453 lines
79 KiB
XML
453 lines
79 KiB
XML
<document id="658F38A43216331554FFDC60FA7F7FC4" ID-CLB-Dataset="7413" ID-DOI="10.11646/zootaxa.4951.1.2" ID-GBIF-Dataset="d7a40f10-4145-499c-97be-3f9bc5088d2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655625" ID-ZooBank="557825A0-714A-426A-917F-1C9AB7372C30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617265349256" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Makarkin, Vladimir N." docDate="2021" docId="03DE878EFFE7C459FF67F964C8268436" docLanguage="en" docName="zootaxa.4951.1.2.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archiinocellia protomaculata Archibald & Makarkin 2021, comb. nov." docType="treatment" docVersion="6" lastPageNumber="64" masterDocId="FFE7FFF6FFF6C44EFFF0FFA7CC6F8000" masterDocTitle="Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation" masterLastPageNumber="79" masterPageNumber="41" pageNumber="58" updateTime="1698935824968" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="EA75CB6C953A9B43B0D9E0699F89C3B0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="73489CA555B5F57226ED6146EEE2B94A">
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<mods:title id="6FD44C3947CDCB1CB329F0E1783B802B">Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation</mods:title>
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<mods:name id="E3DFC8232CBE241E1DD3E104853C657F" type="personal">
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<mods:roleTerm id="D38621A9410D4C0C2A3E967536F5692F">Author</mods:roleTerm>
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<mods:namePart id="12B55231CF5521A4AA020334B000F333">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="773E6B50EF087E3861D16190C5A717D1" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation id="1BE3F571D50EC7564043CE5323AA422F">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:roleTerm id="A528F8C157DCEA7E5C7C61656BF2C788">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="89EABF834ED0FF4F4AF0D0C199DEE2A1">Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation id="938DAD530F39CB04B7808FF6349CB406">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivo-</mods:affiliation>
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<mods:typeOfResource id="AE8678841740C95829559692B4F53368">text</mods:typeOfResource>
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<mods:titleInfo id="63655C4721F237E521BFC1DFBBD3EDF2">
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<mods:title id="B94AD731289C4048EFAEDEB42A025D6F">Zootaxa</mods:title>
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<mods:part id="8A96A5A788B7943484B2B349F8CB59EC">
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<mods:date id="5B69A9DF5E088B3729D1BA63A5C636B5">2021</mods:date>
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<mods:number id="5870361EEC207ACABAFEBB68B9CD4F05">2021-04-01</mods:number>
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<mods:number id="73ACC89946D05C4C6D6C3F62D3AEE6B3">4951</mods:number>
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<mods:number id="EEBBEFAAB1CAAA0A73B1669EFAF4D05A">1</mods:number>
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<mods:classification id="D0C2E58E289823FA6566DF050874E59A">journal article</mods:classification>
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<mods:identifier id="544E0CB28452772759B6315C6C409A30" type="DOI">10.11646/zootaxa.4951.1.2</mods:identifier>
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<mods:identifier id="AE91FA46CC20861567D3F2BBDFB0395F" type="GBIF-Dataset">d7a40f10-4145-499c-97be-3f9bc5088d2f</mods:identifier>
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<mods:identifier id="E88AD26AE11DC491AA3EC84D5B8BD636" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F994EEC178182B00DB3363B0AF358C81" type="ZooBank">557825A0-714A-426A-917F-1C9AB7372C30</mods:identifier>
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</mods:mods>
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<treatment id="03DE878EFFE7C459FF67F964C8268436" ID-DOI="http://doi.org/10.5281/zenodo.4655649" ID-GBIF-Taxon="180999352" ID-Zenodo-Dep="4655649" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436" httpUri="http://treatment.plazi.org/id/03DE878EFFE7C459FF67F964C8268436" lastPageId="23" lastPageNumber="64" pageId="17" pageNumber="58">
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<subSubSection id="C36D6513FFE7C45FFF67F964CF5486DD" box="[151,827,1731,1758]" pageId="17" pageNumber="58" type="nomenclature">
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<paragraph id="8BC83698FFE7C45FFF67F964CF5486DD" blockId="17.[151,827,1731,1793]" box="[151,827,1731,1758]" pageId="17" pageNumber="58">
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<heading id="D08081F4FFE7C45FFF67F964CF5486DD" bold="true" box="[151,827,1731,1758]" fontSize="11" level="1" pageId="17" pageNumber="58" reason="1">
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<emphasis id="B903EA8AFFE7C45FFF67F964CF5486DD" bold="true" box="[151,827,1731,1758]" pageId="17" pageNumber="58">
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<taxonomicName id="4C774D1BFFE7C45FFF67F964CEC886DD" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[151,679,1731,1758]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="protomaculata" status="comb. nov.">
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<emphasis id="B903EA8AFFE7C45FFF67F964CD9486DE" bold="true" box="[151,507,1731,1758]" italics="true" pageId="17" pageNumber="58">Archiinocellia protomaculata</emphasis>
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(
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<bibRefCitation id="EFE64B69FFE7C45FFDFBF964CEF086DD" author="Engel, M. S." box="[523,671,1731,1757]" pageId="17" pageNumber="58" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23057F1FFE7C45FFD44F964CF5486DD" box="[692,827,1731,1757]" pageId="17" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36D6513FFE7C45FFF67F940CD7A8701" box="[151,277,1767,1793]" pageId="17" pageNumber="58" type="description">
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<paragraph id="8BC83698FFE7C45FFF67F940CD7A8701" blockId="17.[151,827,1731,1793]" box="[151,277,1767,1793]" pageId="17" pageNumber="58">
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<figureCitation id="134C2A1DFFE7C45FFF67F940CD7A8701" box="[151,277,1767,1793]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStart-4="FIGURE 16" captionStart-5="FIGURE 17" captionStartId-0="18.[151,250,1876,1900]" captionStartId-1="19.[151,250,1577,1601]" captionStartId-2="20.[151,250,1455,1479]" captionStartId-3="21.[151,250,1457,1481]" captionStartId-4="22.[151,250,1687,1711]" captionStartId-5="24.[151,250,1943,1967]" captionTargetBox-0="[151,1436,705,1840]" captionTargetBox-1="[169,1417,181,1553]" captionTargetBox-2="[151,1436,181,1431]" captionTargetBox-3="[151,1436,525,1433]" captionTargetBox-4="[160,1427,326,1646]" captionTargetBox-5="[167,1424,191,1902]" captionTargetId-0="figure-254@18.[151,1436,705,1853]" captionTargetId-1="figure-19@19.[169,1417,181,1553]" captionTargetId-2="figure-26@20.[151,1436,181,1431]" captionTargetId-3="figure-191@21.[151,1436,525,1433]" captionTargetId-4="figure-89@22.[151,1436,309,1664]" captionTargetId-5="figure-26@24.[151,1436,181,1919]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionTargetPageId-3="21" captionTargetPageId-4="22" captionTargetPageId-5="24" captionText-0="FIGURE 12. Archiinocellia protomaculata (Engel, 2011), females. A, holotype USNM 31487 (counterpart); B, UCM 42107. Scale bars = 5 mm." captionText-1="FIGURE 13. Archiinocellia protomaculata (Engel, 2011), abdomen. A, UCM 61231b, male (lateral/slightly dorsal view); B, UCM 42684, male (lateral view); C, UCM 45968, female. e, ectoproct; gx9?, possible gonocoxites 9; ov, ovipositor; S2 to S7, second to seventh sternites; T3 to T9, third to ninth tergites. Scale bars = 1 mm." captionText-2="FIGURE 14. Archiinocellia protomaculata (Engel, 2011).A, UCM 61231, rock with four specimens of A. protomaculata (a–c) and one undeterminable Raphidioptera (d); B, UCM 62523. Scale bars = 1 mm." captionText-3="FIGURE 15. Archiinocellia protomaculata (Engel, 2011), head and prothorax (dorsal view). A, UCM 64727; B, UCM 62525; C, UCM 35717; D, UCM 45969; E, UCM 42122; F, UCM 62687. Scale bars = 1 mm." captionText-4="FIGURE 16. Venation of Archiinocellia protomaculata (Engel, 2011), holotype USNM 31487. A, right forewing; B, left forewing; C, right hind wing. Scale bar = 3 mm (all to scale)." captionText-5="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.4655671" figureDoi-1="http://doi.org/10.5281/zenodo.4655677" figureDoi-2="http://doi.org/10.5281/zenodo.4655680" figureDoi-3="http://doi.org/10.5281/zenodo.4655684" figureDoi-4="http://doi.org/10.5281/zenodo.4655686" figureDoi-5="http://doi.org/10.5281/zenodo.4655688" httpUri-0="https://zenodo.org/record/4655671/files/figure.png" httpUri-1="https://zenodo.org/record/4655677/files/figure.png" httpUri-2="https://zenodo.org/record/4655680/files/figure.png" httpUri-3="https://zenodo.org/record/4655684/files/figure.png" httpUri-4="https://zenodo.org/record/4655686/files/figure.png" httpUri-5="https://zenodo.org/record/4655688/files/figure.png" pageId="17" pageNumber="58">Figs 12–17</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36D6513FFE7C45CFF67F888CFF281AE" lastPageId="18" lastPageNumber="59" pageId="17" pageNumber="58" type="materials_examined">
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<paragraph id="8BC83698FFE7C45CFF67F888C8D48165" blockId="17.[151,1436,1839,2010]" lastBlockId="18.[151,1437,150,681]" lastPageId="18" lastPageNumber="59" pageId="17" pageNumber="58">
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||
<emphasis id="B903EA8AFFE7C45FFF67F888CDEE8749" bold="true" box="[151,385,1839,1865]" pageId="17" pageNumber="58">Material examined.</emphasis>
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Thirty-six specimens collected by David D. Kohls in 2004 to 2009 at three localities in Garfield County (
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<collectingRegion id="49B3F87AFFE7C45FFEC5F8F4CDCD876D" box="[309,418,1875,1901]" country="United States of America" name="Colorado" pageId="17" pageNumber="58">Colorado</collectingRegion>
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,
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<collectingCountry id="F3607608FFE7C45FFE5FF8F4CD90876D" box="[431,511,1875,1901]" name="United States of America" pageId="17" pageNumber="58">U.S.A.</collectingCountry>
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): (1)
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<collectionCode id="ED66AE5DFFE7C45FFDCEF8F4CE11876D" box="[574,638,1875,1901]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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locality 2005026 (Labandeira Site): specimens
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<collectionCode id="ED66AE5DFFE7C45FFB6FF8F4C8B0876D" box="[1183,1247,1875,1901]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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35717 (female);
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<collectionCode id="ED66AE5DFFE7C45FFF67F8D0CCB88791" box="[151,215,1911,1937]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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40659 (male);
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<collectionCode id="ED66AE5DFFE7C45FFE8EF8D0CDD18791" box="[382,446,1911,1937]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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58068 (sex unknown);
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<collectionCode id="ED66AE5DFFE7C45FFD31F8D0CF6E8791" box="[705,769,1911,1937]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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59090 (female);
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<collectionCode id="ED66AE5DFFE7C45FFC4EF8D0CF918791" box="[958,1022,1911,1937]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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59509 (male);
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<collectionCode id="ED66AE5DFFE7C45FFB55F8D0C88A8791" box="[1189,1253,1911,1937]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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60297 (female);
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<collectionCode id="ED66AE5DFFE7C45FFF67F83CCCB887B5" box="[151,215,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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61231a (female);
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<collectionCode id="ED66AE5DFFE7C45FFE54F83CCD8B87B5" box="[420,484,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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61231b (male);
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<collectionCode id="ED66AE5DFFE7C45FFD6CF83CCEB387B5" box="[668,732,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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61231с (sex unknown);
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<collectionCode id="ED66AE5DFFE7C45FFC00F83CC85F87B5" box="[1008,1072,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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62026 (male);
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<collectionCode id="ED66AE5DFFE7C45FFB2BF83CC97487B5" box="[1243,1307,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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62523 (sex unknown);
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<collectionCode id="ED66AE5DFFE7C45FFEEAF818CD3587D9" box="[282,346,1983,2009]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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62525 (three specimens: male, female, and sex unknown);
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<collectionCode id="ED66AE5DFFE7C45FFBF6F818C82987D9" box="[1030,1094,1983,2009]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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62687 (female);
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<collectionCode id="ED66AE5DFFE7C45FFAFDF818C92287D9" box="[1293,1357,1983,2009]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="17" pageNumber="58" type="University or college">UCM</collectionCode>
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||
64319 (sex unknown);
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<collectionCode id="ED66AE5DFFE4C45CFEBDFF30CDE280B1" box="[333,397,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
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||
64727 (female);
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<collectionCode id="ED66AE5DFFE4C45CFDBEFF30CEE180B1" box="[590,654,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
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65146 (sex unknown);
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<collectionCode id="ED66AE5DFFE4C45CFC67FF30CFB880B1" box="[919,983,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
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66134 (sex unknown);
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<collectionCode id="ED66AE5DFFE4C45CFB10FF30C94F80B1" box="[1248,1312,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
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67986 (female);
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<collectionCode id="ED66AE5DFFE4C45CFF15FF1CCD4A80D5" box="[229,293,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
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||
72989 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFE12FF1CCE4D80D5" box="[482,546,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
73247 (female); (2)
|
||
<collectionCode id="ED66AE5DFFE4C45CFCF7FF1CCF2880D5" box="[775,839,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
locality 2006032 (Donnell Site):
|
||
<collectionCode id="ED66AE5DFFE4C45CFB4AFF1CC89580D5" box="[1210,1274,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
40057 (male);
|
||
<collectionCode id="ED66AE5DFFE4C45CFF67FF78CCB880F9" box="[151,215,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
42684 (male);
|
||
<collectionCode id="ED66AE5DFFE4C45CFE79FF78CDA680F9" box="[393,457,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
42744 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFD60FF78CEBF80F9" box="[656,720,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
45968 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFC67FF78CFB880F9" box="[919,983,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
45969 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFB6EFF78C8B180F9" box="[1182,1246,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
45970 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFF67FEA4CCB8811D" box="[151,215,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
53268 (female); (3)
|
||
<collectionCode id="ED66AE5DFFE4C45CFE4FFEA4CD90811D" box="[447,511,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
locality 2007238 (Denson Site):
|
||
<collectionCode id="ED66AE5DFFE4C45CFC81FEA4CFDE811D" box="[881,945,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
41697 (male);
|
||
<collectionCode id="ED66AE5DFFE4C45CFBACFEA4C8F3811D" box="[1116,1180,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
42105 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFAACFEA4C9F3811D" box="[1372,1436,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
42107 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFEA0FE80CDFF8141" box="[336,400,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
42122 (sex unknown);
|
||
<collectionCode id="ED66AE5DFFE4C45CFD65FE80CEBA8141" box="[661,725,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
54806 (male);
|
||
<collectionCode id="ED66AE5DFFE4C45CFC8EFE80CFD18141" box="[894,958,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
55329 (female);
|
||
<collectionCode id="ED66AE5DFFE4C45CFB8CFE80C8D38141" box="[1148,1212,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
56552 (female); the Parachute Creek Member of the Green River Formation; all are deposited in
|
||
<collectionCode id="ED66AE5DFFE4C45CFC03FEECC85C8165" box="[1011,1075,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="18" pageNumber="59" type="University or college">UCM</collectionCode>
|
||
collections.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE4C45CFF37FEC8CFF281AE" blockId="18.[151,1437,150,681]" pageId="18" pageNumber="59">
|
||
<materialsCitation id="3B1F3CC5FFE4C45CFF37FEC8CFF681AE" ID-GBIF-Occurrence="3071122307" country="United States of America" county="Garfield County" location="Probably Labandeira Site" municipality="Eocene" pageId="18" pageNumber="59" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
|
||
<emphasis id="B903EA8AFFE4C45CFF37FEC8CD918188" bold="true" box="[199,510,366,393]" pageId="18" pageNumber="59">
|
||
<typeStatus id="54CC883AFFE4C45CFF37FEC8CD6D8189" box="[199,258,367,393]" pageId="18" pageNumber="59">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
<location id="8EA86043FFE4C45CFDF5FEC8CF4D8189" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFDF5FEC8CF4D8189" box="[517,802,367,393]" country="United States of America" county="Garfield County" municipality="Eocene" name="Probably Labandeira Site" pageId="18" pageNumber="59" stateProvince="Colorado">Probably Labandeira Site</location>
|
||
in the
|
||
<location id="8EA86043FFE4C45CFC9FFEC8C8E28189" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFC9FFEC8C8E28189" box="[879,1165,367,393]" country="United States of America" county="Garfield County" municipality="Eocene" name="Parachute Creek Member" pageId="18" pageNumber="59" stateProvince="Colorado">
|
||
<location id="8EA86043FFE4C45CFC9FFEC8C8488189" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFC9FFEC8C8488189" box="[879,1063,367,393]" country="United States of America" county="Garfield County" municipality="Eocene" name="Parachute Creek" pageId="18" pageNumber="59" stateProvince="Colorado">Parachute Creek</location>
|
||
Member
|
||
</location>
|
||
of the
|
||
<location id="8EA86043FFE4C45CFB2CFEC8CC8781AD" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFB2CFEC8CC8781AD" country="United States of America" county="Garfield County" municipality="Eocene" name="Green River Formation" pageId="18" pageNumber="59" stateProvince="Colorado">
|
||
<location id="8EA86043FFE4C45CFB2CFEC8C90A8189" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFB2CFEC8C90A8189" box="[1244,1381,367,393]" country="United States of America" county="Garfield County" municipality="Eocene" name="Green River" pageId="18" pageNumber="59" stateProvince="Colorado">
|
||
Green
|
||
<location id="8EA86043FFE4C45CFAD7FEC8C90A8189" LSID="urn:lsid:plazi:treatment:03DE878EFFE7C459FF67F964C8268436:8EA86043FFE4C45CFAD7FEC8C90A8189" box="[1319,1381,367,393]" country="United States of America" county="Garfield County" municipality="Eocene" name="River Formation" pageId="18" pageNumber="59" stateProvince="Colorado">River</location>
|
||
</location>
|
||
Formation
|
||
</location>
|
||
,
|
||
<collectingCounty id="62A94E14FFE4C45CFF03FE34CDC481AD" box="[243,427,403,429]" pageId="18" pageNumber="59">Garfield County</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B3F87AFFE4C45CFE46FE34CE4E81AD" box="[438,545,403,429]" country="United States of America" name="Colorado" pageId="18" pageNumber="59">Colorado</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3607608FFE4C45CFDDCFE34CE1381AD" box="[556,636,403,429]" name="United States of America" pageId="18" pageNumber="59">U.S.A.</collectingCountry>
|
||
; early
|
||
<collectingMunicipality id="6BACACE2FFE4C45CFD37FE34CF7681AD" box="[711,793,403,429]" pageId="18" pageNumber="59">Eocene</collectingMunicipality>
|
||
(Ypresian)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFE4C45CFF37FE11C9FA81D0" box="[199,1429,438,466]" pageId="18" pageNumber="59" type="diagnosis">
|
||
<paragraph id="8BC83698FFE4C45CFF37FE11C9FA81D0" blockId="18.[151,1437,150,681]" box="[199,1429,438,466]" pageId="18" pageNumber="59">
|
||
<emphasis id="B903EA8AFFE4C45CFF37FE11CD2D81D0" bold="true" box="[199,322,438,464]" pageId="18" pageNumber="59">Diagnosis.</emphasis>
|
||
May be distinguished from
|
||
<taxonomicName id="4C774D1BFFE4C45CFD8EFE10CF7781D1" authorityName="Handlirsch" authorityYear="1910" box="[638,792,439,465]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFE4C45CFD8EFE10CF7781D1" box="[638,792,439,465]" italics="true" pageId="18" pageNumber="59">A. oligoneura</emphasis>
|
||
</taxonomicName>
|
||
by markedly narrower fore- and hind wings (see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFE4C458FF37FE7DC96E811E" lastPageId="22" lastPageNumber="63" pageId="18" pageNumber="59" type="description">
|
||
<paragraph id="8BC83698FFE4C45CFF37FE7DCEB182A9" blockId="18.[151,1437,150,681]" pageId="18" pageNumber="59">
|
||
<emphasis id="B903EA8AFFE4C45CFF37FE7DCD0281F4" bold="true" box="[199,365,474,500]" pageId="18" pageNumber="59">Redescription</emphasis>
|
||
(based on the
|
||
<typeStatus id="54CC883AFFE4C45CFDE1FE7CCE1C81F5" box="[529,627,475,501]" pageId="18" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
and material from UCM listed above): body length of male
|
||
<quantity id="4C8F9B7DFFE4C45CFAE6FE7CC9F281F5" box="[1302,1437,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="8.649999999999999" metricValueMax="9.2" metricValueMin="8.1" pageId="18" pageNumber="59" unit="mm" value="8.649999999999999" valueMax="9.2" valueMin="8.1">8.1–9.2 mm</quantity>
|
||
(five specimens), female (without ovipositor)
|
||
<quantity id="4C8F9B7DFFE4C45CFD53FE58CF448219" box="[675,811,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.1" metricValueMin="8.7" pageId="18" pageNumber="59" unit="mm" value="8.899999999999999" valueMax="9.1" valueMin="8.7">8.7–9.1 mm</quantity>
|
||
(seven specimens). Head broadly oval, somewhat narrowed posteriorly (in dorsal view); dorsal maculation very distinct with extensive pale areas (
|
||
<figureCitation id="134C2A1DFFE4C45CFB32FD84C97A823D" box="[1218,1301,546,573]" captionStart="FIGURE 15" captionStartId="21.[151,250,1457,1481]" captionTargetBox="[151,1436,525,1433]" captionTargetId="figure-191@21.[151,1436,525,1433]" captionTargetPageId="21" captionText="FIGURE 15. Archiinocellia protomaculata (Engel, 2011), head and prothorax (dorsal view). A, UCM 64727; B, UCM 62525; C, UCM 35717; D, UCM 45969; E, UCM 42122; F, UCM 62687. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655684" httpUri="https://zenodo.org/record/4655684/files/figure.png" pageId="18" pageNumber="59">Fig. 15</figureCitation>
|
||
), especially beyond and around eyes, broad medial stripe (sometimes divided by a thin brown medial line:
|
||
<figureCitation id="134C2A1DFFE4C45CFB4FFDE0C94E8261" box="[1215,1313,582,609]" captionStart="FIGURE 15" captionStartId="21.[151,250,1457,1481]" captionTargetBox="[151,1436,525,1433]" captionTargetId="figure-191@21.[151,1436,525,1433]" captionTargetPageId="21" captionText="FIGURE 15. Archiinocellia protomaculata (Engel, 2011), head and prothorax (dorsal view). A, UCM 64727; B, UCM 62525; C, UCM 35717; D, UCM 45969; E, UCM 42122; F, UCM 62687. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655684" httpUri="https://zenodo.org/record/4655684/files/figure.png" pageId="18" pageNumber="59">Fig. 15F</figureCitation>
|
||
), and most areas before the eyes. Palpi brownish. Antennae: scapus brown to dark brown; flagellum distinctly paler, with slightly more than 40 segments. Ocelli not detected.
|
||
</paragraph>
|
||
<caption id="DF086610FFE4C45CFF67F8F3CD34878F" ID-DOI="http://doi.org/10.5281/zenodo.4655671" ID-Zenodo-Dep="4655671" httpUri="https://zenodo.org/record/4655671/files/figure.png" pageId="18" pageNumber="59" startId="18.[151,250,1876,1900]" targetBox="[151,1436,705,1840]" targetPageId="18">
|
||
<paragraph id="8BC83698FFE4C45CFF67F8F3CD34878F" blockId="18.[151,1437,1876,1936]" pageId="18" pageNumber="59">
|
||
<emphasis id="B903EA8AFFE4C45CFF67F8F3CD4E876C" bold="true" box="[151,289,1876,1900]" pageId="18" pageNumber="59">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C774D1BFFE4C45CFED7F8F3CE88876C" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[295,743,1876,1900]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE4C45CFED7F8F3CE3C876C" box="[295,595,1876,1900]" italics="true" pageId="18" pageNumber="59">Archiinocellia protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE4C45CFD91F8F3CE8F876C" author="Engel, M. S." box="[609,736,1876,1900]" pageId="18" pageNumber="59" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, females. A, holotype USNM 31487 (counterpart); B, UCM 42107. Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF086610FFE5C45DFF67F98ECFD28689" ID-DOI="http://doi.org/10.5281/zenodo.4655677" ID-Zenodo-Dep="4655677" httpUri="https://zenodo.org/record/4655677/files/figure.png" pageId="19" pageNumber="60" startId="19.[151,250,1577,1601]" targetBox="[169,1417,181,1553]" targetPageId="19">
|
||
<paragraph id="8BC83698FFE5C45DFF67F98ECFD28689" blockId="19.[151,1437,1577,1673]" pageId="19" pageNumber="60">
|
||
<emphasis id="B903EA8AFFE5C45DFF67F98ECD4E8641" bold="true" box="[151,289,1577,1601]" pageId="19" pageNumber="60">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C774D1BFFE5C45DFED9F98ECE858641" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[297,746,1577,1601]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="19" pageNumber="60" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE5C45DFED9F98ECE3B8641" box="[297,596,1577,1601]" italics="true" pageId="19" pageNumber="60">Archiinocellia protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE5C45DFD94F98ECE8C8641" author="Engel, M. S." box="[612,739,1577,1601]" pageId="19" pageNumber="60" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, abdomen. A, UCM 61231b, male (lateral/slightly dorsal view); B, UCM 42684, male (lateral view); C, UCM 45968, female. e, ectoproct; gx9?, possible gonocoxites 9; ov, ovipositor; S2 to S7, second to seventh sternites; T3 to T9, third to ninth tergites. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83698FFE5C45DFF37F91FCEA186F5" blockId="19.[151,1437,1719,2034]" pageId="19" pageNumber="60">
|
||
Pronotum dark brown, relatively short,
|
||
<quantity id="4C8F9B7DFFE5C45DFD8FF910CF4D86D2" box="[639,802,1719,1746]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="19" pageNumber="60" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.20–1.50 mm</quantity>
|
||
long; maximum width
|
||
<quantity id="4C8F9B7DFFE5C45DFBD6F910C8E586D2" box="[1062,1162,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="19" pageNumber="60" unit="mm" value="0.57">0.57 mm</quantity>
|
||
in dorsal view,
|
||
<quantity id="4C8F9B7DFFE5C45DFAC9F910C9F286D2" box="[1337,1437,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="60" unit="mm" value="0.65">0.65 mm</quantity>
|
||
in lateral view. Meso- and metathorax dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE5C45DFF37F8A7C8698719" blockId="19.[151,1437,1719,2034]" box="[199,1030,1791,1818]" pageId="19" pageNumber="60">
|
||
Legs pale except brown in proximal half of femora and in coxae (
|
||
<figureCitation id="134C2A1DFFE5C45DFC56F958CF968719" box="[934,1017,1791,1817]" captionStart="FIGURE 12" captionStartId="18.[151,250,1876,1900]" captionTargetBox="[151,1436,705,1840]" captionTargetId="figure-254@18.[151,1436,705,1853]" captionTargetPageId="18" captionText="FIGURE 12. Archiinocellia protomaculata (Engel, 2011), females. A, holotype USNM 31487 (counterpart); B, UCM 42107. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655671" httpUri="https://zenodo.org/record/4655671/files/figure.png" pageId="19" pageNumber="60">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE5C45DFF37F884CFFB87F2" blockId="19.[151,1437,1719,2034]" pageId="19" pageNumber="60">
|
||
Abdomen of male (
|
||
<figureCitation id="134C2A1DFFE5C45DFE52F884CD99873D" box="[418,502,1827,1853]" captionStart="FIGURE 13" captionStartId="19.[151,250,1577,1601]" captionTargetBox="[169,1417,181,1553]" captionTargetId="figure-19@19.[169,1417,181,1553]" captionTargetPageId="19" captionText="FIGURE 13. Archiinocellia protomaculata (Engel, 2011), abdomen. A, UCM 61231b, male (lateral/slightly dorsal view); B, UCM 42684, male (lateral view); C, UCM 45968, female. e, ectoproct; gx9?, possible gonocoxites 9; ov, ovipositor; S2 to S7, second to seventh sternites; T3 to T9, third to ninth tergites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655677" httpUri="https://zenodo.org/record/4655677/files/figure.png" pageId="19" pageNumber="60">Fig. 13</figureCitation>
|
||
): Tergites, sternites dark, easily distinguished; T3 to T7, and S3 to S7 nearly equal, moderately long; T8 slightly shorter than T3 to T7, extending laterally notably more than these tergites; T9 short, extending laterally to ventral margin; ectoproct short, elongate in lateral view, ventrally appears narrowed; gonocoxites 9 not discernible (as in
|
||
<figureCitation id="134C2A1DFFE5C45DFE06F828CE2287A9" box="[502,589,1935,1961]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[206,1380,192,1913]" captionTargetId="figure-26@4.[195,1393,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Wing venation of Megaraphidia antiquissima sp. nov., holotype RBCM P1555. A, right forewing, B, left forewing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655631" httpUri="https://zenodo.org/record/4655631/files/figure.png" pageId="19" pageNumber="60">Fig. 2A</figureCitation>
|
||
) or apparently weakly sclerotized (see
|
||
<figureCitation id="134C2A1DFFE5C45DFBF6F828C80387A9" box="[1030,1132,1935,1961]" captionStart="FIGURE 13" captionStartId="19.[151,250,1577,1601]" captionTargetBox="[169,1417,181,1553]" captionTargetId="figure-19@19.[169,1417,181,1553]" captionTargetPageId="19" captionText="FIGURE 13. Archiinocellia protomaculata (Engel, 2011), abdomen. A, UCM 61231b, male (lateral/slightly dorsal view); B, UCM 42684, male (lateral view); C, UCM 45968, female. e, ectoproct; gx9?, possible gonocoxites 9; ov, ovipositor; S2 to S7, second to seventh sternites; T3 to T9, third to ninth tergites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655677" httpUri="https://zenodo.org/record/4655677/files/figure.png" pageId="19" pageNumber="60">Fig. 13B</figureCitation>
|
||
). Abdomen of female (
|
||
<figureCitation id="134C2A1DFFE5C45DFA9FF828CCA587CD" captionStart="FIGURE 13" captionStartId="19.[151,250,1577,1601]" captionTargetBox="[169,1417,181,1553]" captionTargetId="figure-19@19.[169,1417,181,1553]" captionTargetPageId="19" captionText="FIGURE 13. Archiinocellia protomaculata (Engel, 2011), abdomen. A, UCM 61231b, male (lateral/slightly dorsal view); B, UCM 42684, male (lateral view); C, UCM 45968, female. e, ectoproct; gx9?, possible gonocoxites 9; ov, ovipositor; S2 to S7, second to seventh sternites; T3 to T9, third to ninth tergites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655677" httpUri="https://zenodo.org/record/4655677/files/figure.png" pageId="19" pageNumber="60">Fig. 13C</figureCitation>
|
||
): Tergites, sternites dark, easily distinguished; T8, T9 relatively short; ectoproct small, elongate in lateral view; ovipositor moderately long,
|
||
<quantity id="4C8F9B7DFFE5C45DFE25F870CE1787F2" box="[469,632,2007,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.525" metricValueMax="3.8" metricValueMin="3.25" pageId="19" pageNumber="60" unit="mm" value="3.525" valueMax="3.8" valueMin="3.25">3.25–3.80 mm</quantity>
|
||
(0.27–0.30 body length).
|
||
</paragraph>
|
||
<caption id="DF086610FFE2C45AFF67FA08CFD085EB" ID-DOI="http://doi.org/10.5281/zenodo.4655680" ID-Zenodo-Dep="4655680" httpUri="https://zenodo.org/record/4655680/files/figure.png" pageId="20" pageNumber="61" startId="20.[151,250,1455,1479]" targetBox="[151,1436,181,1431]" targetPageId="20">
|
||
<paragraph id="8BC83698FFE2C45AFF67FA08CFD085EB" blockId="20.[151,1436,1455,1515]" pageId="20" pageNumber="61">
|
||
<emphasis id="B903EA8AFFE2C45AFF67FA08CD7085C7" bold="true" box="[151,287,1455,1479]" pageId="20" pageNumber="61">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C774D1BFFE2C45AFED5FA08CE8E85C7" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[293,737,1455,1479]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="61" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE2C45AFED5FA08CE2085C7" box="[293,591,1455,1479]" italics="true" pageId="20" pageNumber="61">Archiinocellia protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE2C45AFDACFA08CEB585C7" author="Engel, M. S." box="[604,730,1455,1479]" pageId="20" pageNumber="61" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A, UCM 61231, rock with four specimens of
|
||
<taxonomicName id="4C774D1BFFE2C45AFB43FA17C90D85C7" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[1203,1378,1455,1479]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="61" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE2C45AFB43FA17C90D85C7" box="[1203,1378,1455,1479]" italics="true" pageId="20" pageNumber="61">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
(a–c) and one undeterminable
|
||
<taxonomicName id="4C774D1BFFE2C45AFE61FA74CE4E85EB" authorityName="Navas" authorityYear="1916" box="[401,545,1491,1515]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="61" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
|
||
(d); B, UCM 62523. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83698FFE2C45BFF37F9BECE0381F5" blockId="20.[151,1437,1561,2020]" lastBlockId="21.[151,1437,151,501]" lastPageId="21" lastPageNumber="62" pageId="20" pageNumber="61">
|
||
Forewing relatively narrow, 7.4–8.0 mm long,
|
||
<quantity id="4C8F9B7DFFE2C45AFD12F9BDCF048634" box="[738,875,1562,1588]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="20" pageNumber="61" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
|
||
wide (length/width: 3.63–4.15, measured in five females). Costal space narrow, most dilated at proximal 1/6 wing length; at maximum four subcostal veinlets simple preserved (basal regions not preserved, probably six total), widely spaced. ScP very short, terminating on costal margin approximately at just basad mid-wing, strongly distad crossvein 2ra-rp; from termination of ScP to pterostigma approximately double length of pterostigma. Subcostal space: basal crossvein 1scp-r closer to divergence of M, CuA than to origin of RP; distal crossvein (2scp-r, forming basal margin of pterostigma) straight to incurved. Pterostigma elongate (4.9–5.1 times longer than width measured along to costal margin), lightly pigmented (from very pale to slightly brownish as preserved). Two branches of RA: one within pterostigma usually easily visible, strongly inclined, in middle of pterostigma; branch forming distal margin of pterostigma usually straight; no branch distad pterostigma; portion of RA distad pterostigma short. RA space with two crossveins forming two radial cells nearly equal in length: 2ra-rp usually relatively short, located about mid-way between pterostigma, termination of ScP; 3ra-rp located in distal part of pterostigma along RA. RP originating at nearly half (0.43–0.48) wing length, usually with four pectinate branches (rarely three): RP1 deeply forked (at about half or somewhat less its length), with anterior branch simple, posterior forked; other branches (RP2 to RP4) simple. One long intraradial crossvein rp1-rp2 between stems of RP, RP1. Three crossveins between RP, MA: 1r-m, 2r-m connecting stem of RP, MA; 3r-m connecting RP1, anterior branch of MA. M close to R (appears fused) for considerable distance, and then fused with CuA for short distance; forked well proximad origin of RP. MA deeply dichotomously forked twice. MP strongly zigzagged, anterior trace with two (rarely one) simple branches. Two intramedian crossveins form two
|
||
<emphasis id="B903EA8AFFE3C45BFAFCFE80C95F8141" box="[1292,1328,295,321]" italics="true" pageId="21" pageNumber="62">doi</emphasis>
|
||
, approximately equal in length (sometimes distal
|
||
<emphasis id="B903EA8AFFE3C45BFD95FEECCEE68165" box="[613,649,331,357]" italics="true" pageId="21" pageNumber="62">doi</emphasis>
|
||
longer): 2im very long, joining posterior branch of MA near origin or at forking of MA. CuA fused with M for short distance; anterior trace CuA strongly zigzagged, fused with MP for long distance to form part of posterior margin of basal
|
||
<emphasis id="B903EA8AFFE3C45BFCF0FE34CF4B81AD" box="[768,804,403,429]" italics="true" pageId="21" pageNumber="62">doi</emphasis>
|
||
, with one long simple branch. CuP simple, aligned with Cu. Crossvein icu between CuA, CuP long. Crossveins between CuP, AA1, between AA1, AA2 short. AA1, AA2 simple; region of AA3 poorly discernible.
|
||
</paragraph>
|
||
<caption id="DF086610FFE3C45BFF67FA16CF9085EC" ID-DOI="http://doi.org/10.5281/zenodo.4655684" ID-Zenodo-Dep="4655684" httpUri="https://zenodo.org/record/4655684/files/figure.png" pageId="21" pageNumber="62" startId="21.[151,250,1457,1481]" targetBox="[151,1436,525,1433]" targetPageId="21">
|
||
<paragraph id="8BC83698FFE3C45BFF67FA16CF9085EC" blockId="21.[151,1436,1457,1517]" pageId="21" pageNumber="62">
|
||
<emphasis id="B903EA8AFFE3C45BFF67FA16CD4F85C9" bold="true" box="[151,288,1457,1481]" pageId="21" pageNumber="62">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C774D1BFFE3C45BFED6FA16CE8B85C9" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[294,740,1457,1481]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="21" pageNumber="62" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE3C45BFED6FA16CE3E85C9" box="[294,593,1457,1481]" italics="true" pageId="21" pageNumber="62">Archiinocellia protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE3C45BFDAFFA16CEB285C9" author="Engel, M. S." box="[607,733,1457,1481]" pageId="21" pageNumber="62" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, head and prothorax (dorsal view). A, UCM 64727; B, UCM 62525; C, UCM 35717; D, UCM 45969; E, UCM 42122; F, UCM 62687. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83698FFE3C458FF37F9BCC96E811E" blockId="21.[151,1437,1563,2022]" lastBlockId="22.[151,1437,151,286]" lastPageId="22" lastPageNumber="63" pageId="21" pageNumber="62">
|
||
Hind wing
|
||
<quantity id="4C8F9B7DFFE3C45BFEB8F9BCCDBF8636" box="[328,464,1563,1590]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.1" metricValueMin="6.5" pageId="21" pageNumber="62" unit="mm" value="6.8" valueMax="7.1" valueMin="6.5">6.5–7.1 mm</quantity>
|
||
long,
|
||
<quantity id="4C8F9B7DFFE3C45BFDE9F9BCCE1F8636" box="[537,624,1563,1590]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="62" unit="mm" value="1.7">1.7 mm</quantity>
|
||
wide (length/width: 3.82). Costal space narrow; at maximum three discernible simple subcostal veinlets, very widely spaced (slightly wider than in forewing). ScP very short, terminating at approximately mid-wing; from termination of ScP to pterostigma approximately twice length of pterostigma. One crossvein detected in subcostal space, 2scp-r, forming basal margin of pterostigma, straight to incurved. Pterostigma elongate (4.5–5.1 times longer than width measured along to costal margin). RA with two branches: one within pterostigma usually clearly visible, strongly inclined, located medially in pterostigma; branch forming distal margin of pterostigma usually straight; no branches distad pterostigma; portion of RA distad pterostigma short. Three crossveins between RA, RP: 1ra-rp located slightly proximad 2r-m (rarer probably distad:
|
||
<figureCitation id="134C2A1DFFE3C45BFB77F8B0C8838731" box="[1159,1260,1815,1841]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="21" pageNumber="62">Fig. 17B</figureCitation>
|
||
); 2ra-rp located far distad termination of ScP; 3ra-rp located just distad middle of pterostigma along RA. RP originating 0.33 wing length, with three to four pectinate branches, two proximad 3ra-rp, one to two distad. RP1 deeply forked, anterior branch simple, posterior forked; other branches (RP2 to RP4) simple. One intraradial crossvein rp1-rp2 between stems of RP, RP1. Three crossveins between R, M: 1r-m long, subparallel to R connecting to RP near its origin (its base unclear); 2r-m connecting stems of RP, MA; 3r-m connecting RP1, anterior branch of MA. M forked distad origin of RP. MA deeply dichotomously forked twice. MP: anterior trace zigzagged, simple, with three simple, pectinate branches. Two intramedian crossveins form two
|
||
<emphasis id="B903EA8AFFE0C458FCDDFF1CCF3E80D5" box="[813,849,187,213]" italics="true" pageId="22" pageNumber="63">doi</emphasis>
|
||
, proximal much shorter than distal. Two crossveins between M, Cu: basal 1m-cu relatively short, oblique, connecting M, CuA; 2m-cu connecting MP, CuA. Anterior trace of CuA simple distally, with one simple branch. CuP fused with AA1 (incompletely preserved).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF086610FFE0C458FF67F930CEB486D3" ID-DOI="http://doi.org/10.5281/zenodo.4655686" ID-Zenodo-Dep="4655686" httpUri="https://zenodo.org/record/4655686/files/figure.png" pageId="22" pageNumber="63" startId="22.[151,250,1687,1711]" targetBox="[160,1427,326,1646]" targetPageId="22">
|
||
<paragraph id="8BC83698FFE0C458FF67F930CEB486D3" blockId="22.[151,1436,1687,1747]" pageId="22" pageNumber="63">
|
||
<emphasis id="B903EA8AFFE0C458FF67F930CD4F86AF" bold="true" box="[151,288,1687,1711]" pageId="22" pageNumber="63">FIGURE 16.</emphasis>
|
||
Venation of
|
||
<taxonomicName id="4C774D1BFFE0C458FE53F930CF0E86AF" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[419,865,1687,1711]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE0C458FE53F930CEA286AF" box="[419,717,1687,1711]" italics="true" pageId="22" pageNumber="63">Archiinocellia protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE0C458FD2BF930CF3586AF" author="Engel, M. S." box="[731,858,1687,1711]" pageId="22" pageNumber="63" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, holotype USNM 31487. A, right forewing; B, left forewing; C, right hind wing. Scale bar = 3 mm (all to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36D6513FFE0C459FF37F8A6C8268436" lastPageId="23" lastPageNumber="64" pageId="22" pageNumber="63" type="discussion">
|
||
<paragraph id="8BC83698FFE0C459FF37F8A6CE7B81AE" blockId="22.[151,1437,1793,2036]" lastBlockId="23.[151,1437,151,1078]" lastPageId="23" lastPageNumber="64" pageId="22" pageNumber="63">
|
||
<emphasis id="B903EA8AFFE0C458FF37F8A6CD54871B" bold="true" box="[199,315,1793,1819]" pageId="22" pageNumber="63">Remarks.</emphasis>
|
||
This species was described based on
|
||
<specimenCount id="9D71FD11FFE0C458FD1CF8A6CFE9871B" box="[748,902,1793,1819]" pageId="22" pageNumber="63" type="generic">19 specimens</specimenCount>
|
||
from the National Museum of Natural History (
|
||
<bibRefCitation id="EFE64B69FFE0C458FF6FF882CD43873F" author="Engel, M. S." box="[159,300,1829,1855]" pageId="22" pageNumber="63" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
). The specimen mentioned and figured by
|
||
<bibRefCitation id="EFE64B69FFE0C458FCFAF882CF928740" author="Dayvault, R. D. & Codington, L. A. & Kohls, D. & Hawes, W. D. & Ott, P. M." box="[778,1021,1829,1856]" pageId="22" pageNumber="63" pagination="97 - 115" refId="ref22558" refString="Dayvault, R. D., Codington, L. A., Kohls, D., Hawes, W. D. & Ott, P. M. (1995) Fossil insects and spiders from three locations in the Green River Formation of the Piceance Creek Basin, Colorado. In: Averett, W. R. (Ed.), The Green River Formation in Piceance Creek and Eastern Uinta Basins. Grand Junction Geological Society, Grand Junction, Colorado, pp. 97 - 115." type="book chapter" year="1995">
|
||
Dayvault
|
||
<emphasis id="B903EA8AFFE0C458FC88F881CFC6873F" box="[888,937,1829,1855]" italics="true" pageId="22" pageNumber="63">et al</emphasis>
|
||
. (1995
|
||
</bibRefCitation>
|
||
: Fig. 35) belongs to this species (
|
||
<bibRefCitation id="EFE64B69FFE0C458FA82F882CD6D8763" author="Engel, M. S." pageId="22" pageNumber="63" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
), as does
|
||
<bibRefCitation id="EFE64B69FFE0C458FE9CF8EECE4E8763" author="Aspock, H." box="[364,545,1865,1891]" pageId="22" pageNumber="63" pagination="33 - 44" refId="ref20619" refString="Aspock, H. (1998) Distribution and biogeography of the order Raphidioptera: updated facts and a new hypothesis. Acta Zoologica Fennica, 209, 33 - 44." type="journal article" year="1998">Aspöck’s (1998</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE0C458FDDBF8EECE018764" box="[555,622,1865,1892]" captionStart="FIGURE 7" captionStartId="11.[151,250,1581,1605]" captionTargetBox="[151,1430,1137,1553]" captionTargetId="figure-445@11.[151,1436,1137,1558]" captionTargetPageId="11" captionText="FIGURE 7. Megaraphidia klondika sp. nov., specimen SR16-004-001-01.01. A, specimen as preserved; B, drawing. Scale bar = 3 mm (both to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655655" httpUri="https://zenodo.org/record/4655655/files/figure.png" pageId="22" pageNumber="63">Fig. 7</figureCitation>
|
||
) specimen (determination here by head maculation). We examined highquality photographs of
|
||
<specimenCount id="9D71FD11FFE0C458FE6EF8CACE588787" box="[414,567,1901,1928]" pageId="22" pageNumber="63" type="generic">51 specimens</specimenCount>
|
||
in the UCM collections. Most are only partially visible pending preparation, as the rock in these beds does not split along fine, thin laminae bearing the fossil as is usually the case in Okanagan Highlands fossils, and portions often protrude into the matrix. We confidently assign 36 of these to
|
||
<taxonomicName id="4C774D1BFFE0C458FB2BF811C9F387CF" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[1243,1436,1973,2000]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE0C458FB2BF811C9F387CF" box="[1243,1436,1973,2000]" italics="true" pageId="22" pageNumber="63">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
by their characteristic head maculation and/or wing venation. Five further fossils possibly belong to this species, but cannot be assigned with certainty by preservation: three females from the Labandeira Site (UCM 60298; UCM 62430; UCM 65597), a probable male (UCM 42748) from the Donnell Site, and a female (UCM 75788) from the UCM locality 2009063 (Claudia’s Place in Garfield County). Characters present on the remaining ten more poorly preserved specimens suggest that at least some of them might also belong to the species, including four from the Labandeira Site: UCM 61231d (sex unknown); UCM 61320 (male), UCM 62686 (sex unknown), UCM 62689 (sex unknown), UCM 68352 (sex unknown); two from the Donnell Site: UCM 39661 (sex unknown) and UCM 39662 (female); two from Claudia’s Place: UCM 75468 (sex unknown) and UCM 75790 (male); and one from the Denson Site: UCM 56842 (sex unknown).
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE1C459FF37FE10CD1E82A9" blockId="23.[151,1437,151,1078]" pageId="23" pageNumber="64">
|
||
The maculation of the head in
|
||
<taxonomicName id="4C774D1BFFE1C459FDEBFE10CF0A81D1" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[539,869,439,465]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE1C459FDEBFE10CF0A81D1" box="[539,869,439,465]" italics="true" pageId="23" pageNumber="64">Archiinocellia protomaculata</emphasis>
|
||
</taxonomicName>
|
||
is distinctive (
|
||
<figureCitation id="134C2A1DFFE1C459FBF9FE10C83381D1" box="[1033,1116,439,465]" captionStart="FIGURE 14" captionStartId="20.[151,250,1455,1479]" captionTargetBox="[151,1436,181,1431]" captionTargetId="figure-26@20.[151,1436,181,1431]" captionTargetPageId="20" captionText="FIGURE 14. Archiinocellia protomaculata (Engel, 2011).A, UCM 61231, rock with four specimens of A. protomaculata (a–c) and one undeterminable Raphidioptera (d); B, UCM 62523. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4655680" httpUri="https://zenodo.org/record/4655680/files/figure.png" pageId="23" pageNumber="64">Fig. 14</figureCitation>
|
||
). Paler areas occupy most of the lateral part of its head, whereas there are paler streaks in some extant genera of both families; others are mostly dark, lacking reddish regions (see
|
||
<emphasis id="B903EA8AFFE1C459FDEAFE58CE548218" box="[538,571,511,536]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<bibRefCitation id="EFE64B69FFE1C459FDBFFE58CE828219" author="Albarda, H." box="[591,749,511,537]" pageId="23" pageNumber="64" pagination="65 - 184" refId="ref19852" refString="Albarda, H. (1891) Revision des Rhaphidides. Tijdschrift voor Entomologie, 34, 65 - 184." type="journal article" year="1891">Albarda 1891</bibRefCitation>
|
||
). While the life colour of these pale regions of the head of
|
||
<taxonomicName id="4C774D1BFFE1C459FA74FE58CD54823D" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE1C459FA74FE58CD54823D" italics="true" pageId="23" pageNumber="64">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
is unknown, lighter regions in the heads of extant species are reddish.
|
||
<bibRefCitation id="EFE64B69FFE1C459FB90FD84C94E823E" author="Carpenter, F. M." box="[1120,1313,547,574]" pageId="23" pageNumber="64" pagination="89 - 157" refId="ref21922" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157. https: // doi. org / 10.2307 / 20023217" type="journal article" year="1936">Carpenter (1936)</bibRefCitation>
|
||
stated that the size of the reddish patches varies freely within
|
||
<taxonomicName id="4C774D1BFFE1C459FD3DFDE0CF3B8261" authorityName="Latreille" authorityYear="1810" box="[717,852,583,609]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
||
species (p. 9), but the colour patterning of the heads of
|
||
<taxonomicName id="4C774D1BFFE1C459FF44FDCCCD198285" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[180,374,619,645]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE1C459FF44FDCCCD198285" box="[180,374,619,645]" italics="true" pageId="23" pageNumber="64">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
is strongly consistent among individuals, and so we consider it to be a good diagnostic character state of the species.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE1C459FF37FD14C8D58315" blockId="23.[151,1437,151,1078]" pageId="23" pageNumber="64">
|
||
The maculation of the
|
||
<typeStatus id="54CC883AFFE1C459FE35FD14CE4882CD" box="[453,551,691,717]" pageId="23" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
head is typical of the species, but its wings are overlapping and many portions are obscured by the abdomen (
|
||
<figureCitation id="134C2A1DFFE1C459FE18FD70CE2182F1" box="[488,590,727,753]" captionStart="FIGURE 12" captionStartId="18.[151,250,1876,1900]" captionTargetBox="[151,1436,705,1840]" captionTargetId="figure-254@18.[151,1436,705,1853]" captionTargetPageId="18" captionText="FIGURE 12. Archiinocellia protomaculata (Engel, 2011), females. A, holotype USNM 31487 (counterpart); B, UCM 42107. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655671" httpUri="https://zenodo.org/record/4655671/files/figure.png" pageId="23" pageNumber="64">Fig. 12A</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFE1C459FDA8FD70CE8E82F1" author="Engel, M. S." box="[600,737,727,753]" pageId="23" pageNumber="64" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE1C459FD1AFD70CF4182F1" box="[746,814,727,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="23" pageNumber="64">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="134C2A1DFFE1C459FCCBFD70CF2382F1" box="[827,844,727,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[206,1380,192,1913]" captionTargetId="figure-26@4.[195,1393,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Wing venation of Megaraphidia antiquissima sp. nov., holotype RBCM P1555. A, right forewing, B, left forewing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655631" httpUri="https://zenodo.org/record/4655631/files/figure.png" pageId="23" pageNumber="64">2</figureCitation>
|
||
). The venation of both the fore- and hind wings (
|
||
<figureCitation id="134C2A1DFFE1C459FA94FD70CC858315" captionStart="FIGURE 16" captionStartId="22.[151,250,1687,1711]" captionTargetBox="[160,1427,326,1646]" captionTargetId="figure-89@22.[151,1436,309,1664]" captionTargetPageId="22" captionText="FIGURE 16. Venation of Archiinocellia protomaculata (Engel, 2011), holotype USNM 31487. A, right forewing; B, left forewing; C, right hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655686" httpUri="https://zenodo.org/record/4655686/files/figure.png" pageId="23" pageNumber="64">Figs. 16A, B</figureCitation>
|
||
), is generally similar to that of other examined specimens (
|
||
<emphasis id="B903EA8AFFE1C459FC72FD5CCFCB8314" box="[898,932,763,788]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<figureCitation id="134C2A1DFFE1C459FC49FD5CC8248315" box="[953,1099,763,789]" captionStart="FIGURE 16" captionStartId="22.[151,250,1687,1711]" captionTargetBox="[160,1427,326,1646]" captionTargetId="figure-89@22.[151,1436,309,1664]" captionTargetPageId="22" captionText="FIGURE 16. Venation of Archiinocellia protomaculata (Engel, 2011), holotype USNM 31487. A, right forewing; B, left forewing; C, right hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655686" httpUri="https://zenodo.org/record/4655686/files/figure.png" pageId="23" pageNumber="64">Figs. 16C–D</figureCitation>
|
||
;
|
||
<figureCitation id="134C2A1DFFE1C459FBA7FD5CC8C18315" box="[1111,1198,763,789]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">17A–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE1C459FF37FCB8C8268436" blockId="23.[151,1437,151,1078]" pageId="23" pageNumber="64">
|
||
Wing venation shows little variation among the specimens examined. There are usually two branches of RP distad 3ra-rp, but in one specimen there is only one (
|
||
<figureCitation id="134C2A1DFFE1C459FD29FCE4CF06835D" box="[729,873,835,861]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Figs. 17C–D</figureCitation>
|
||
) in both the forewing and hind wing. The length of 3ra-rp varies from long (
|
||
<emphasis id="B903EA8AFFE1C459FE5DFCC0CDA18380" box="[429,462,871,896]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<figureCitation id="134C2A1DFFE1C459FE14FCC0CE088381" box="[484,615,871,897]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17B–C</figureCitation>
|
||
) to short (
|
||
<emphasis id="B903EA8AFFE1C459FD2DFCC0CF6A8380" box="[733,773,871,896]" italics="true" pageId="23" pageNumber="64">e.g.</emphasis>
|
||
,
|
||
<figureCitation id="134C2A1DFFE1C459FCE3FCC0CFCA8381" box="[787,933,871,897]" captionStart="FIGURE 16" captionStartId="22.[151,250,1687,1711]" captionTargetBox="[160,1427,326,1646]" captionTargetId="figure-89@22.[151,1436,309,1664]" captionTargetPageId="22" captionText="FIGURE 16. Venation of Archiinocellia protomaculata (Engel, 2011), holotype USNM 31487. A, right forewing; B, left forewing; C, right hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655686" httpUri="https://zenodo.org/record/4655686/files/figure.png" pageId="23" pageNumber="64">Figs. 16A, C</figureCitation>
|
||
), also in both wings. In forewings, crossvein 2im usually joins the posterior branch of MA (
|
||
<emphasis id="B903EA8AFFE1C459FD53FC2CCEAB83A4" box="[675,708,907,932]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<figureCitation id="134C2A1DFFE1C459FD2AFC2CCF5083A5" box="[730,831,907,933]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17C</figureCitation>
|
||
), but may rarely be at its forking (
|
||
<emphasis id="B903EA8AFFE1C459FB31FC2CC88C83A4" box="[1217,1251,907,932]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<figureCitation id="134C2A1DFFE1C459FB08FC2CC93083A5" box="[1272,1375,907,933]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17A</figureCitation>
|
||
), and while MP usually has two branches, it may rarely have one (
|
||
<bibRefCitation id="EFE64B69FFE1C459FCB2FC08CFA183C9" author="Engel, M. S." box="[834,974,943,969]" pageId="23" pageNumber="64" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE1C459FC2AFC08C84D83C9" box="[986,1058,943,969]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1407,190,1909]" captionTargetId="figure-26@6.[165,1423,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Wing venation of Megaraphidia ootsa sp. nov., holotype RBCM P1556. A, right forewing, B, left forewing; C, right hind wing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655637" httpUri="https://zenodo.org/record/4655637/files/figure.png" pageId="23" pageNumber="64">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="134C2A1DFFE1C459FBDEFC08C85183C9" box="[1070,1086,943,969]" captionStart="FIGURE 7" captionStartId="11.[151,250,1581,1605]" captionTargetBox="[151,1430,1137,1553]" captionTargetId="figure-445@11.[151,1436,1137,1558]" captionTargetPageId="11" captionText="FIGURE 7. Megaraphidia klondika sp. nov., specimen SR16-004-001-01.01. A, specimen as preserved; B, drawing. Scale bar = 3 mm (both to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655655" httpUri="https://zenodo.org/record/4655655/files/figure.png" pageId="23" pageNumber="64">7</figureCitation>
|
||
). In hind wings, the size of the basal
|
||
<emphasis id="B903EA8AFFE1C459FF29FC74CC9283ED" box="[217,253,979,1005]" italics="true" pageId="23" pageNumber="64">doi</emphasis>
|
||
varies from relatively long (
|
||
<figureCitation id="134C2A1DFFE1C459FDB6FC74CEC283ED" box="[582,685,979,1005]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17D</figureCitation>
|
||
) to short (
|
||
<figureCitation id="134C2A1DFFE1C459FCD5FC74CFE383ED" box="[805,908,979,1005]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17B</figureCitation>
|
||
), and crossvein 2r-m may be placed relatively distally (distad 2ra-rp:
|
||
<figureCitation id="134C2A1DFFE1C459FE6AFC50CE6E8411" box="[410,513,1015,1041]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17D</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFE1C459FDFEFC50CEF68411" author="Engel, M. S." box="[526,665,1015,1041]" pageId="23" pageNumber="64" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE1C459FD54FC50CE878411" box="[676,744,1015,1041]" captionStart="FIGURE 7" captionStartId="11.[151,250,1581,1605]" captionTargetBox="[151,1430,1137,1553]" captionTargetId="figure-445@11.[151,1436,1137,1558]" captionTargetPageId="11" captionText="FIGURE 7. Megaraphidia klondika sp. nov., specimen SR16-004-001-01.01. A, specimen as preserved; B, drawing. Scale bar = 3 mm (both to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655655" httpUri="https://zenodo.org/record/4655655/files/figure.png" pageId="23" pageNumber="64">Fig. 7</figureCitation>
|
||
) to proximally (proximad 2ra-rp:
|
||
<figureCitation id="134C2A1DFFE1C459FB99FC50C8A18411" box="[1129,1230,1015,1041]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="23" pageNumber="64">Fig. 17B</figureCitation>
|
||
). These variations are normal for the suborder, and in some species are greater (see
|
||
<emphasis id="B903EA8AFFE1C459FC9FFBBCCFFF8434" box="[879,912,1051,1076]" italics="true" pageId="23" pageNumber="64">e.g</emphasis>
|
||
.,
|
||
<bibRefCitation id="EFE64B69FFE1C459FC55FBBCC8528436" author="Zeleny, J." box="[933,1085,1051,1078]" pageId="23" pageNumber="64" pagination="15 - 38" refId="ref26145" refString="Zeleny, J. (1969) Variability in the species Raphidia ophiopsis Linnaeus, and Agulla xanthostigma (Schummel) (Raphidioptera). Acta Entomologica Bohemoslovaca, 66, 15 - 38." type="journal article" year="1969">Zelený, 1969</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |