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<document id="25DBBBBCED0AE8A018ED894ECD86B661" ID-DOI="10.11646/zootaxa.4880.1.1" ID-GBIF-Dataset="50f2ef69-fec5-45e5-8005-623e2cb991dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425522" ID-ZooBank="33BE91BC-DC0F-4CBB-85AB-CA7BF1891C0C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610041656870" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2020" docId="03DE878CFFE2FFCAFF3B723AD568FA78" docLanguage="en" docName="zootaxa.4880.1.1.pdf.imf" docOrigin="Zootaxa 4880 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ariasa Distant 1905" docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="FFE7FFF4FFF8FFD0FFAC7050D010FFCE" masterDocTitle="The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records" masterLastPageNumber="80" masterPageNumber="1" pageNumber="27" updateTime="1700279468176" updateUser="plazi">
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<mods:title id="F69164D76CD5325E701F957D0E19F920">The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records</mods:title>
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<mods:namePart id="8CFF2F24774AB89FB098204AC0DEA9A6">Sanborn, Allen F.</mods:namePart>
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<mods:date id="FEE0DACC98C17BC44EDF65D3CD8E08D1">2020</mods:date>
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<treatment id="03DE878CFFE2FFCAFF3B723AD568FA78" ID-DOI="http://doi.org/10.5281/zenodo.4573734" ID-GBIF-Taxon="180323579" ID-Zenodo-Dep="4573734" LSID="urn:lsid:plazi:treatment:03DE878CFFE2FFCAFF3B723AD568FA78" httpUri="http://treatment.plazi.org/id/03DE878CFFE2FFCAFF3B723AD568FA78" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection id="C36D6511FFE2FFCAFF3B723AD29DFD4B" box="[151,653,618,645]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BC8369AFFE2FFCAFF3B723AD29DFD4B" blockId="26.[151,653,618,645]" box="[151,653,618,645]" pageId="26" pageNumber="27">
<emphasis id="B903EA88FFE2FFCAFF3B723AD0F7FD4A" bold="true" box="[151,231,618,644]" pageId="26" pageNumber="27">Genus</emphasis>
<taxonomicName id="4C774D19FFE2FFCAFF42723BD1E9FD4A" ID-CoL="8HCS4" ID-ENA="2218926" authority="Distant, 1905" authorityName="Distant" authorityYear="1905" box="[238,505,618,645]" class="Insecta" family="Cicadidae" genus="Ariasa" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFF42723BD12CFD4B" box="[238,316,619,645]" italics="true" pageId="26" pageNumber="27">Ariasa</emphasis>
<bibRefCitation id="EFE64B6BFFE2FFCAFEE8723AD1E9FD4A" author="Distant, W. L." box="[324,505,618,644]" pageId="26" pageNumber="27" pagination="304 - 319" refId="ref69306" refString="Distant, W. L. (1905 d) Rhynchotal notes XXX. Annals and Magazine of Natural History, Series 7, 15, 304 - 319. https: // doi. org / 10.1080 / 03745480509443047" type="journal article" year="1905">
<emphasis id="B903EA88FFE2FFCAFEE8723AD1E9FD4A" bold="true" box="[324,505,618,644]" pageId="26" pageNumber="27">Distant, 1905d</emphasis>
</bibRefCitation>
</taxonomicName>
<emphasis id="B903EA88FFE2FFCAFDAC723BD29DFD4B" bold="true" box="[512,653,619,645]" pageId="26" pageNumber="27">new record</emphasis>
</paragraph>
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<subSubSection id="C36D6511FFE2FFCAFF3B72FFD1BAFD09" box="[151,426,687,711]" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="8BC8369AFFE2FFCAFF3B72FFD1BAFD09" blockId="26.[151,888,687,742]" box="[151,426,687,711]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB6711B4FFE2FFCAFF3B72FFD1BAFD09" box="[151,426,687,711]" pageId="26" pageNumber="27">
<treatmentCitation id="0AD6108BFFE2FFCAFF3B72FFD1B6FD09" author="Distant, W. L." box="[151,422,687,711]" page="314" pageId="26" pageNumber="27" year="1905">
<taxonomicName id="4C774D19FFE2FFCAFF3B72FFD1B6FD09" ID-CoL="8HCS4" authority="Distant, 1905" authorityName="Distant" authorityPageNumber="314" authorityYear="1905" box="[151,422,687,711]" class="Insecta" family="Cicadidae" genus="Ariasa" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFF3B72FFD0CBFD08" box="[151,219,687,710]" italics="true" pageId="26" pageNumber="27">Ariasa</emphasis>
<bibRefCitation id="EFE64B6BFFE2FFCAFF4D72FFD1B6FD09" author="Distant, W. L." box="[225,422,687,711]" pageId="26" pageNumber="27" pagination="304 - 319" refId="ref69306" refString="Distant, W. L. (1905 d) Rhynchotal notes XXX. Annals and Magazine of Natural History, Series 7, 15, 304 - 319. https: // doi. org / 10.1080 / 03745480509443047" type="journal article" year="1905">Distant 1905d: 314</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D6511FFE2FFCAFF3B729ED368FD28" box="[151,888,718,742]" pageId="26" pageNumber="27" type="type_taxon">
<paragraph id="8BC8369AFFE2FFCAFF3B729ED368FD28" blockId="26.[151,888,687,742]" box="[151,888,718,742]" pageId="26" pageNumber="27">
<typeStatus id="54CC8838FFE2FFCAFF3B729ED0D6FD2A" box="[151,198,718,741]" pageId="26" pageNumber="27">TYPE</typeStatus>
SPECIES.—
<taxonomicName id="4C774D19FFE2FFCAFE9D729FD364FD28" ID-CoL="7DB3W" authority="Distant, 1892" authorityName="Distant" authorityPageNumber="60" authorityYear="1892" box="[305,884,718,742]" class="Insecta" family="Cicadidae" genus="Tympanoterpes" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="colombiae">
<treatmentCitationGroup id="AB6711B4FFE2FFCAFE9D729FD2E6FD28" box="[305,758,718,742]" pageId="26" pageNumber="27">
<emphasis id="B903EA88FFE2FFCAFE9D729FD22CFD28" box="[305,572,718,742]" italics="true" pageId="26" pageNumber="27">Tympanoterpes colombiae</emphasis>
<treatmentCitation id="0AD6108BFFE2FFCAFDEE729ED2E6FD28" author="Distant, W. L." box="[578,758,718,742]" page="60" pageId="26" pageNumber="27" year="1892">
<bibRefCitation id="EFE64B6BFFE2FFCAFDEE729ED2E6FD28" author="Distant, W. L." box="[578,758,718,742]" pageId="26" pageNumber="27" pagination="54 - 67" refId="ref69055" refString="Distant, W. L. (1892 a) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant 1892a: 60</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
(
<collectingCountry id="F360760AFFE2FFCAFCAF729ED37DFD28" box="[771,877,718,742]" name="Colombia" pageId="26" pageNumber="27">Colombia</collectingCountry>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6511FFE2FFCAFF3B7345D489FA85" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BC8369AFFE2FFCAFF3B7345D3FFFB58" blockId="26.[151,1437,788,1463]" pageId="26" pageNumber="27">
REMARKS.—
<taxonomicName id="4C774D19FFE2FFCAFEB37344D17AFCE0" authorityName="Distant" authorityYear="1905" box="[287,362,788,814]" class="Insecta" family="Cicadidae" genus="Ariasa" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFEB37344D17AFCE0" box="[287,362,788,814]" italics="true" pageId="26" pageNumber="27">Ariasa</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C774D19FFE2FFCAFD0B7344D316FCE0" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[679,774,788,814]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFD0B7344D316FCE0" box="[679,774,788,814]" italics="true" pageId="26" pageNumber="27">Fidicina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D19FFE2FFCAFCEE7344D3DEFCE0" authorityName="Boulard &amp; Martinelli" authorityYear="1996" box="[834,974,788,814]" class="Insecta" family="Cicadidae" genus="Fidicinoides" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFCEE7344D3DEFCE0" box="[834,974,788,814]" italics="true" pageId="26" pageNumber="27">Fidicinoides</emphasis>
</taxonomicName>
by the presence of three-part tarsi.
<taxonomicName id="4C774D19FFE2FFCAFAC27345D18AFC9D" authority="Distant, 1905 d" authorityName="Distant" authorityYear="1905" class="Insecta" family="Cicadidae" genus="Majeorona" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFAC27345D0FDFC9C" italics="true" pageId="26" pageNumber="27">Majeorona</emphasis>
<bibRefCitation id="EFE64B6BFFE2FFCAFF597368D18AFC9D" author="Distant, W. L." box="[245,410,824,851]" pageId="26" pageNumber="27" pagination="304 - 319" refId="ref69306" refString="Distant, W. L. (1905 d) Rhynchotal notes XXX. Annals and Magazine of Natural History, Series 7, 15, 304 - 319. https: // doi. org / 10.1080 / 03745480509443047" type="journal article" year="1905">Distant, 1905d</bibRefCitation>
</taxonomicName>
has a head width wider than the mesonotum, with the prominent eyes stylate and extended anteriorly. The genus
<taxonomicName id="4C774D19FFE2FFCAFE27730DD1F9FCB8" authorityName="Stal" authorityYear="1864" box="[395,489,861,886]" class="Insecta" family="Cicadidae" genus="Proarna" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFE27730DD1F9FCB8" box="[395,489,861,886]" italics="true" pageId="26" pageNumber="27">Proarna</emphasis>
</taxonomicName>
possesses a head that is about as wide as the mesonotum with the eyes barely or not extending beyond the anterior angle of the pronotum, radial crossvein vertically oriented to the radius anterior 2 and radius posterior veins with a radiomedial crossvein that is obliquely oriented, large semicircular male opercula, and triangular or small, curved timbal covers.
<taxonomicName id="4C774D19FFE2FFCAFDC77398D2DBFC2C" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[619,715,968,994]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFDC77398D2DBFC2C" box="[619,715,968,994]" italics="true" pageId="26" pageNumber="27">Guyalna</emphasis>
</taxonomicName>
can be distinguished by the head being as wide or slightly wider than the mesonotum, eyes that only protrude slightly beyond the anterior angles of the pronotum, parallel sides of the abdomen, and the median uncus lobes are bent forming a horse-shoe shape when viewed from the posterior.
<taxonomicName id="4C774D19FFE2FFCAFF3B7465D119FB80" authorityName="Metcalf" authorityYear="1952" box="[151,265,1077,1102]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFF3B7465D119FB80" box="[151,265,1077,1102]" italics="true" pageId="26" pageNumber="27">Dorisiana</emphasis>
</taxonomicName>
can be distinguished by its head being as wide or wider than the mesonotum with the eyes not extending beyond the lateral edges of the pronotum, triangular timbal covers, ventral margin of the timbal cover angled dorsally not parallel to the long body axis, and the median uncus lobes are bent.
</paragraph>
<paragraph id="8BC8369AFFE2FFCAFF6B74F0D489FA85" blockId="26.[151,1437,788,1463]" pageId="26" pageNumber="27">
<taxonomicName id="4C774D19FFE2FFCAFF6B74F0D102FB74" authorityName="Distant" authorityYear="1905" box="[199,274,1184,1210]" class="Insecta" family="Cicadidae" genus="Ariasa" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFF6B74F0D102FB74" box="[199,274,1184,1210]" italics="true" pageId="26" pageNumber="27">Ariasa</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C774D19FFE2FFCAFE4974F0D356FB74" authority="Sanborn, 2016 c" authorityName="Sanborn" authorityYear="2016" box="[485,838,1184,1210]" class="Insecta" family="Cicadidae" genus="Cracenpsaltria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFE4974F0D29FFB74" box="[485,655,1184,1210]" italics="true" pageId="26" pageNumber="27">Cracenpsaltria</emphasis>
<bibRefCitation id="EFE64B6BFFE2FFCAFD3A74F0D356FB74" author="Sanborn, A. F." box="[662,838,1184,1210]" pageId="26" pageNumber="27" pagination="501 - 519" refId="ref73465" refString="Sanborn, A. F. (2016 c) Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa, 4137 (4), 501 - 519. https: // doi. org / 10.11646 / zootaxa. 4137.4.3" type="journal article" year="2016">Sanborn, 2016c</bibRefCitation>
</taxonomicName>
also represented in the Ecuadorian fauna. Species of
<taxonomicName id="4C774D19FFE2FFCAFF3B7494D0F2FB10" authorityName="Distant" authorityYear="1905" box="[151,226,1220,1246]" class="Insecta" family="Cicadidae" genus="Ariasa" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFF3B7494D0F2FB10" box="[151,226,1220,1246]" italics="true" pageId="26" pageNumber="27">Ariasa</emphasis>
</taxonomicName>
can be distinguished by the posteriorly extending median uncus lobe rather than the separate, long hooked lateral uncal lobes found in
<taxonomicName id="4C774D19FFE2FFCAFE7574B8D293FACC" authorityName="Sanborn" authorityYear="2016" box="[473,643,1256,1282]" class="Insecta" family="Cicadidae" genus="Cracenpsaltria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFE7574B8D293FACC" box="[473,643,1256,1282]" italics="true" pageId="26" pageNumber="27">Cracenpsaltria</emphasis>
</taxonomicName>
, the pygofer distal shoulder is angled in
<taxonomicName id="4C774D19FFE2FFCAFBF074B8D516FACC" authorityName="Sanborn" authorityYear="2016" box="[1116,1286,1256,1282]" class="Insecta" family="Cicadidae" genus="Cracenpsaltria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFBF074B8D516FACC" box="[1116,1286,1256,1282]" italics="true" pageId="26" pageNumber="27">Cracenpsaltria</emphasis>
</taxonomicName>
, the laterally expanding abdominal segments are absent in
<taxonomicName id="4C774D19FFE2FFCAFD3F755CD32DFAE8" authorityName="Sanborn" authorityYear="2016" box="[659,829,1292,1318]" class="Insecta" family="Cicadidae" genus="Cracenpsaltria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFD3F755CD32DFAE8" box="[659,829,1292,1318]" italics="true" pageId="26" pageNumber="27">Cracenpsaltria</emphasis>
</taxonomicName>
, and the length of the abdomen is greater than the distance between the apex of the head and the posterior cruciform elevation in
<taxonomicName id="4C774D19FFE2FFCAFC457560D483FA84" authorityName="Sanborn" authorityYear="2016" box="[1001,1171,1328,1354]" class="Insecta" family="Cicadidae" genus="Cracenpsaltria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFE2FFCAFC457560D483FA84" box="[1001,1171,1328,1354]" italics="true" pageId="26" pageNumber="27">Cracenpsaltria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6511FFE2FFCAFF6B7505D568FA78" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BC8369AFFE2FFCAFF6B7505D568FA78" blockId="26.[151,1437,788,1463]" pageId="26" pageNumber="27">
DISTRIBUTION.—Species of the genus have been recorded previously from
<collectingCountry id="F360760AFFE2FFCAFC557504D47BFAA0" box="[1017,1131,1364,1390]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFBD97504D4DAFAA0" box="[1141,1226,1364,1390]" name="Bolivia" pageId="26" pageNumber="27">Bolivia</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFB797504D50CFAA0" box="[1237,1308,1364,1390]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFA8B7504D588FAA0" box="[1319,1432,1364,1390]" name="Colombia" pageId="26" pageNumber="27">Colombia</collectingCountry>
, French Guiana,
<collectingCountry id="F360760AFFE2FFCAFEFC7528D1BAFA5C" box="[336,426,1400,1426]" name="Guyana" pageId="26" pageNumber="27">Guyana</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFE1B7529D230FA5D" box="[439,544,1401,1427]" name="Paraguay" pageId="26" pageNumber="27">Paraguay</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFD827529D274FA5D" box="[558,612,1401,1427]" name="Peru" pageId="26" pageNumber="27">Peru</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFDDE7528D2F0FA5C" box="[626,736,1400,1426]" name="Suriname" pageId="26" pageNumber="27">Suriname</collectingCountry>
,
<collectingCountry id="F360760AFFE2FFCAFD427529D341FA5D" box="[750,849,1401,1427]" name="Uruguay" pageId="26" pageNumber="27">Uruguay</collectingCountry>
, and
<collectingCountry id="F360760AFFE2FFCAFC3C7528D413FA5C" box="[912,1027,1400,1426]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
(
<bibRefCitation id="EFE64B6BFFE2FFCAFBB87528D4AEFA5C" author="Metcalf, Z. P." box="[1044,1214,1400,1426]" pageId="26" pageNumber="27" pagination="1 - 585" refId="ref71633" refString="Metcalf, Z. P. (1963 a) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i-vii + 1 - 585." type="journal article" year="1963">Metcalf 1963a</bibRefCitation>
; Sanborn 2011a, b; 2013; 2016b; 2016c; 2019b; 2020a, b;
<bibRefCitation id="EFE64B6BFFE2FFCAFDE675CCD342FA78" author="Sanborn, A. F. &amp; Heath, M. S." box="[586,850,1436,1462]" pageId="26" pageNumber="27" pagination="1 - 94" refId="ref74381" refString="Sanborn, A. F. &amp; Heath, M. S. (2014) The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 3883 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 3883.1.1" type="journal article" year="2014">Sanborn &amp; Heath 2014</bibRefCitation>
). The range is expanded here to include
<collectingCountry id="F360760AFFE2FFCAFAB975CCD564FA78" box="[1301,1396,1436,1462]" name="Ecuador" pageId="26" pageNumber="27">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>