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<mods:title id="D385E3D94A6CAC4BE6086A3015BB512D">An annotated and illustrated checklist of the porcelain crabs of Panama (Decapoda: Anomura)</mods:title>
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<taxonomicName id="4C7FE6FE7759FFFD4EF95B51FD9DABF1" ID-CoL="39935d99-618f-48a1-a1ae-eb3316772241" authority="Nobili, 1901" authorityName="Nobili" authorityYear="1901" box="[151,562,1384,1411]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="vicarius">
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<emphasis id="B90B416F7759FFFD4EF95B51FE3BABF1" bold="true" box="[151,404,1384,1411]" italics="true" pageId="58" pageNumber="59">Pachycheles vicarius</emphasis>
<bibRefCitation id="EFEEE08C7759FFFD4FF25B51FD9DABF1" author="Nobili, G." box="[412,562,1384,1410]" pageId="58" pageNumber="59" pagination="1 - 16" refId="ref75569" refString="Nobili, G. (1901) Decapodi raccolti dal Dr. Filippo Silvestri nell' America Meridionale. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Universita di Torino, 16 (402), 1 - 16." type="journal article" year="1901">Nobili, 1901</bibRefCitation>
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<paragraph id="8BC09D7D7759FFFD4EF95BB5FEC3ABD4" blockId="58.[151,562,1384,1447]" box="[151,364,1420,1447]" pageId="58" pageNumber="59">
(
<figureCitation id="134481F87759FFFD4EF15BB5FF52ABD4" box="[159,253,1420,1447]" captionStart="FIGURE 32" captionStartId="61.[151,250,1935,1959]" captionTargetBox="[189,1398,201,1912]" captionTargetId="figure-19@61.[189,1398,201,1912]" captionTargetPageId="61" captionText="FIGURE 32. Pachycheles vicarius Nobili, 1901: A, B, ovigerous female (cl 4.6 mm, cw 5.4 mm) from Punta Mala, Panama (MZUSP 33466), dorsal (A) and ventral (B) views [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532273" httpUri="https://zenodo.org/record/5532273/files/figure.png" pageId="58" pageNumber="59">Figs. 32</figureCitation>
,
<figureCitation id="134481F87759FFFD4F675BB4FE88ABD4" box="[265,295,1421,1447]" captionStart="FIGURE 33" captionStartId="62.[151,250,1811,1835]" captionTargetBox="[151,1435,181,1787]" captionTargetId="figure-19@62.[151,1435,181,1787]" captionTargetPageId="62" captionText="FIGURE 33. Pachycheles vicarius Nobili, 1901: A, male (cl 4.9 mm, cw 5.3 mm) from Coiba Island, Panama (FLMNH UF 57705), dorsal view; B, extra-limital specimen, male (cl 4.0 mm, cw 4.6 mm) from Los Negritos off Bahía Málaga, Colombia (CRBM-UV-cr:2009-106), dorsal view [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532275" httpUri="https://zenodo.org/record/5532275/files/figure.png" pageId="58" pageNumber="59">33</figureCitation>
,
<figureCitation id="134481F87759FFFD4F5D5BB4FECBABD4" box="[307,356,1421,1447]" captionStart="FIGURE 89" captionStartId="139.[151,250,1809,1833]" captionTargetBox="[189,1398,181,1785]" captionTargetId="figure-19@139.[189,1398,181,1785]" captionTargetPageId="139" captionText="FIGURE 89. Some eastern Pacific Porcellanidae (all species reported from Panama) photographed in situ: A, Pachycheles vicarius Nobili, 1901, Bahía Málaga, Colombia; B, C, Petrolisthes edwardsii (de Saussure, 1853), Los Cabos, Baja California, Mexico (B), and Malpelo Island, Colombia (C); D, Petrolisthes donadio Hiller &amp; Werding, 2007, Malpelo Island, Colombia, near sea urchin, Diadema mexicanum Agassiz; E, Petrolisthes glasselli Haig, 1957, Malpelo Island, Colombia; F, Petrolisthes haigae Chace, 1962, Malpelo Island, Colombia [photographs courtesy of Juan-Felipe Lazarus-Agudelo (A, C–F) and Jeff Goddard (B)]." figureDoi="http://doi.org/10.5281/zenodo.5532432" httpUri="https://zenodo.org/record/5532432/files/figure.png" pageId="58" pageNumber="59">89A</figureCitation>
)
</paragraph>
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<treatmentCitation id="0ADEBB6C7759FFFD4EF95BE8FDBAAB9A" author="Nobili, G." box="[151,533,1489,1513]" page="19" pageId="58" pageNumber="59" year="1901">
<taxonomicName id="4C7FE6FE7759FFFD4EF95BE8FDBAAB9A" ID-CoL="39935d99-618f-48a1-a1ae-eb3316772241" authority="Nobili 1901: 19" authorityName="Nobili" authorityPageNumber="19" authorityYear="1901" box="[151,533,1489,1513]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B90B416F7759FFFD4EF95BE8FEC0AB9A" box="[151,367,1489,1513]" italics="true" pageId="58" pageNumber="59">Pachycheles vicarius</emphasis>
<bibRefCitation id="EFEEE08C7759FFFD4F185BE8FDBAAB9A" author="Nobili, G." box="[374,533,1489,1513]" pageId="58" pageNumber="59" pagination="1 - 16" refId="ref75569" refString="Nobili, G. (1901) Decapodi raccolti dal Dr. Filippo Silvestri nell' America Meridionale. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Universita di Torino, 16 (402), 1 - 16." type="journal article" year="1901">Nobili 1901:19</bibRefCitation>
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;
<treatmentCitation id="0ADEBB6C7759FFFD4C4E5BE8FD6DAB9A" author="Haig, J." box="[544,706,1489,1513]" page="147" pageId="58" pageNumber="59" year="1960">
<bibRefCitation id="EFEEE08C7759FFFD4C4E5BE8FD6DAB9A" author="Haig, J." box="[544,706,1489,1513]" pageId="58" pageNumber="59" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960: 147</bibRefCitation>
</treatmentCitation>
, pl. 32, fig. 2;
<treatmentCitation id="0ADEBB6C7759FFFD4D325BE8FC5DAB9A" author="Haig, J." box="[860,1010,1489,1513]" page="69" pageId="58" pageNumber="59" year="1968">
<bibRefCitation id="EFEEE08C7759FFFD4D325BE8FC5DAB9A" author="Haig, J." box="[860,1010,1489,1513]" pageId="58" pageNumber="59" pagination="57 - 74" refId="ref72890" refString="Haig, J. (1968) Eastern Pacific expeditions of the New York Zoological Society. Porcellanid crabs (Crustacea: Anomura) from the west coast of tropical America. Zoologica, New York, 53 (2), 57 - 74." type="journal article" year="1968">Haig 1968: 69</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C7759FFFD4D905BE8FAAFAB9A" author="Gore, R. H. &amp; Abele, L. G." box="[1022,1280,1489,1513]" page="563" pageId="58" pageNumber="59" year="1974">
<bibRefCitation id="EFEEE08C7759FFFD4D905BE8FAAFAB9A" author="Gore, R. H. &amp; Abele, L. G." box="[1022,1280,1489,1513]" pageId="58" pageNumber="59" pagination="559 - 573" refId="ref72254" refString="Gore, R. H. &amp; Abele, L. G. (1974) Three new species of porcellanid crabs (Crustacea, Decapoda, Porcellanidae) from the Bay of Panama and adjacent Caribbean waters. Bulletin of Marine Science, 23 (3), 559 - 573." type="journal article" year="1974">Gore &amp; Abele 1974: 563</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C7759FFFD4B625BE8FE8CA87B" author="Gore, R. H. &amp; Abele, L. G." page="20" pageId="58" pageNumber="59" year="1976">
<bibRefCitation id="EFEEE08C7759FFFD4B625BE8FE8CA87B" author="Gore, R. H. &amp; Abele, L. G." pageId="58" pageNumber="59" pagination="1 - 30" refId="ref72307" refString="Gore, R. H. &amp; Abele, L. G. (1976) Shallow water porcelain crabs from the Pacific coast of Panama and adjacent Caribbean waters (Crustacea: Anomura: Porcellanidae). Smithsonian Contributions to Zoology, 237, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.237" type="journal article" year="1976">Gore &amp; Abele 1976: 20</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C7759FFFD4F435BC9FD97A874" author="Werding, B. &amp; Haig, H." box="[301,568,1519,1544]" page="65" pageId="58" pageNumber="59" year="1982">
<bibRefCitation id="EFEEE08C7759FFFD4F435BC9FD97A874" author="Werding, B. &amp; Haig, H." box="[301,568,1519,1544]" pageId="58" pageNumber="59" pagination="57 - 70" refId="ref77511" refString="Werding, B. &amp; Haig, H. (1982) The porcellanid crabs of the Isla Gorgona, Pacific coast of Colombia, with a description of Clastotoechus gorgonensis sp. nov. (Crustacea: Anomura). Anales del Instituto de Investigaciones Marinas de Punta Betin, 12, 57 - 70. https: // doi. org / 10.25268 / bimc. invemar. 1982.12.0.495" type="journal article" year="1982">Werding &amp; Haig 1982: 65</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C7759FFFD4C2D5BC9FBBDA87B" author="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R." box="[579,1042,1520,1544]" page="229" pageId="58" pageNumber="59" year="2007">
<bibRefCitation id="EFEEE08C7759FFFD4C2D5BC9FBBDA87B" author="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R." box="[579,1042,1520,1544]" pageId="58" pageNumber="59" pagination="221 - 239" refId="ref73986" refString="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R. (2007) Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico colombiano). Biota Colombiana, 8 (2), 221 - 239." type="journal article" year="2007">Lazarus-Agudelo &amp; Cantera-Kintz 2007: 229</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C7759FFFD4A725BC9FB75A87B" author="Moscoso, V." box="[1052,1242,1520,1544]" page="79" pageId="58" pageNumber="59" year="2012">
<bibRefCitation id="EFEEE08C7759FFFD4A725BC9FB75A87B" author="Moscoso, V." box="[1052,1242,1520,1544]" pageId="58" pageNumber="59" pagination="1 - 208" refId="ref75462" refString="Moscoso, V. (2012) Catalogo de crustaceos decapodos y estomatopodos del Peru. Boletin Instituto del Mar del Peru, 27 (1 / 2), 1 - 208. https: // doi. org / 10.5962 / bhl. part. 4563" type="journal article" year="2012">Moscoso 2012: 79</bibRefCitation>
</treatmentCitation>
, fig.
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</paragraph>
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<emphasis id="B90B416F7759FFFD4EF95809FED5A839" bold="true" box="[151,378,1584,1610]" pageId="58" pageNumber="59">Material examined</emphasis>
.
<materialsCitation id="3B1797207759FFFD4FE65808FB69A838" box="[392,1222,1584,1611]" collectionCode="FLMNH" collectorName="Pacific" country="Panama" location="Coiba Is" pageId="58" pageNumber="59" specimenCode="UF 57135" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F368DDED7759FFFD4FE65808FE4EA838" box="[392,481,1585,1611]" name="Panama" pageId="58" pageNumber="59">Panama</collectingCountry>
[
<collectorName id="268AF8AB7759FFFD4F815809FDE9A838" box="[495,582,1584,1611]" pageId="58" pageNumber="59">Pacific</collectorName>
]:
<specimenCount id="9D7956F47759FFFD4C365808FD13A839" box="[600,700,1584,1611]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
, cl 4.7, cw 5.7 (
<collectionCode id="ED6E05B87759FFFD4D015808FC7CA838" box="[879,979,1585,1611]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="58" pageNumber="59" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBD935067759FFFD4DB45808FBE0A838" box="[986,1103,1584,1611]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="58" pageNumber="59" type="Museum">UF 57135</specimenCode>
),
<location id="8EA0CBA67759FFFD4A0D5809FB69A838" LSID="urn:lsid:plazi:treatment:03D62C6B7759FFFC4EF95B51FA2FAF87:8EA0CBA67759FFFD4A0D5809FB69A838" box="[1123,1222,1584,1611]" country="Panama" name="Coiba Is" pageId="58" pageNumber="59">Coiba Is</location>
</materialsCitation>
.,
<materialsCitation id="3B1797207759FFFD4AB95809FF1FA8C4" collectingDate="2019-02-20" collectionCode="FLMNH" collectorName="M. Leray &amp; Coiba Is" country="Iceland" latitude="7.6315002" location="Coiba" municipality="Bahia Santa Cruz" pageId="58" pageNumber="59" specimenCode="UF 57705" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F368DDED7759FFFD4AB95809FB50A839" box="[1239,1279,1584,1610]" name="Iceland" pageId="58" pageNumber="59">Isla</collectingCountry>
<location id="8EA0CBA67759FFFD4B685809FAE3A839" LSID="urn:lsid:plazi:treatment:03D62C6B7759FFFC4EF95B51FA2FAF87:8EA0CBA67759FFFD4B685809FAE3A839" box="[1286,1356,1584,1610]" country="Iceland" latitude="7.6315002" municipality="Bahia Santa Cruz" name="Coiba" pageId="58" pageNumber="59">Coiba</location>
, northwest coast, small rocky island off
<collectingMunicipality id="6BA407077759FFFD4C77586DFD4DA81C" box="[537,738,1620,1647]" pageId="58" pageNumber="59">Bahía Santa Cruz</collectingMunicipality>
,
<geoCoordinate id="EE4BFBBA7759FFFD4C80586DFC2EA81C" box="[750,897,1620,1647]" degrees="7" direction="north" minutes="37" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="53.4" value="7.6315002">7°37’53.4”N</geoCoordinate>
, 81°46”44.9”W, rocky plateau, under rocks and in rock crevices, leg.
<collectorName id="268AF8AB7759FFFD4FEA5840FE44A8E0" box="[388,491,1657,1683]" pageId="58" pageNumber="59">M. Leray</collectorName>
<emphasis id="B90B416F7759FFFD4F9C5840FD8DA8E1" box="[498,546,1656,1682]" italics="true" pageId="58" pageNumber="59">et al</emphasis>
.,
<date id="FFC1BBBD7759FFFD4C585840FD1BA8E0" box="[566,692,1657,1683]" pageId="58" pageNumber="59" value="2019-02-20">
<collectingDate id="EF8542557759FFFD4C585840FD1BA8E0" box="[566,692,1657,1683]" pageId="58" pageNumber="59" value="2019-02-20">20.02.2019</collectingDate>
</date>
(fcn PAN-150);
<specimenCount id="9D7956F47759FFFD4D005840FC14A8E1" box="[878,955,1656,1683]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
, cl 4.9, cw 5.3 (
<collectionCode id="ED6E05B87759FFFD4A065840FB63A8E0" box="[1128,1228,1657,1683]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="58" pageNumber="59" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBD935067759FFFD4ABD5840FAE8A8E0" box="[1235,1351,1656,1683]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="58" pageNumber="59" type="Museum">UF 57705</specimenCode>
),
<collectorName id="268AF8AB7759FFFD4B375841FF1FA8C4" pageId="58" pageNumber="59">Coiba Is</collectorName>
</materialsCitation>
.,
<materialsCitation id="3B1797207759FFFD4EAF58A5FE26A951" collectingDate="2015-03-21" collectingDateMax="2019-02-22" collectingDateMin="2015-03-21" collectionCode="MZUSP" collectorName="M. Leray &amp; Punta Mala &amp; A. Anker &amp; J. F. Lazarus-Agudelo" country="Iceland" county="Extra-limital" latitude="7.6315002" location="Coiba" municipality="Bahia Santa Cruz" pageId="58" pageNumber="59" specimenCode="MZUSP 33466" specimenCount="1">
<collectingCountry id="F368DDED7759FFFD4EAF58A5FF46A8C5" box="[193,233,1692,1718]" name="Iceland" pageId="58" pageNumber="59">Isla</collectingCountry>
<location id="8EA0CBA67759FFFD4E9C58A5FE97A8C5" LSID="urn:lsid:plazi:treatment:03D62C6B7759FFFC4EF95B51FA2FAF87:8EA0CBA67759FFFD4E9C58A5FE97A8C5" box="[242,312,1692,1718]" country="Iceland" county="Extra-limital" latitude="7.6315002" municipality="Bahia Santa Cruz" name="Coiba" pageId="58" pageNumber="59">Coiba</location>
, northwest coast, small rocky island off
<collectingMunicipality id="6BA407077759FFFD4D6F58A5FC65A8C4" box="[769,970,1692,1719]" pageId="58" pageNumber="59">Bahía Santa Cruz</collectingMunicipality>
,
<geoCoordinate id="EE4BFBBA7759FFFD4DB858A5FBC6A8C4" box="[982,1129,1692,1719]" degrees="7" direction="north" minutes="37" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="53.4" value="7.6315002">7°37’53.4”N</geoCoordinate>
, 81°46”44.9”W, rocky plateau, under rocks and in rock crevices, leg.
<collectorName id="268AF8AB7759FFFD4C1058F8FD49A8A8" box="[638,742,1729,1755]" pageId="58" pageNumber="59">M. Leray</collectorName>
<emphasis id="B90B416F7759FFFD4C8358F8FCB1A8A9" box="[749,798,1728,1754]" italics="true" pageId="58" pageNumber="59">et al</emphasis>
.,
<date id="FFC1BBBD7759FFFD4D5D58F8FC1EA8A8" box="[819,945,1729,1755]" pageId="58" pageNumber="59" value="2019-02-22">
<collectingDate id="EF8542557759FFFD4D5D58F8FC1EA8A8" box="[819,945,1729,1755]" pageId="58" pageNumber="59" value="2019-02-22">22.02.2019</collectingDate>
</date>
(fcn PAN-224); 1 ov. female, cl 4.6, cw 5.4 (
<specimenCode id="DBD935067759FFFD4EF158DCFEFEA88C" box="[159,337,1764,1791]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="58" pageNumber="59">MZUSP 33466</specimenCode>
),
<collectorName id="268AF8AB7759FFFD4F0958DCFE43A88D" box="[359,492,1764,1791]" pageId="58" pageNumber="59">Punta Mala</collectorName>
, rocky intertidal, under rocks, leg.
<collectorName id="268AF8AB7759FFFD4DEC58DCFC44A88D" box="[898,1003,1764,1791]" pageId="58" pageNumber="59">A. Anker</collectorName>
&amp;
<collectorName id="268AF8AB7759FFFD4A7A58DCFAA6A88D" box="[1044,1289,1764,1791]" pageId="58" pageNumber="59">J.F. Lazarus-Agudelo</collectorName>
,
<date id="FFC1BBBD7759FFFD4B7958DDFA37A88C" box="[1303,1432,1764,1791]" pageId="58" pageNumber="59" value="2015-03-21">
<collectingDate id="EF8542557759FFFD4B7958DDFA37A88C" box="[1303,1432,1764,1791]" pageId="58" pageNumber="59" value="2015-03-21">21.03.2015</collectingDate>
</date>
.
<collectingCounty id="62A1E5F17759FFFD4EF95931FE89A951" box="[151,294,1800,1826]" pageId="58" pageNumber="59">Extra-limital</collectingCounty>
material
</materialsCitation>
.
<materialsCitation id="3B1797207759FFFD4FFB5931FBCFA918" collectingDate="2009-04-27" collectorName="A. Anker" country="Colombia" elevation="5" latitude="3.8965278" location="Los Negritos" longLatPrecision="1" longitude="-77.402725" municipality="Bahia Malaga" pageId="58" pageNumber="59" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F368DDED7759FFFD4FFB5931FDACA951" box="[405,515,1800,1826]" name="Colombia" pageId="58" pageNumber="59">Colombia</collectingCountry>
[Pacific]:
<specimenCount id="9D7956F47759FFFD4C175930FD69A951" box="[633,710,1800,1827]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
, cl 4.0, cw 4.6 (CRBM-UV-cr:2009-106),
<location id="8EA0CBA67759FFFD4AF05930FA81A951" LSID="urn:lsid:plazi:treatment:03D62C6B7759FFFC4EF95B51FA2FAF87:8EA0CBA67759FFFD4AF05930FA81A951" box="[1182,1326,1800,1827]" country="Colombia" latitude="3.8965278" longLatPrecision="1" longitude="-77.402725" municipality="Bahia Malaga" name="Los Negritos" pageId="58" pageNumber="59">Los Negritos</location>
off
<collectingMunicipality id="6BA407077759FFFD4B355931FF41A935" pageId="58" pageNumber="59">Bahía Málaga</collectingMunicipality>
,
<geoCoordinate id="EE4BFBBA7759FFFD4E955915FE33A934" box="[251,412,1836,1863]" degrees="03" direction="north" minutes="53" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="47.5" value="3.8965278">03°53’47.5”N</geoCoordinate>
,
<geoCoordinate id="EE4BFBBA7759FFFD4FC75914FDE1A934" box="[425,590,1837,1863]" degrees="77" direction="west" minutes="24" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="09.8" value="-77.402725">77°24’09.8”W</geoCoordinate>
, depth
<quantity id="4C8730987759FFFD4CCD5914FD41A934" box="[675,750,1837,1863]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="58" pageNumber="59" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="00527A4E7759FFFD4CCD5914FD41A934" box="[675,750,1837,1863]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="58" pageNumber="59" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</elevation>
</quantity>
, large rocky outcrop with large boulders, under large rocks, many cemented by sponges, on coarse sand, leg.
<collectorName id="268AF8AB7759FFFD4CD55968FC8BA919" box="[699,804,1872,1899]" pageId="58" pageNumber="59">A. Anker</collectorName>
,
<date id="FFC1BBBD7759FFFD4D415968FC03A918" box="[815,940,1873,1899]" pageId="58" pageNumber="59" value="2009-04-27">
<collectingDate id="EF8542557759FFFD4D415968FC03A918" box="[815,940,1873,1899]" pageId="58" pageNumber="59" value="2009-04-27">27.04.2009</collectingDate>
</date>
(fcn COL-254)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC09D7D7759FFFD4EA9594DFBCEA9FC" blockId="58.[151,1437,1584,2043]" box="[199,1121,1908,1935]" pageId="58" pageNumber="59">
<emphasis id="B90B416F7759FFFD4EA9594DFD95A9FC" bold="true" box="[199,570,1908,1935]" pageId="58" pageNumber="59">
Previous records from
<collectingCountry id="F368DDED7759FFFD4FB6594CFD95A9FC" box="[472,570,1909,1935]" name="Panama" pageId="58" pageNumber="59">Panama</collectingCountry>
</emphasis>
.
<bibRefCitation id="EFEEE08C7759FFFD4C26594DFD65A9FD" author="Haig, J." box="[584,714,1908,1934]" pageId="58" pageNumber="59" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960</bibRefCitation>
,
<bibRefCitation id="EFEEE08C7759FFFD4CBB594DFCBEA9FC" author="Haig, J." box="[725,785,1908,1935]" pageId="58" pageNumber="59" pagination="57 - 74" refId="ref72890" refString="Haig, J. (1968) Eastern Pacific expeditions of the New York Zoological Society. Porcellanid crabs (Crustacea: Anomura) from the west coast of tropical America. Zoologica, New York, 53 (2), 57 - 74." type="journal article" year="1968">1968</bibRefCitation>
);
<bibRefCitation id="EFEEE08C7759FFFD4D4B594CFBA2A9FC" author="Gore, R. H. &amp; Abele, L. G." box="[805,1037,1908,1935]" pageId="58" pageNumber="59" pagination="559 - 573" refId="ref72254" refString="Gore, R. H. &amp; Abele, L. G. (1974) Three new species of porcellanid crabs (Crustacea, Decapoda, Porcellanidae) from the Bay of Panama and adjacent Caribbean waters. Bulletin of Marine Science, 23 (3), 559 - 573." type="journal article" year="1974">Gore &amp; Abele (1974</bibRefCitation>
,
<bibRefCitation id="EFEEE08C7759FFFD4A76594DFBFBA9FC" author="Gore, R. H. &amp; Abele, L. G." box="[1048,1108,1908,1935]" pageId="58" pageNumber="59" pagination="1 - 30" refId="ref72307" refString="Gore, R. H. &amp; Abele, L. G. (1976) Shallow water porcelain crabs from the Pacific coast of Panama and adjacent Caribbean waters (Crustacea: Anomura: Porcellanidae). Smithsonian Contributions to Zoology, 237, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.237" type="journal article" year="1976">1976</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF67759FFFD4EA959A1FDD4A988" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BC09D7D7759FFFD4EA959A1FDD4A988" blockId="58.[151,1437,1584,2043]" pageId="58" pageNumber="59">
<emphasis id="B90B416F7759FFFD4EA959A1FEF6A9C1" bold="true" box="[199,345,1944,1970]" pageId="58" pageNumber="59">Distribution</emphasis>
. East Pacific:
<collectingCountry id="F368DDED7759FFFD4F9459A1FDD0A9C1" box="[506,639,1944,1970]" name="El Salvador" pageId="58" pageNumber="59">El Salvador</collectingCountry>
,
<collectingCountry id="F368DDED7759FFFD4CE459A0FCA9A9C1" box="[650,774,1944,1971]" name="Costa Rica" pageId="58" pageNumber="59">Costa Rica</collectingCountry>
,
<collectingCountry id="F368DDED7759FFFD4D7F59A0FCC5A9C0" box="[785,874,1945,1971]" name="Panama" pageId="58" pageNumber="59">Panama</collectingCountry>
(Coiba Is., Bahía Honda in
<collectingRegion id="49BB539F7759FFFD4ACF59A0FAA4A9C0" box="[1185,1291,1945,1971]" country="Panama" name="Veraguas" pageId="58" pageNumber="59">Veraguas</collectingRegion>
, Punta Mala, Las Perlas Is.)
<collectingCountry id="F368DDED7759FFFD4F535985FE01A9A5" box="[317,430,1980,2006]" name="Colombia" pageId="58" pageNumber="59">Colombia</collectingCountry>
,
<collectingCountry id="F368DDED7759FFFD4FD75985FDB9A9A5" box="[441,534,1980,2006]" name="Ecuador" pageId="58" pageNumber="59">Ecuador</collectingCountry>
and
<collectingCountry id="F368DDED7759FFFD4C225984FDD0A9A4" box="[588,639,1981,2007]" name="Peru" pageId="58" pageNumber="59">Peru</collectingCountry>
(
<bibRefCitation id="EFEEE08C7759FFFD4CE05985FCA7A9A5" author="Haig, J." box="[654,776,1980,2006]" pageId="58" pageNumber="59" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960</bibRefCitation>
,
<bibRefCitation id="EFEEE08C7759FFFD4D7D5985FCE1A9A5" author="Haig, J." box="[787,846,1980,2006]" pageId="58" pageNumber="59" pagination="57 - 74" refId="ref72890" refString="Haig, J. (1968) Eastern Pacific expeditions of the New York Zoological Society. Porcellanid crabs (Crustacea: Anomura) from the west coast of tropical America. Zoologica, New York, 53 (2), 57 - 74." type="journal article" year="1968">1968</bibRefCitation>
;
<bibRefCitation id="EFEEE08C7759FFFD4D375984FB97A9A5" author="Gore, R. H. &amp; Abele, L. G." box="[857,1080,1980,2007]" pageId="58" pageNumber="59" pagination="1 - 30" refId="ref72307" refString="Gore, R. H. &amp; Abele, L. G. (1976) Shallow water porcelain crabs from the Pacific coast of Panama and adjacent Caribbean waters (Crustacea: Anomura: Porcellanidae). Smithsonian Contributions to Zoology, 237, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.237" type="journal article" year="1976">Gore &amp; Abele 1976</bibRefCitation>
; Lazarus-Agudelo and Cantera- Kintz 2007;
<bibRefCitation id="EFEEE08C7759FFFD4F4D59D8FE64A988" author="Moscoso, V." box="[291,459,2017,2043]" pageId="58" pageNumber="59" pagination="1 - 208" refId="ref75462" refString="Moscoso, V. (2012) Catalogo de crustaceos decapodos y estomatopodos del Peru. Boletin Instituto del Mar del Peru, 27 (1 / 2), 1 - 208. https: // doi. org / 10.5962 / bhl. part. 4563" type="journal article" year="2012">Moscoso 2012</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF67758FFFC4EA95EAFFF4AAE8A" pageId="59" pageNumber="60" type="biology_ecology">
<paragraph id="8BC09D7D7758FFFC4EA95EAFFF4AAE8A" blockId="59.[151,1437,150,501]" pageId="59" pageNumber="60">
<emphasis id="B90B416F7758FFFC4EA95EAFFE89AEC3" bold="true" box="[199,294,150,176]" pageId="59" pageNumber="60">Ecology</emphasis>
. Intertidal and shallow subtidal, known depth range:
<quantity id="4C8730987758FFFC4DFE5EAFFC73AEC2" box="[912,988,150,177]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="59" pageNumber="60" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">0–8 m</quantity>
; typically under intertidal rocks and in crevices on rocky shores, but also in crevices of coral in deeper water (
<bibRefCitation id="EFEEE08C7758FFFC4DA35E83FBE5AEA7" author="Haig, J." box="[973,1098,186,212]" pageId="59" pageNumber="60" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960</bibRefCitation>
;
<bibRefCitation id="EFEEE08C7758FFFC4A365E83FA92AEA7" author="Gore, R. H. &amp; Abele, L. G." box="[1112,1341,186,212]" pageId="59" pageNumber="60" pagination="1 - 30" refId="ref72307" refString="Gore, R. H. &amp; Abele, L. G. (1976) Shallow water porcelain crabs from the Pacific coast of Panama and adjacent Caribbean waters (Crustacea: Anomura: Porcellanidae). Smithsonian Contributions to Zoology, 237, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.237" type="journal article" year="1976">Gore &amp; Abele 1976</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF67758FFFC4EA95F3BFA2FAF87" pageId="59" pageNumber="60" type="discussion">
<paragraph id="8BC09D7D7758FFFC4EA95F3BFA2FAF87" blockId="59.[151,1437,150,501]" pageId="59" pageNumber="60">
<emphasis id="B90B416F7758FFFC4EA95F3BFE9BAF6F" bold="true" box="[199,308,258,284]" pageId="59" pageNumber="60">Remarks</emphasis>
. The presumed western Atlantic sister of
<taxonomicName id="4C7FE6FE7758FFFC4D755F3AFC3BAF6F" authorityName="Nobili" authorityYear="1901" box="[795,916,259,284]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B90B416F7758FFFC4D755F3AFC3BAF6F" box="[795,916,259,284]" italics="true" pageId="59" pageNumber="60">P. vicarius</emphasis>
</taxonomicName>
is
<taxonomicName id="4C7FE6FE7758FFFC4DD55F3AFB9CAF6F" authorityName="Gore &amp; Abele" authorityYear="1974" box="[955,1075,259,284]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B90B416F7758FFFC4DD55F3AFB9CAF6F" box="[955,1075,259,284]" italics="true" pageId="59" pageNumber="60">P. susanae</emphasis>
</taxonomicName>
(see above). Indeed, these two species share numerous morphological characters, including the presence of strong longitudinal crests on the dorsal surface of the P1 carpus and palm; the shape and number of the mesial teeth on the P1 carpus; and the telson composed of seven plates. They differ, however, in several important morphological characters, as summarised by
<bibRefCitation id="EFEEE08C7758FFFC4EF95FABFE26AFDF" author="Gore, R. H. &amp; Abele, L. G." box="[151,393,402,428]" pageId="59" pageNumber="60" pagination="559 - 573" refId="ref72254" refString="Gore, R. H. &amp; Abele, L. G. (1974) Three new species of porcellanid crabs (Crustacea, Decapoda, Porcellanidae) from the Bay of Panama and adjacent Caribbean waters. Bulletin of Marine Science, 23 (3), 559 - 573." type="journal article" year="1974">Gore &amp; Abele (1974)</bibRefCitation>
, for instance, in the development of the male pleopods (present in
<taxonomicName id="4C7FE6FE7758FFFC4A195FAAFB42AFDF" authorityName="Nobili" authorityYear="1901" box="[1143,1261,403,428]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B90B416F7758FFFC4A195FAAFB42AFDF" box="[1143,1261,403,428]" italics="true" pageId="59" pageNumber="60">P. vicarius</emphasis>
</taxonomicName>
vs. absent in
<taxonomicName id="4C7FE6FE7758FFFC4BE65FAAFF5EAFA3" authorityName="Gore &amp; Abele" authorityYear="1974" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B90B416F7758FFFC4BE65FAAFF5EAFA3" italics="true" pageId="59" pageNumber="60">P. susanae</emphasis>
</taxonomicName>
) and the number of longitudinal crests on the P1 (four in
<taxonomicName id="4C7FE6FE7758FFFC4D045F8EFC71AFA3" authorityName="Nobili" authorityYear="1901" box="[874,990,439,464]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B90B416F7758FFFC4D045F8EFC71AFA3" box="[874,990,439,464]" italics="true" pageId="59" pageNumber="60">P. vicarius</emphasis>
</taxonomicName>
vs. three in
<taxonomicName id="4C7FE6FE7758FFFC4A0A5F8EFB77AFA3" authorityName="Gore &amp; Abele" authorityYear="1974" box="[1124,1240,439,464]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B90B416F7758FFFC4A0A5F8EFB77AFA3" box="[1124,1240,439,464]" italics="true" pageId="59" pageNumber="60">P. susanae</emphasis>
</taxonomicName>
). As expected for sister taxa, the colour pattern of
<taxonomicName id="4C7FE6FE7758FFFC4C6F5FE2FDD9AF87" authorityName="Nobili" authorityYear="1901" box="[513,630,475,500]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B90B416F7758FFFC4C6F5FE2FDD9AF87" box="[513,630,475,500]" italics="true" pageId="59" pageNumber="60">P. vicarius</emphasis>
</taxonomicName>
(
<figureCitation id="134481F87758FFFC4CE85FE3FD4CAF87" box="[646,739,474,500]" captionStart="FIGURE 32" captionStartId="61.[151,250,1935,1959]" captionTargetBox="[189,1398,201,1912]" captionTargetId="figure-19@61.[189,1398,201,1912]" captionTargetPageId="61" captionText="FIGURE 32. Pachycheles vicarius Nobili, 1901: A, B, ovigerous female (cl 4.6 mm, cw 5.4 mm) from Punta Mala, Panama (MZUSP 33466), dorsal (A) and ventral (B) views [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532273" httpUri="https://zenodo.org/record/5532273/files/figure.png" pageId="59" pageNumber="60">Figs. 32</figureCitation>
,
<figureCitation id="134481F87758FFFC4C815FE3FCA3AF87" box="[751,780,474,500]" captionStart="FIGURE 33" captionStartId="62.[151,250,1811,1835]" captionTargetBox="[151,1435,181,1787]" captionTargetId="figure-19@62.[151,1435,181,1787]" captionTargetPageId="62" captionText="FIGURE 33. Pachycheles vicarius Nobili, 1901: A, male (cl 4.9 mm, cw 5.3 mm) from Coiba Island, Panama (FLMNH UF 57705), dorsal view; B, extra-limital specimen, male (cl 4.0 mm, cw 4.6 mm) from Los Negritos off Bahía Málaga, Colombia (CRBM-UV-cr:2009-106), dorsal view [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532275" httpUri="https://zenodo.org/record/5532275/files/figure.png" pageId="59" pageNumber="60">33</figureCitation>
) is generally similar to that of
<taxonomicName id="4C7FE6FE7758FFFC4A0B5FE2FB74AF87" authorityName="Gore &amp; Abele" authorityYear="1974" box="[1125,1243,475,500]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B90B416F7758FFFC4A0B5FE2FB74AF87" box="[1125,1243,475,500]" italics="true" pageId="59" pageNumber="60">P. susanae</emphasis>
</taxonomicName>
(
<figureCitation id="134481F87758FFFC4A845FE3FAE8AF87" box="[1258,1351,474,500]" captionStart="FIGURE 28" captionStartId="56.[151,250,1944,1968]" captionTargetBox="[179,1407,181,1920]" captionTargetId="figure-19@56.[179,1407,181,1920]" captionTargetPageId="56" captionText="FIGURE 28. Pachycheles susanae Gore &amp; Abele, 1974: A, B, male (cl 4.4 mm, cw 4.8 mm) from Bocas del Toro, Panama (FLMNH UF 57133), dorsal (A) and ventral (B) view [photographs courtesy of Paulo P.G Pachelle]." figureDoi="http://doi.org/10.5281/zenodo.5532263" httpUri="https://zenodo.org/record/5532263/files/figure.png" pageId="59" pageNumber="60">Figs. 28</figureCitation>
,
<figureCitation id="134481F87758FFFC4B3D5FE3FADCAF87" box="[1363,1395,474,500]" captionStart="FIGURE 29" captionStartId="57.[151,250,1089,1113]" captionTargetBox="[151,1435,181,1065]" captionTargetId="figure-20@57.[151,1435,181,1065]" captionTargetPageId="57" captionText="FIGURE 29. Pachycheles susanae Gore &amp; Abele, 1974: ovigerous female (cl 4.8 mm, cw 5.6 mm) from Isla Grande, Panama (MZUSP 33470), dorsal view [photograph by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532265" httpUri="https://zenodo.org/record/5532265/files/figure.png" pageId="59" pageNumber="60">29</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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