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<document ID-DOI="10.5733/afin.053.0115" ID-ISSN="2305-2562" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1683581940806" checkinUser="felipe" docAuthor="Mathis, Wayne N. &amp; Freidberg, Amnon" docDate="2012" docId="03DE879FDF4BCF6E448211EFFCFE4111" docLanguage="en" docName="AfrInvertebr.53.1.231-238.pdf" docOrigin="African Invertebrates 53 (1)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.053.0115" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Periscelididae Oldenberg 1914" docType="treatment" docVersion="2" lastPageNumber="232" masterDocId="FFE7FFE7DF4ACF6F4538136BFF86471F" masterDocTitle="Periscelis stuckenbergi sp. n., the first record of the genus from the Afrotropical Region (Diptera: Periscelididae: Periscelidinae)" masterLastPageNumber="231" masterPageNumber="231" pageNumber="232" updateTime="1683638392222" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Periscelis stuckenbergi sp. n., the first record of the genus from the Afrotropical Region (Diptera: Periscelididae: Periscelidinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mathis, Wayne N.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Freidberg, Amnon</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>231</mods:start>
<mods:end>231</mods:end>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.053.0115</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.053.0115</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DE879FDF4BCF6E448211EFFCFE4111" httpUri="http://treatment.plazi.org/id/03DE879FDF4BCF6E448211EFFCFE4111" lastPageNumber="232" pageId="1" pageNumber="232">
<subSubSection box="[442,880,644,670]" pageId="1" pageNumber="232" type="nomenclature">
<paragraph blockId="1.[442,880,644,670]" box="[442,880,644,670]" pageId="1" pageNumber="232">
<heading box="[442,880,644,670]" centered="true" fontSize="36" level="2" pageId="1" pageNumber="232" reason="7">
Family
<taxonomicName ID-CoL="625V3" ID-ENA="286481" authority="Oldenberg, 1914" authorityName="Oldenberg" authorityYear="1914" box="[529,880,644,670]" class="Insecta" family="Periscelididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">
Periscelididae
<bibRefCitation author="OLDENBERG, L." box="[693,880,644,670]" pageId="1" pageNumber="232" pagination="1 - 42" refId="ref4365" refString="OLDENBERG, L. 1914. Beitrag zur Kenntnis der europaischen Drosophiliden (Dipt.). Archiv fur Naturgeschichte 80 A (2): 1 - 42." type="journal article" year="1914">Oldenberg, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="232" type="diagnosis">
<paragraph blockId="1.[181,1142,683,1550]" pageId="1" pageNumber="232">
Diagnosis: Adult. Small flies, body length 2.55.0 mm, moderately broad to slender.
<emphasis box="[181,242,716,742]" italics="true" pageId="1" pageNumber="232">Head</emphasis>
: Width equal to that of thorax, distinctly wider than high; frons wider than long, becoming narrower toward anterior margin, bearing 1 or 2 fronto­orbital setae; postocellar setae present and divergent or absent. Pedicel cap­like and with a dorsal cleft, bearing 1 or more dorsoapical setae; flagellomere 1 frequently sharply deflexed, extended from ventral surface of pedicel; arista bipectinate. Face uniformly sclerotised and arched, setose laterally.
<emphasis box="[361,442,876,902]" italics="true" pageId="1" pageNumber="232">Thorax</emphasis>
: Dorsocentral setae usually 2 (0+2), sometimes 1 (0+1), none presutural; posterior intra­alar seta reduced; scutellum with 1 or 2 pairs of marginal setae; scutellar disc bare; anepisternal seta usually lacking (present in
<taxonomicName authority="Cresson, 1914" authorityName="Cresson" authorityYear="1914" class="Insecta" family="Periscelididae" genus="Planinasus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[911,1035,940,966]" italics="true" pageId="1" pageNumber="232">Planinasus</emphasis>
Cresson, 1914
</taxonomicName>
). Wing: Hyaline or with infuscate markings. Subcosta rudimentary, not reaching costal margin, and not fused apically with
<emphasis box="[651,676,1004,1036]" italics="true" pageId="1" pageNumber="232">
R
<subScript attach="left" box="[668,676,1022,1036]" fontSize="6" pageId="1" pageNumber="232">1</subScript>
</emphasis>
; no costal breaks (a weakness in the costa just apicad of the humeral crossvein in
<taxonomicName authorityName="Cresson" authorityYear="1914" box="[622,746,1036,1062]" class="Insecta" family="Periscelididae" genus="Planinasus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[622,746,1036,1062]" italics="true" pageId="1" pageNumber="232">Planinasus</emphasis>
</taxonomicName>
); costa extended to vein
<emphasis box="[1028,1073,1036,1068]" italics="true" pageId="1" pageNumber="232">
R
<subScript attach="left" box="[1045,1073,1054,1068]" fontSize="6" pageId="1" pageNumber="232">4+5</subScript>
</emphasis>
or
<emphasis box="[1110,1133,1036,1061]" italics="true" pageId="1" pageNumber="232">M</emphasis>
; cell
<emphasis box="[229,263,1068,1094]" italics="true" pageId="1" pageNumber="232">dm</emphasis>
with shallow fold running entire length; cell
<emphasis box="[767,807,1068,1093]" italics="true" pageId="1" pageNumber="232">cup</emphasis>
present, although
<emphasis box="[1013,1071,1068,1100]" italics="true" pageId="1" pageNumber="232">
CuA
<subScript attach="left" box="[1063,1071,1086,1100]" fontSize="6" pageId="1" pageNumber="232">2</subScript>
</emphasis>
either well developed or extremely reduced. Mid tibia bearing prominent, apicoventral seta. Classification: The concept of
<taxonomicName authorityName="Oldenberg" authorityYear="1914" box="[514,673,1140,1166]" class="Insecta" family="Periscelididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Periscelididae</taxonomicName>
, as adopted here, follows
<bibRefCitation author="MCALPINE, D. K." box="[952,1137,1140,1166]" pageId="1" pageNumber="232" pagination="273 - 295" refId="ref4204" refString="MCALPINE, D. K. 1978. Description and biology of a new genus of flies related to Anthoclusia and representing a new family (Diptera, Schizophora, Neurochaetidae). Annals of the Natal Museum 23: 273 - 295." type="journal article" year="1978">McAlpine (1978</bibRefCitation>
, 1983) and includes a few genera previously assigned to
<taxonomicName authority="Duda, 1924" authorityName="Duda" authorityYear="1924" box="[824,1141,1172,1198]" class="Insecta" family="Aulacigastridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Aulacigastridae Duda, 1924</taxonomicName>
(
<taxonomicName authorityName="Melander" authorityYear="1913" box="[191,295,1204,1230]" class="Insecta" family="Periscelididae" genus="Cyamops" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[191,295,1204,1230]" italics="true" pageId="1" pageNumber="232">Cyamops</emphasis>
</taxonomicName>
,
<taxonomicName authority=", Stenocyamops Papp, 2006" authorityName=", Stenocyamops Papp" authorityYear="2006" box="[308,737,1204,1230]" class="Insecta" family="Periscelididae" genus="Planinasus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[308,432,1204,1230]" italics="true" pageId="1" pageNumber="232">Planinasus</emphasis>
,
<emphasis box="[446,606,1204,1230]" italics="true" pageId="1" pageNumber="232">Stenocyamops</emphasis>
Papp, 2006
</taxonomicName>
and
<taxonomicName authorityName="Coquillett" authorityYear="1900" box="[790,918,1204,1230]" class="Insecta" family="Periscelididae" genus="Stenomicra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[790,918,1204,1230]" italics="true" pageId="1" pageNumber="232">Stenomicra</emphasis>
</taxonomicName>
). McAlpine characterised
<taxonomicName authorityName="Oldenberg" authorityYear="1914" box="[264,421,1236,1262]" class="Insecta" family="Periscelididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Periscelididae</taxonomicName>
primarily by the cap­like pedicel, which has a dorsal cleft, and its relationship to the first flagellomere. Although these characters are common to all
<taxonomicName authorityName="Oldenberg" authorityYear="1914" box="[181,341,1300,1326]" class="Insecta" family="Periscelididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Periscelididae</taxonomicName>
, they also occur in
<taxonomicName authority="McAlpine, 1978" authorityName="McAlpine" authorityYear="1978" box="[548,910,1300,1326]" class="Insecta" family="Neurochaetidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">
Neurochaetidae
<bibRefCitation author="MCALPINE, D. K." box="[729,910,1300,1326]" pageId="1" pageNumber="232" pagination="273 - 295" refId="ref4204" refString="MCALPINE, D. K. 1978. Description and biology of a new genus of flies related to Anthoclusia and representing a new family (Diptera, Schizophora, Neurochaetidae). Annals of the Natal Museum 23: 273 - 295." type="journal article" year="1978">McAlpine, 1978</bibRefCitation>
</taxonomicName>
(
<bibRefCitation author="WOODLEY, N. E." box="[923,1087,1300,1326]" pageId="1" pageNumber="232" pagination="211 - 218" refId="ref4688" refString="WOODLEY, N. E. 1982. Two new species of Neurochaeta McAlpine (Diptera: Neurochaetidae), with notes on cladistic relationships within the genus. Memoirs of the Entomological Society of Washington 10: 211 - 218." type="journal article" year="1982">Woodley 1982</bibRefCitation>
) and some other acalyptrate genera. In a recent phylogenetic study of the Opomyzoidea Fallén, 1820, using 28S ribosomal DNA and
<collectingCountry box="[601,660,1364,1390]" name="Chad" pageId="1" pageNumber="232">CAD</collectingCountry>
(rudimentary) genes (
<bibRefCitation author="WINKLER, I. S. &amp; RUNG, R. &amp; SCHEFFER, S. J." box="[907,1129,1364,1390]" pageId="1" pageNumber="232" pagination="746 - 762" refId="ref4645" refString="WINKLER, I. S., RUNG, R. &amp; SCHEFFER, S. J. 2010. Hennig's orphans revisited: testing morphological hypotheses in the &quot; Opomyzoidea &quot;. Molecular Phylogenetics and Evolution 54: 746 - 762." type="journal article" year="2010">
Winkler
<emphasis box="[1007,1063,1364,1390]" italics="true" pageId="1" pageNumber="232">et al.</emphasis>
2010
</bibRefCitation>
),
<taxonomicName authority=", Cyamops" authorityName="Cyamops" box="[181,425,1396,1422]" class="Insecta" family="Periscelididae" genus="Stenomicra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[181,308,1396,1422]" italics="true" pageId="1" pageNumber="232">Stenomicra</emphasis>
,
<emphasis box="[321,425,1396,1422]" italics="true" pageId="1" pageNumber="232">Cyamops</emphasis>
</taxonomicName>
and
<taxonomicName authorityName=", Stenocyamops Papp" authorityYear="2006" box="[475,599,1396,1422]" class="Insecta" family="Periscelididae" genus="Planinasus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[475,599,1396,1422]" italics="true" pageId="1" pageNumber="232">Planinasus</emphasis>
</taxonomicName>
grouped consistently with moderate support with the genus
<taxonomicName authority="Macquart, 1835" authorityName="Macquart" authorityYear="1835" box="[290,612,1427,1454]" class="Insecta" family="Aulacigastridae" genus="Aulacigaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis box="[290,432,1428,1454]" italics="true" pageId="1" pageNumber="232">Aulacigaster</emphasis>
Macquart, 1835
</taxonomicName>
and not with
<taxonomicName authorityName=": Oldenberg" authorityYear="1914" box="[757,916,1428,1454]" class="Globothalamea" family="Spiroplectamminidae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Lituolida" pageId="1" pageNumber="233" phylum="Foraminifera" rank="subFamily" subFamily="Periscelidinae">Periscelidinae</taxonomicName>
. Moreover, the same analysis failed to find any support for a sister­group relationship between
<taxonomicName authorityName="Oldenberg" authorityYear="1914" box="[985,1142,1460,1486]" class="Insecta" family="Periscelididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Periscelididae</taxonomicName>
and
<taxonomicName authorityName="McAlpine" authorityYear="1978" box="[230,410,1492,1518]" class="Insecta" family="Neurochaetidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Neurochaetidae</taxonomicName>
. Their results highlight the need to study the phylogeny of these groups in greater detail and using different suites of characters.
</paragraph>
</subSubSection>
</treatment>
</document>