93F519CF90A" box="[1161,1235,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="27" pageNumber="478" unit="m" value="260.0">260 m</quantity>
, pitfall millipede,
<geoCoordinate id="EE4350048529FF9A5991F9135461F8D7" box="[151,302,1770,1795]" direction="south" orientation="latitude" pageId="27" pageNumber="478" precision="1" value="-12.373722">12°22'25.4&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048529FF9A583CF9125496F8D7" box="[314,473,1771,1795]" direction="west" orientation="longitude" pageId="27" pageNumber="478" precision="1" value="-70.37034">70°22'13.2&quot;W</geoCoordinate>
, T. Larsen,
<date id="FFC910038529FF9A5B58F91257E1F8D7" box="[606,686,1771,1795]" pageId="27" pageNumber="478" value="2000-05">V.2000</date>
(
<specimenCount id="9D71FD4A8529FF9A5BB9F91257ADF8D7" box="[703,738,1771,1795]" pageId="27" pageNumber="478" type="generic">1♂</specimenCount>
<collectionCode id="ED66AE068529FF9A5BEFF9125663F8D7" box="[745,812,1771,1795]" pageId="27" pageNumber="478">AFIC</collectionCode>
, 1♀
<collectionCode id="ED66AE068529FF9A5A65F91256F9F8D7" box="[867,950,1771,1795]" pageId="27" pageNumber="478">CEMT</collectionCode>
); same but Rio Palma Real Grande, Limon Camp,
<geoCoordinate id="EE4350048529FF9A59EEF8F65426F8F3" box="[232,361,1807,1831]" direction="south" orientation="latitude" pageId="27" pageNumber="478" precision="15" value="-12.538889">12°32'20&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048529FF9A5873F8F754B0F8F3" box="[373,511,1806,1831]" direction="west" orientation="longitude" pageId="27" pageNumber="478" precision="15" value="-68.86111">68°51'40&quot;W</geoCoordinate>
, Flight intercept trap,
<quantity id="4C8F9B268529FF9A5BF1F8F6560FF8F2" box="[759,832,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="478" unit="m" value="220.0">220 m</quantity>
, T. Larsen,
<date id="FFC910038529FF9A5AC5F8F6512CF8F3" box="[963,1123,1807,1831]" pageId="27" pageNumber="478" value="1999-10-11" valueMax="1999-10-12" valueMin="1999-10-11">1112.X.1999</date>
(1♀
<collectionCode id="ED66AE068529FF9A5D9BF8F651BFF8F3" box="[1181,1264,1807,1831]" pageId="27" pageNumber="478">CEMT</collectionCode>
); same but pitf. crished millipede,
<date id="FFC910038529FF9A587FF8CA5755F89F" box="[377,538,1843,1867]" pageId="27" pageNumber="478" value="1999-10-12" valueMax="1999-10-13" valueMin="1999-10-12">1213.X.1999</date>
(1♀
<collectionCode id="ED66AE068529FF9A5B61F8CA57E3F89F" box="[615,684,1843,1867]" pageId="27" pageNumber="478">AFIC</collectionCode>
); same but perching night,
<date id="FFC910038529FF9A5D0CF8CA51CFF89F" box="[1034,1152,1843,1867]" pageId="27" pageNumber="478" value="1999-10-11">11.X.1999</date>
(1♀
<collectionCode id="ED66AE068529FF9A5DCBF8CA505DF89F" box="[1229,1298,1843,1867]" pageId="27" pageNumber="478">AFIC</collectionCode>
); same but
<geoCoordinate id="EE4350048529FF9A5991F8AE5457F8BB" box="[151,280,1879,1903]" direction="south" orientation="latitude" pageId="27" pageNumber="478" precision="15" value="-12.538889">12°32'20&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048529FF9A5822F8AF54E1F8BB" box="[292,430,1878,1903]" direction="west" orientation="longitude" pageId="27" pageNumber="478" precision="15" value="-68.86139">68°51'41&quot;W</geoCoordinate>
, Flight interception trap,
<quantity id="4C8F9B268529FF9A5BCCF8AE565CF8BA" box="[714,787,1879,1903]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="478" unit="m" value="400.0">400 m</quantity>
,
<date id="FFC910038529FF9A5A19F8AE56E6F8BB" box="[799,937,1879,1903]" pageId="27" pageNumber="478" value="1999-04-02" valueMax="1999-04-03" valueMin="1999-04-02">23.IV.1999</date>
(1♀
<collectionCode id="ED66AE068529FF9A5AE5F8AE5167F8BB" box="[995,1064,1879,1903]" pageId="27" pageNumber="478">AFIC</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38529FF9A59C1F8835041F847" blockId="27.[151,1437,582,2011]" box="[199,1294,1914,1939]" pageId="27" pageNumber="478">
<emphasis id="B903EAD18529FF9A59C1F8835465F847" bold="true" box="[199,298,1914,1939]" pageId="27" pageNumber="478">Habitat.</emphasis>
<collectingCountry id="F36076538529FF9A5837F88354C1F847" box="[305,398,1914,1939]" name="Ecuador" pageId="27" pageNumber="478">Ecuador</collectingCountry>
and
<collectingCountry id="F36076538529FF9A58C2F88254B7F846" box="[452,504,1915,1938]" name="Peru" pageId="27" pageNumber="478">Peru</collectingCountry>
(
<figureCitation id="134C2A468529FF9A5B01F8835715F847" box="[519,602,1914,1939]" captionStart="FIGURES 89 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="27" pageNumber="478">Fig. 91</figureCitation>
), from available data this species inhabits Amazon rainforest.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488529FF9D59C1F8675601FE5C" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478" type="diagnosis">
<paragraph id="8BC836C38529FF9D59C1F8675601FE5C" blockId="27.[151,1437,582,2011]" lastBlockId="28.[151,1436,151,392]" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478">
<emphasis id="B903EAD18529FF9A59C1F8675495F863" bold="true" box="[199,474,1950,1975]" pageId="27" pageNumber="478">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408529FF9A5B67F8665787F862" box="[609,712,1951,1974]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD18529FF9A5B67F8665787F862" box="[609,712,1951,1974]" italics="true" pageId="27" pageNumber="478">acropyge</emphasis>
</taxonomicName>
complex): length &lt;
<quantity id="4C8F9B268529FF9A5AADF86656B3F862" box="[939,1020,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="478" unit="mm" value="13.0">13 mm</quantity>
; punctures of pronotal disc separated by approximately one diameter; posteromedial punctures of pronotum usually separated by at least one diameter; pronotal margin, between lateral and posterior angles, with length somewhat shorter than margin between anterior and lateral angles; elytral striae with carinate margins clearly separated; strial punctures separated by regular spacing; most of strial punctures round intermingled with some slightly elliptical; apical carina or tubercle of second interstria absent; pseudepipleural carina incomplete, effaced medially (as in Fig. 80). The following aedeagus characters are also diagnostic: parameres symmetric and short, length shorter than phallobase; in lateral view forming an angle of approximately 45° with phallobase (
<figureCitation id="134C2A46852EFF9D5A62FEB556F6FEB0" box="[868,953,332,357]" captionStart="FIGURES 12 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="28" pageNumber="479">Fig. 14</figureCitation>
). Lamella copulatrix (LC): subrectangular sclerites. Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.
</paragraph>
</subSubSection>
</treatment>
</document>