treatments-xml/data/03/D5/87/03D58791FFC4FF91A1B285BB3ABDBDD3.xml
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<document id="5601E7E43D790B10F27C1E5059C6D23D" ID-DOI="10.11646/zootaxa.5315.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142338" ID-ZooBank="9034FE10-0022-4009-BC46-11EE68A9B59F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689236579947" checkinUser="plazi" docAuthor="Antić, Dragan" docDate="2023" docId="03D58791FFC4FF91A1B285BB3ABDBDD3" docLanguage="en" docName="zootaxa.5315.5.2.pdf" docOrigin="Zootaxa 5315 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Karadenizia merti Antić 2023, gen. nov., sp. nov." docType="treatment" docVersion="2" lastPageNumber="465" masterDocId="FFECFFE9FFC5FF98A12580523840BD06" masterDocTitle="Karadenizia, a new monospecific pachyiuline genus (Diplopoda: Julida: Julidae) from Turkey" masterLastPageNumber="468" masterPageNumber="456" pageNumber="457" updateTime="1689236979077" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C379FF02BD6727572015E1658788F7AB">Karadenizia, a new monospecific pachyiuline genus (Diplopoda: Julida: Julidae) from Turkey</mods:title>
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<mods:namePart id="3A09733B972BE01DF6DCB9B4082C4036">Antić, Dragan</mods:namePart>
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<mods:date id="9859BAF1E18F8DCC19EC5E11B56D19BB">2023</mods:date>
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<treatment id="03D58791FFC4FF91A1B285BB3ABDBDD3" LSID="urn:lsid:plazi:treatment:03D58791FFC4FF91A1B285BB3ABDBDD3" httpUri="http://treatment.plazi.org/id/03D58791FFC4FF91A1B285BB3ABDBDD3" lastPageId="9" lastPageNumber="465" pageId="1" pageNumber="457">
<subSubSection id="C366650CFFC4FF99A1B285BB3A12BB05" box="[151,594,1513,1540]" pageId="1" pageNumber="457" type="nomenclature">
<paragraph id="8BC33687FFC4FF99A1B285BB3A12BB05" blockId="1.[151,594,1513,1576]" box="[151,594,1513,1540]" pageId="1" pageNumber="457">
<heading id="D08B81EBFFC4FF99A1B285BB3A12BB05" bold="true" box="[151,594,1513,1540]" fontSize="11" level="1" pageId="1" pageNumber="457" reason="1">
<emphasis id="B908EA95FFC4FF99A1B285BB3A12BB05" bold="true" box="[151,594,1513,1540]" pageId="1" pageNumber="457">
<taxonomicName id="4C7C4D04FFC4FF99A1B285BB3931BB05" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[151,369,1513,1540]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="1" pageNumber="457" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC4FF99A1B285BB3931BB05" bold="true" box="[151,369,1513,1540]" italics="true" pageId="1" pageNumber="457">Karadenizia merti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57EEFFC4FF99A05D85BB3A12BB05" box="[376,594,1513,1539]" pageId="1" pageNumber="457" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFC4FF99A1B2865F38B8BB2E" box="[151,248,1549,1576]" pageId="1" pageNumber="457" type="description">
<paragraph id="8BC33687FFC4FF99A1B2865F38B8BB2E" blockId="1.[151,594,1513,1576]" box="[151,248,1549,1576]" pageId="1" pageNumber="457">
<figureCitation id="13472A02FFC4FF99A1B2865F38B8BB2E" box="[151,248,1549,1576]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1870,1894]" captionStartId-1="3.[151,250,1874,1898]" captionStartId-2="4.[151,250,1943,1967]" captionStartId-3="5.[151,250,1909,1933]" captionStartId-4="6.[151,250,1907,1931]" captionTargetBox-0="[151,1436,179,1844]" captionTargetBox-1="[226,1360,181,1851]" captionTargetBox-2="[185,1401,181,1919]" captionTargetBox-3="[231,1357,181,1885]" captionTargetBox-4="[169,1423,189,1877]" captionTargetId-0="figure-15@2.[151,1436,179,1844]" captionTargetId-1="figure-19@3.[226,1360,181,1851]" captionTargetId-2="figure-15@4.[185,1401,181,1919]" captionTargetId-3="figure-19@5.[231,1357,181,1885]" captionTargetId-4="figure-15@6.[158,1430,181,1883]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." captionText-1="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." captionText-2="FIGURE 3. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), left promere (anterior gonopod). A. Posterior view. B. Lateral view. C. Anterolateral view. D. Anterior view. E. Anteromesal view. F. Mesal view. Abbreviations: lr: lateral ridge; t: triangular tooth. Scale bar: 0.1 mm." captionText-3="FIGURE 4. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), left opisthomere (posterior gonopod). A. Posteromesal view. B. Mesal view. C. Anterior view. D. Anterolateral view. E. Lateral view. F. Posterolateral view. G. Distal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; m: mesomeral process; o: opisthomere; s: solenomere; sg: sperm groove; v: velum. Scale bar: 0.1 mm." captionText-4="FIGURE 5. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), right gonopods. A. Right gonopods, lateral view. B. Right gonopods, mesal view. C. Right promere, posterior view. D. Right gonopods, anterior view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8142342" figureDoi-1="http://doi.org/10.5281/zenodo.8142344" figureDoi-2="http://doi.org/10.5281/zenodo.8142346" figureDoi-3="http://doi.org/10.5281/zenodo.8142348" figureDoi-4="http://doi.org/10.5281/zenodo.8142350" httpUri-0="https://zenodo.org/record/8142342/files/figure.png" httpUri-1="https://zenodo.org/record/8142344/files/figure.png" httpUri-2="https://zenodo.org/record/8142346/files/figure.png" httpUri-3="https://zenodo.org/record/8142348/files/figure.png" httpUri-4="https://zenodo.org/record/8142350/files/figure.png" pageId="1" pageNumber="457">Figs 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFC4FF99A1B286073B1ABBDA" pageId="1" pageNumber="457" type="diagnosis">
<paragraph id="8BC33687FFC4FF99A1B28607395EBB95" blockId="1.[151,1437,1621,2008]" pageId="1" pageNumber="457">
<emphasis id="B908EA95FFC4FF99A1B286073952BB69" bold="true" box="[151,274,1621,1647]" pageId="1" pageNumber="457">Diagnosis.</emphasis>
As for the genus, with the addition that the metazonal setae are very long, ca 45% of the vertical diameter of the rings.
</paragraph>
<paragraph id="8BC33687FFC4FF99A1E286CF3B1ABBDA" blockId="1.[151,1437,1621,2008]" pageId="1" pageNumber="457">
<emphasis id="B908EA95FFC4FF99A1E286CF3954BBB1" bold="true" box="[199,276,1693,1719]" pageId="1" pageNumber="457">Name.</emphasis>
The new species is dedicated to Mert Elverici, a Turkish biospeleologist and the collector of this beautiful animal. The epithet, a name in the genitive case, is a patronym.
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFC4FF99A1E286B73B97BA6A" pageId="1" pageNumber="457" type="materials_examined">
<paragraph id="8BC33687FFC4FF99A1E286B739F1BBF9" blockId="1.[151,1437,1621,2008]" box="[199,433,1765,1791]" pageId="1" pageNumber="457">
<emphasis id="B908EA95FFC4FF99A1E286B739F1BBF9" bold="true" box="[199,433,1765,1791]" pageId="1" pageNumber="457">Material examined.</emphasis>
</paragraph>
<paragraph id="8BC33687FFC4FF99A1E2875B3A0DBA4E" blockId="1.[151,1437,1621,2008]" pageId="1" pageNumber="457">
<materialsCitation id="3B143CDAFFC4FF99A1E2875B3A0DBA4E" collectingDate="2020-08-12" collectionCode="ABBM" collectorName="M. Elverici" country="Turkey" elevation="260" latitude="41.4358" location="Ercek cave" longLatPrecision="6" longitude="31.8631" municipality="Ercek village" pageId="1" pageNumber="457" specimenCount="1" stateProvince="Zonguldak" typeStatus="holotype">
<emphasis id="B908EA95FFC4FF99A1E2875B397DBA25" bold="true" box="[199,317,1801,1827]" pageId="1" pageNumber="457">
<typeStatus id="54C78825FFC4FF99A1E2875B3978BA25" box="[199,312,1801,1827]" pageId="1" pageNumber="457">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F36B7617FFC4FF99A063875839F7BA22" box="[326,439,1802,1828]" name="Turkey" pageId="1" pageNumber="457">TURKEY</collectingCountry>
● ♁;
<collectingRegion id="49B8F865FFC4FF99A0D9875B3AA4BA25" box="[508,740,1801,1827]" country="Turkey" name="Zonguldak" pageId="1" pageNumber="457">Zonguldak Province</collectingRegion>
,
<collectingMunicipality id="6BA7ACFDFFC4FF99A3D5875B3BC9BA25" box="[752,905,1801,1827]" pageId="1" pageNumber="457">Erçek village</collectingMunicipality>
,
<location id="8EA3605CFFC4FF99A2B0875B3C52BA22" LSID="urn:lsid:plazi:treatment:03D58791FFC4FF91A1B285BB3ABDBDD3:8EA3605CFFC4FF99A2B0875B3C52BA22" box="[917,1042,1801,1828]" country="Turkey" latitude="41.4358" longLatPrecision="6" longitude="31.8631" municipality="Ercek village" name="Ercek cave" pageId="1" pageNumber="457" stateProvince="Zonguldak">Erçek cave</location>
(
<geoCoordinate id="EE485040FFC4FF99A507875B3CC1BA22" box="[1058,1153,1801,1828]" degrees="41.4358" direction="north" orientation="latitude" pageId="1" pageNumber="457" precision="5" value="41.4358">41.4358</geoCoordinate>
,
<geoCoordinate id="EE485040FFC4FF99A5AB875B3CADBA25" box="[1166,1261,1801,1827]" degrees="31.8631" direction="east" orientation="longitude" pageId="1" pageNumber="457" precision="5" value="31.8631">31.8631</geoCoordinate>
),
<elevation id="0051D1B4FFC4FF99A424875B3D33BA25" box="[1281,1395,1801,1828]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="457" unit="m" value="260.0">
<quantity id="4C849B62FFC4FF99A424875B3D09BA22" box="[1281,1353,1801,1828]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="457" unit="m" value="260.0">260 m</quantity>
asl
</elevation>
;
<date id="FFC21047FFC4FF99A4A5875B396BBA41" pageId="1" pageNumber="457" value="2020-08-12">
<collectingDate id="EF86E9AFFFC4FF99A4A5875B396BBA41" pageId="1" pageNumber="457" value="2020-08-12">12 August 2020</collectingDate>
</date>
;
<collectorName id="26895351FFC4FF99A013877C39F6BA41" box="[310,438,1837,1864]" pageId="1" pageNumber="457">M. Elverici</collectorName>
leg.;
<collectionCode id="ED6DAE42FFC4FF99A0D6877C3A09BA4E" box="[499,585,1838,1864]" pageId="1" pageNumber="457">ABBM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC33687FFC4FF99A1E287033B97BA6A" blockId="1.[151,1437,1621,2008]" box="[199,983,1873,1900]" pageId="1" pageNumber="457">
<materialsCitation id="3B143CDAFFC4FF99A1E287033B93BA6A" box="[199,979,1873,1900]" collectingDate="2020-08-12" collectionCode="ABBM" collectorName="M. Elverici" country="Turkey" elevation="260" latitude="41.4358" location="Ercek cave" longLatPrecision="6" longitude="31.8631" municipality="Ercek village" pageId="1" pageNumber="457" specimenCount="1" specimenCount-juvenile="1" stateProvince="Zonguldak" typeStatus="paratype">
<emphasis id="B908EA95FFC4FF99A1E28703397DBA6D" bold="true" box="[199,317,1873,1899]" pageId="1" pageNumber="457">
<typeStatus id="54C78825FFC4FF99A1E287033978BA6D" box="[199,312,1873,1899]" pageId="1" pageNumber="457">Paratype</typeStatus>
:
</emphasis>
<collectingCountry id="F36B7617FFC4FF99A061870039F5BA6A" box="[324,437,1874,1900]" name="Turkey" pageId="1" pageNumber="457">TURKEY</collectingCountry>
<specimenCount id="9D7AFD0EFFC4FF99A0F187033A07BA6D" box="[468,583,1873,1899]" pageId="1" pageNumber="457" type="juvenile">1 juvenile</specimenCount>
; same data as for holotype;
<collectionCode id="ED6DAE42FFC4FF99A25887003B93BA6A" box="[893,979,1874,1900]" pageId="1" pageNumber="457">ABBM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFC4FF9FA1E287273D1DB89B" lastPageId="7" lastPageNumber="463" pageId="1" pageNumber="457" type="description">
<paragraph id="8BC33687FFC4FF99A1E287273A7FBA89" blockId="1.[151,1437,1621,2008]" box="[199,575,1909,1936]" pageId="1" pageNumber="457">
<emphasis id="B908EA95FFC4FF99A1E287273911BA89" bold="true" box="[199,337,1909,1935]" pageId="1" pageNumber="457">Description</emphasis>
(based on
<typeStatus id="54C78825FFC4FF99A0E987273A73BA89" box="[460,563,1909,1935]" pageId="1" pageNumber="457">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BC33687FFC4FF99A1E287C83AB9BAD1" blockId="1.[151,1437,1621,2008]" pageId="1" pageNumber="457">
<emphasis id="B908EA95FFC4FF99A1E287C83A62BAB5" box="[199,546,1945,1972]" italics="true" pageId="1" pageNumber="457">Size and number of body rings.</emphasis>
Body moniliform,
<quantity id="4C849B62FFC4FF99A3DC87CB3B08BAB2" box="[761,840,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="457" unit="mm" value="12.0">12 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4C849B62FFC4FF99A41787CB3DD8BAB2" box="[1330,1432,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="457" unit="mm" value="1.1">1.10 mm</quantity>
, body with 34 podous rings + 3 apodous rings + telson.
</paragraph>
<caption id="DF03660FFFC7FF9AA1B2871C3A34BAD4" ID-DOI="http://doi.org/10.5281/zenodo.8142342" ID-Zenodo-Dep="8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="2" pageNumber="458" startId="2.[151,250,1870,1894]" targetBox="[151,1436,179,1844]" targetPageId="2" targetType="figure">
<paragraph id="8BC33687FFC7FF9AA1B2871C3A34BAD4" blockId="2.[151,1437,1870,2002]" pageId="2" pageNumber="458">
<emphasis id="B908EA95FFC7FF9AA1B2871C3954BA60" bold="true" box="[151,276,1870,1894]" pageId="2" pageNumber="458">FIGURE 1.</emphasis>
<taxonomicName id="4C7C4D04FFC7FF9AA03E871C3994BA60" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[283,468,1870,1894]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="2" pageNumber="458" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC7FF9AA03E871C3994BA60" box="[283,468,1870,1894]" italics="true" pageId="2" pageNumber="458">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC7FF9AA0FE871C3ADABA60" bold="true" box="[475,666,1870,1894]" pageId="2" pageNumber="458">
<taxonomicNameLabel id="A23B57EEFFC7FF9AA0FE871C3ADABA60" box="[475,666,1870,1894]" pageId="2" pageNumber="458" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
, ♁ holotype (ABBM), habitus.
<emphasis id="B908EA95FFC7FF9AA2C5871C3BB2BA60" bold="true" box="[992,1010,1870,1894]" pageId="2" pageNumber="458">A</emphasis>
. Whole body, lateral view.
<emphasis id="B908EA95FFC7FF9AA42E871C3D5CBA60" bold="true" box="[1291,1308,1870,1894]" pageId="2" pageNumber="458">B</emphasis>
. Head, right side, lateral, slightly ventral view.
<emphasis id="B908EA95FFC7FF9AA0DF87203A4CBA8C" bold="true" box="[506,524,1906,1930]" pageId="2" pageNumber="458">C</emphasis>
. Anterior part of body, lateral view.
<emphasis id="B908EA95FFC7FF9AA2A087203BD7BA8C" bold="true" box="[901,919,1906,1930]" pageId="2" pageNumber="458">D</emphasis>
. Head and rings 13, left side, lateral view, arrow shows mandibular stipes.
<emphasis id="B908EA95FFC7FF9AA0BB87C439EFBAA8" bold="true" box="[414,431,1942,1966]" pageId="2" pageNumber="458">E</emphasis>
. Midbody rings, lateral view.
<emphasis id="B908EA95FFC7FF9AA3C487C43AB0BAA8" bold="true" box="[737,752,1942,1966]" pageId="2" pageNumber="458">F</emphasis>
. Right half of pleurotergum 7, lateral view.
<emphasis id="B908EA95FFC7FF9AA59587C43C84BAA8" bold="true" box="[1200,1220,1942,1966]" pageId="2" pageNumber="458">G</emphasis>
. Telson, lateral view.
<emphasis id="B908EA95FFC7FF9AA1B287E838EBBAD4" bold="true" box="[151,171,1978,2002]" pageId="2" pageNumber="458">H</emphasis>
. Telson, posterior view.
<emphasis id="B908EA95FFC7FF9AA08187E83A52BAD4" bold="true" box="[420,530,1978,2002]" pageId="2" pageNumber="458">Scale bars</emphasis>
: 0.5 mm.
</paragraph>
</caption>
<caption id="DF03660FFFC6FF9BA1B287003928BAFC" ID-DOI="http://doi.org/10.5281/zenodo.8142344" ID-Zenodo-Dep="8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="3" pageNumber="459" startId="3.[151,250,1874,1898]" targetBox="[226,1360,181,1851]" targetPageId="3" targetType="figure">
<paragraph id="8BC33687FFC6FF9BA1B287003928BAFC" blockId="3.[151,1437,1874,2043]" pageId="3" pageNumber="459">
<emphasis id="B908EA95FFC6FF9BA1B287003955BA6C" bold="true" box="[151,277,1874,1898]" pageId="3" pageNumber="459">FIGURE 2.</emphasis>
<taxonomicName id="4C7C4D04FFC6FF9BA03887013998BA6C" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[285,472,1875,1899]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="3" pageNumber="459" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC6FF9BA03887013998BA6C" box="[285,472,1875,1899]" italics="true" pageId="3" pageNumber="459">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC6FF9BA0C487013AE3BA6D" bold="true" box="[481,675,1875,1899]" pageId="3" pageNumber="459">
<taxonomicNameLabel id="A23B57EEFFC6FF9BA0C487013AE3BA6D" box="[481,675,1875,1899]" pageId="3" pageNumber="459" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
, ♁ holotype (ABBM).
<emphasis id="B908EA95FFC6FF9BA2BC87003BEBBA6C" bold="true" box="[921,939,1874,1898]" pageId="3" pageNumber="459">A</emphasis>
. Gnathochilarium, ventral view.
<emphasis id="B908EA95FFC6FF9BA42787013D53BA6D" bold="true" box="[1282,1299,1875,1899]" pageId="3" pageNumber="459">B</emphasis>
. Right leg 1, lateral view.
<emphasis id="B908EA95FFC6FF9BA03E8724396DBA88" bold="true" box="[283,301,1910,1934]" pageId="3" pageNumber="459">C</emphasis>
. Distal part of right leg 2, posterior view.
<emphasis id="B908EA95FFC6FF9BA3C687253AB5BA89" bold="true" box="[739,757,1911,1935]" pageId="3" pageNumber="459">D</emphasis>
. Penis, posterior view.
<emphasis id="B908EA95FFC6FF9BA2C287253BB8BA89" bold="true" box="[999,1016,1911,1935]" pageId="3" pageNumber="459">E</emphasis>
. Right promere, posterior view.
<emphasis id="B908EA95FFC6FF9BA46287253D16BA89" bold="true" box="[1351,1366,1911,1935]" pageId="3" pageNumber="459">F</emphasis>
. Right gonopods, lateral view.
<emphasis id="B908EA95FFC6FF9BA0AB87C839E2BAB4" bold="true" box="[398,418,1946,1970]" pageId="3" pageNumber="459">G</emphasis>
. Right gonopods, mesal view.
<emphasis id="B908EA95FFC6FF9BA3C087C83B3EBAB4" bold="true" box="[741,894,1946,1970]" pageId="3" pageNumber="459">Abbreviations</emphasis>
:
<emphasis id="B908EA95FFC6FF9BA2A887C93BEFBAB5" bold="true" box="[909,943,1947,1971]" pageId="3" pageNumber="459">am</emphasis>
: accessory membrane;
<emphasis id="B908EA95FFC6FF9BA58687C93CEABAB5" bold="true" box="[1187,1194,1947,1971]" pageId="3" pageNumber="459">l</emphasis>
: subtriangular lobe;
<emphasis id="B908EA95FFC6FF9BA4A687C93DD5BAB5" bold="true" box="[1411,1429,1947,1971]" pageId="3" pageNumber="459">lr</emphasis>
: lateral ridge;
<emphasis id="B908EA95FFC6FF9BA03887ED3972BAD1" bold="true" box="[285,306,1983,2007]" pageId="3" pageNumber="459">m</emphasis>
: mesomeral process;
<emphasis id="B908EA95FFC6FF9BA32887ED3A5ABAD1" bold="true" box="[525,538,1983,2007]" pageId="3" pageNumber="459">o</emphasis>
: opisthomere;
<emphasis id="B908EA95FFC6FF9BA39587ED3AFEBAD1" bold="true" box="[688,702,1983,2007]" pageId="3" pageNumber="459">p</emphasis>
: promere;
<emphasis id="B908EA95FFC6FF9BA20987ED3B76BAD1" bold="true" box="[812,822,1983,2007]" pageId="3" pageNumber="459">s</emphasis>
: solenomere;
<emphasis id="B908EA95FFC6FF9BA2E087ED3B9CBAD1" bold="true" box="[965,988,1983,2007]" pageId="3" pageNumber="459">sg</emphasis>
: sperm groove;
<emphasis id="B908EA95FFC6FF9BA55A87ED3CC7BAD1" bold="true" box="[1151,1159,1983,2007]" pageId="3" pageNumber="459">t</emphasis>
: triangular tooth;
<emphasis id="B908EA95FFC6FF9BA41887ED3D0ABAD1" bold="true" box="[1341,1354,1983,2007]" pageId="3" pageNumber="459">v</emphasis>
: velum.
<emphasis id="B908EA95FFC6FF9BA1B287B03946BAFD" bold="true" box="[151,262,2018,2043]" pageId="3" pageNumber="459">Scale bars</emphasis>
: 0.1 mm.
</paragraph>
</caption>
<caption id="DF03660FFFC1FF9CA1B287C53A5DBAF0" ID-DOI="http://doi.org/10.5281/zenodo.8142346" ID-Zenodo-Dep="8142346" httpUri="https://zenodo.org/record/8142346/files/figure.png" pageId="4" pageNumber="460" startId="4.[151,250,1943,1967]" targetBox="[185,1401,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="8BC33687FFC1FF9CA1B287C53A5DBAF0" blockId="4.[151,1437,1943,2039]" pageId="4" pageNumber="460">
<emphasis id="B908EA95FFC1FF9CA1B287C53954BAA9" bold="true" box="[151,276,1943,1967]" pageId="4" pageNumber="460">FIGURE 3.</emphasis>
<taxonomicName id="4C7C4D04FFC1FF9CA03987C53996BAA9" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[284,470,1943,1967]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="4" pageNumber="460" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC1FF9CA03987C53996BAA9" box="[284,470,1943,1967]" italics="true" pageId="4" pageNumber="460">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC1FF9CA0FB87C53ADEBAA9" bold="true" box="[478,670,1943,1967]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A23B57EEFFC1FF9CA0FB87C53ADEBAA9" box="[478,670,1943,1967]" pageId="4" pageNumber="460" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
, ♁ holotype (ABBM), left promere (anterior gonopod).
<emphasis id="B908EA95FFC1FF9CA5C687C53CB5BAA9" bold="true" box="[1251,1269,1943,1967]" pageId="4" pageNumber="460">A</emphasis>
. Posterior view.
<emphasis id="B908EA95FFC1FF9CA1B287E938E8BAD5" bold="true" box="[151,168,1979,2003]" pageId="4" pageNumber="460">B</emphasis>
. Lateral view.
<emphasis id="B908EA95FFC1FF9CA01A87E93911BAD5" bold="true" box="[319,337,1979,2003]" pageId="4" pageNumber="460">C</emphasis>
. Anterolateral view.
<emphasis id="B908EA95FFC1FF9CA30187E93A76BAD5" bold="true" box="[548,566,1979,2003]" pageId="4" pageNumber="460">D</emphasis>
. Anterior view.
<emphasis id="B908EA95FFC1FF9CA3FF87E93AABBAD5" bold="true" box="[730,747,1979,2003]" pageId="4" pageNumber="460">E</emphasis>
. Anteromesal view.
<emphasis id="B908EA95FFC1FF9CA29F87E93B89BAD5" bold="true" box="[954,969,1979,2003]" pageId="4" pageNumber="460">F</emphasis>
. Mesal view.
<emphasis id="B908EA95FFC1FF9CA57387E93CAFBAD5" bold="true" box="[1110,1263,1979,2003]" pageId="4" pageNumber="460">Abbreviations</emphasis>
:
<emphasis id="B908EA95FFC1FF9CA5D987E93D4EBAD5" bold="true" box="[1276,1294,1979,2003]" pageId="4" pageNumber="460">lr</emphasis>
: lateral ridge;
<emphasis id="B908EA95FFC1FF9CA1B2878D38DFBAF1" bold="true" box="[151,159,2015,2039]" pageId="4" pageNumber="460">t</emphasis>
: triangular tooth.
<emphasis id="B908EA95FFC1FF9CA072878D39FBBAF1" bold="true" box="[343,443,2015,2039]" pageId="4" pageNumber="460">Scale bar</emphasis>
: 0.1 mm.
</paragraph>
</caption>
<caption id="DF03660FFFC0FF9DA1B287273A1CBAFF" ID-DOI="http://doi.org/10.5281/zenodo.8142348" ID-Zenodo-Dep="8142348" httpUri="https://zenodo.org/record/8142348/files/figure.png" pageId="5" pageNumber="461" startId="5.[151,250,1909,1933]" targetBox="[231,1357,181,1885]" targetPageId="5" targetType="figure">
<paragraph id="8BC33687FFC0FF9DA1B287273A1CBAFF" blockId="5.[151,1437,1909,2042]" pageId="5" pageNumber="461">
<emphasis id="B908EA95FFC0FF9DA1B287273952BA8B" bold="true" box="[151,274,1909,1933]" pageId="5" pageNumber="461">FIGURE 4.</emphasis>
<taxonomicName id="4C7C4D04FFC0FF9DA03D8727398FBA8B" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[280,463,1909,1933]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="5" pageNumber="461" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC0FF9DA03D8727398FBA8B" box="[280,463,1909,1933]" italics="true" pageId="5" pageNumber="461">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC0FF9DA0F087273ACFBA8B" bold="true" box="[469,655,1909,1933]" pageId="5" pageNumber="461">
<taxonomicNameLabel id="A23B57EEFFC0FF9DA0F087273ACFBA8B" box="[469,655,1909,1933]" pageId="5" pageNumber="461" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
, ♁ holotype (ABBM), left opisthomere (posterior gonopod).
<emphasis id="B908EA95FFC0FF9DA5D287273D49BA8B" bold="true" box="[1271,1289,1909,1933]" pageId="5" pageNumber="461">A</emphasis>
. Posteromesal view.
<emphasis id="B908EA95FFC0FF9DA1FC87CB38AABAB7" bold="true" box="[217,234,1945,1969]" pageId="5" pageNumber="461">B</emphasis>
. Mesal view.
<emphasis id="B908EA95FFC0FF9DA0AD87CB39DABAB7" bold="true" box="[392,410,1945,1969]" pageId="5" pageNumber="461">C</emphasis>
. Anterior view.
<emphasis id="B908EA95FFC0FF9DA36B87CB3A20BAB7" bold="true" box="[590,608,1945,1969]" pageId="5" pageNumber="461">D</emphasis>
. Anterolateral view.
<emphasis id="B908EA95FFC0FF9DA26087CB3B16BAB7" bold="true" box="[837,854,1945,1969]" pageId="5" pageNumber="461">E</emphasis>
. Lateral view.
<emphasis id="B908EA95FFC0FF9DA2D887CB3C4CBAB7" bold="true" box="[1021,1036,1945,1969]" pageId="5" pageNumber="461">F</emphasis>
. Posterolateral view.
<emphasis id="B908EA95FFC0FF9DA5DD87CB3D4CBAB7" bold="true" box="[1272,1292,1945,1969]" pageId="5" pageNumber="461">G</emphasis>
. Distal view.
<emphasis id="B908EA95FFC0FF9DA1B287EF3970BAD3" bold="true" box="[151,304,1981,2005]" pageId="5" pageNumber="461">Abbreviations</emphasis>
:
<emphasis id="B908EA95FFC0FF9DA06587EF3922BAD3" bold="true" box="[320,354,1981,2005]" pageId="5" pageNumber="461">am</emphasis>
: accessory membrane;
<emphasis id="B908EA95FFC0FF9DA37C87EF3A20BAD3" bold="true" box="[601,608,1981,2005]" pageId="5" pageNumber="461">l</emphasis>
: subtriangular lobe;
<emphasis id="B908EA95FFC0FF9DA21E87EF3B10BAD3" bold="true" box="[827,848,1981,2005]" pageId="5" pageNumber="461">m</emphasis>
: mesomeral process;
<emphasis id="B908EA95FFC0FF9DA51087EF3C02BAD3" bold="true" box="[1077,1090,1981,2005]" pageId="5" pageNumber="461">o</emphasis>
: opisthomere;
<emphasis id="B908EA95FFC0FF9DA5C587EF3CAABAD3" bold="true" box="[1248,1258,1981,2005]" pageId="5" pageNumber="461">s</emphasis>
: solenomere;
<emphasis id="B908EA95FFC0FF9DA45A87EF3DD6BAD3" bold="true" box="[1407,1430,1981,2005]" pageId="5" pageNumber="461">sg</emphasis>
: sperm groove;
<emphasis id="B908EA95FFC0FF9DA00A87B3397CBAFF" bold="true" box="[303,316,2017,2041]" pageId="5" pageNumber="461">v</emphasis>
: velum.
<emphasis id="B908EA95FFC0FF9DA0B087B339BABAFF" bold="true" box="[405,506,2017,2041]" pageId="5" pageNumber="461">Scale bar</emphasis>
: 0.1 mm.
</paragraph>
</caption>
<caption id="DF03660FFFC3FF9EA1B287213B00BAF0" ID-DOI="http://doi.org/10.5281/zenodo.8142350" ID-Zenodo-Dep="8142350" httpUri="https://zenodo.org/record/8142350/files/figure.png" pageId="6" pageNumber="462" startId="6.[151,250,1907,1931]" targetBox="[169,1423,189,1877]" targetPageId="6" targetType="figure">
<paragraph id="8BC33687FFC3FF9EA1B287213B00BAF0" blockId="6.[151,1437,1907,2039]" pageId="6" pageNumber="462">
<emphasis id="B908EA95FFC3FF9EA1B287213955BA8D" bold="true" box="[151,277,1907,1931]" pageId="6" pageNumber="462">FIGURE 5.</emphasis>
<taxonomicName id="4C7C4D04FFC3FF9EA03A8721399ABA8D" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[287,474,1907,1931]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="6" pageNumber="462" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC3FF9EA03A8721399ABA8D" box="[287,474,1907,1931]" italics="true" pageId="6" pageNumber="462">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC3FF9EA0C687213AE7BA8D" bold="true" box="[483,679,1907,1931]" pageId="6" pageNumber="462">
<taxonomicNameLabel id="A23B57EEFFC3FF9EA0C687213AE7BA8D" box="[483,679,1907,1931]" pageId="6" pageNumber="462" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
, ♁ holotype (ABBM), right gonopods.
<emphasis id="B908EA95FFC3FF9EA56F87213C1CBA8D" bold="true" box="[1098,1116,1907,1931]" pageId="6" pageNumber="462">A</emphasis>
. Right gonopods, lateral view.
<emphasis id="B908EA95FFC3FF9EA1B287C538E8BAA9" bold="true" box="[151,168,1943,1967]" pageId="6" pageNumber="462">B</emphasis>
. Right gonopods, mesal view.
<emphasis id="B908EA95FFC3FF9EA0D587C53A42BAA9" bold="true" box="[496,514,1943,1967]" pageId="6" pageNumber="462">C</emphasis>
. Right promere, posterior view.
<emphasis id="B908EA95FFC3FF9EA27987C53B2EBAA9" bold="true" box="[860,878,1943,1967]" pageId="6" pageNumber="462">D</emphasis>
. Right gonopods, anterior view.
<emphasis id="B908EA95FFC3FF9EA5EF87C53D23BAA9" bold="true" box="[1226,1379,1943,1967]" pageId="6" pageNumber="462">Abbreviations</emphasis>
:
<emphasis id="B908EA95FFC3FF9EA45687C53DD5BAA9" bold="true" box="[1395,1429,1943,1967]" pageId="6" pageNumber="462">am</emphasis>
: accessory membrane;
<emphasis id="B908EA95FFC3FF9EA05D87E9393FBAD5" bold="true" box="[376,383,1979,2003]" pageId="6" pageNumber="462">l</emphasis>
: subtriangular lobe;
<emphasis id="B908EA95FFC3FF9EA37487E93A23BAD5" bold="true" box="[593,611,1979,2003]" pageId="6" pageNumber="462">lr</emphasis>
: lateral ridge;
<emphasis id="B908EA95FFC3FF9EA3D287E93B4CBAD5" bold="true" box="[759,780,1979,2003]" pageId="6" pageNumber="462">m</emphasis>
: mesomeral process;
<emphasis id="B908EA95FFC3FF9EA2CD87E93BB5BAD5" bold="true" box="[1000,1013,1979,2003]" pageId="6" pageNumber="462">o</emphasis>
: opisthomere;
<emphasis id="B908EA95FFC3FF9EA5A987E93CDABAD5" bold="true" box="[1164,1178,1979,2003]" pageId="6" pageNumber="462">p</emphasis>
: promere;
<emphasis id="B908EA95FFC3FF9EA42C87E93D53BAD5" bold="true" box="[1289,1299,1979,2003]" pageId="6" pageNumber="462">s</emphasis>
: solenomere;
<emphasis id="B908EA95FFC3FF9EA1B2878D38EEBAF1" bold="true" box="[151,174,2015,2039]" pageId="6" pageNumber="462">sg</emphasis>
: sperm groove;
<emphasis id="B908EA95FFC3FF9EA076878D391BBAF1" bold="true" box="[339,347,2015,2039]" pageId="6" pageNumber="462">t</emphasis>
: triangular tooth;
<emphasis id="B908EA95FFC3FF9EA336878D3A60BAF1" bold="true" box="[531,544,2015,2039]" pageId="6" pageNumber="462">v</emphasis>
: velum.
<emphasis id="B908EA95FFC3FF9EA35C878D3A9EBAF1" bold="true" box="[633,734,2015,2039]" pageId="6" pageNumber="462">Scale bar</emphasis>
: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC33687FFC2FF9FA1E280C53D6ABDD3" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E280C53956BDB7" box="[199,278,151,177]" italics="true" pageId="7" pageNumber="463">Colour</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA00380C53928BDB7" box="[294,360,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1</figureCitation>
). Generally dichromatic. Head, first three rings, last two rings and telson orange, the rest of the rings dark indigo with anterior halves of prozonae pale. Antennae and legs greyish. Ommatidia blackish.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E2808D3B2FBF1F" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E2808D3944BDFF" box="[199,260,223,249]" italics="true" pageId="7" pageNumber="463">Head</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA037808D39D1BDFF" box="[274,401,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Figs 1AD</figureCitation>
,
<figureCitation id="13472A02FFC2FF9FA0B9808D3982BDFF" box="[412,450,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">2A</figureCitation>
). Number of ommatidia
<quantity id="4C849B62FFC2FF9FA3F6808C3B75BDFF" box="[723,821,222,249]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="7" pageNumber="463" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">2526 in</quantity>
7 horizontal rows, ocular fields rather trapezoidal (
<figureCitation id="13472A02FFC2FF9FA44A808D38A0BC1B" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1AD</figureCitation>
). With a pair of frontal setae (
<figureCitation id="13472A02FFC2FF9FA30281513AE3BC1B" box="[551,675,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1AD</figureCitation>
). Mandibular stipites expanded (
<figureCitation id="13472A02FFC2FF9FA52881513C24BC1B" box="[1037,1124,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1D</figureCitation>
, white arrow). Labrum with three labral teeth, 9+9 labral and 3+3 supralabral setae. Gnathochilarium (
<figureCitation id="13472A02FFC2FF9FA2C881753C08BC47" box="[1005,1096,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Fig. 2A</figureCitation>
) with a subdeltoid, elongated promentum; lamellae linguales with 5+5 setae in one longitudinal row, stipites with 3+3 long distolateral setae and 4+3 short medial setae. Antennae
<quantity id="4C849B62FFC2FF9FA334813D3A27BC8F" box="[529,615,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="463" unit="mm" value="1.2">1.2 mm</quantity>
long, their length 110% of vertical diameter of widest body ring. Lengths of antennomeres IVIII (in mm): 0.12 (I), 0.25 (II), 0.20 (III), 0.20 (IV), 0.25 (V), 0.10 (VI), 0.05 (VII) and 0.03 (VIII). Length/width ratio of antennomeres IVII: 1 (I), 2.3 (II), 1.7 (III), 1.7 (IV), 1.7 (V), 1 (VI) and 0.7 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E282713C85BF67" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E282713902BF3A" box="[199,322,547,573]" italics="true" pageId="7" pageNumber="463">Body rings</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA074827139ADBF3B" box="[337,493,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1A, C, E</figureCitation>
). Entire metazonal area with longitudinal striations. Metazonal setae long, length of midbody setae ca 45% of vertical diameter of rings, ca 20 per ring. Ozopores behind the suture.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E282393D76BFAF" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E28239394FBF83" box="[199,271,619,645]" italics="true" pageId="7" pageNumber="463">Telson</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA005823939DEBF83" box="[288,414,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1G, H</figureCitation>
). Epiproct with a short and rather blunt process. Paraprocts rounded, each with ca 15 long setae over the entire surface. Hypoproct lanceolate, without any modifications, with two long distal setae.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E282E13A84BE3F" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E282E138BCBFCA" box="[199,252,691,716]" italics="true" pageId="7" pageNumber="463">Legs</emphasis>
. First pair of legs modified, hook-shaped (
<figureCitation id="13472A02FFC2FF9FA3C482E13B7ABFCB" box="[737,826,691,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Fig. 2B</figureCitation>
), with three complete podomeres; coxa with one seta; prefemur with four setae; femur, postfemur and tibiotarsus coalesced; femur with two setae; postfemur with one seta; no other peculiarities.All other legs with postfemoral and tibial ventral pads (
<figureCitation id="13472A02FFC2FF9FA50682A93C3ABE13" box="[1059,1146,763,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Fig. 2C</figureCitation>
), and a setiform accessory tarsal claw almost twice as long as the main claw.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E283113C7BBE5B" blockId="7.[151,1437,151,1941]" box="[199,1083,835,861]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E283113936BE5B" box="[199,374,835,861]" italics="true" pageId="7" pageNumber="463">Pleurotergum 7</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA0A083113A5DBE5B" box="[389,541,835,861]" captionStart="FIGURE 1" captionStartId="2.[151,250,1870,1894]" captionTargetBox="[151,1436,179,1844]" captionTargetId="figure-15@2.[151,1436,179,1844]" captionTargetPageId="2" captionText="FIGURE 1. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), habitus. A. Whole body, lateral view. B. Head, right side, lateral, slightly ventral view. C. Anterior part of body, lateral view. D. Head and rings 13, left side, lateral view, arrow shows mandibular stipes. E. Midbody rings, lateral view. F. Right half of pleurotergum 7, lateral view. G. Telson, lateral view. H. Telson, posterior view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8142342" httpUri="https://zenodo.org/record/8142342/files/figure.png" pageId="7" pageNumber="463">Fig. 1A, C, F</figureCitation>
). With high, well-developed, subtriangular lobe.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E283353A7BBEA3" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E283353945BE86" box="[199,261,871,896]" italics="true" pageId="7" pageNumber="463">Penis</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA03083353930BE87" box="[277,368,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Fig. 2D</figureCitation>
). Small, not sclerotized, set deeply below coxae 2, rather trapezoidal, with two elongated lobes fanlike distally and a short basal part.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E283FD3D1DB89B" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E283FD397ABECF" box="[199,314,943,969]" italics="true" pageId="7" pageNumber="463">Gonopods</emphasis>
(
<figureCitation id="13472A02FFC2FF9FA06983FD398BBECF" box="[332,459,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Figs 2EG</figureCitation>
,
<figureCitation id="13472A02FFC2FF9FA0FC83FC3A46BECF" box="[473,518,942,969]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="4.[151,250,1943,1967]" captionStartId-1="5.[151,250,1909,1933]" captionStartId-2="6.[151,250,1907,1931]" captionTargetBox-0="[185,1401,181,1919]" captionTargetBox-1="[231,1357,181,1885]" captionTargetBox-2="[169,1423,189,1877]" captionTargetId-0="figure-15@4.[185,1401,181,1919]" captionTargetId-1="figure-19@5.[231,1357,181,1885]" captionTargetId-2="figure-15@6.[158,1430,181,1883]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 3. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), left promere (anterior gonopod). A. Posterior view. B. Lateral view. C. Anterolateral view. D. Anterior view. E. Anteromesal view. F. Mesal view. Abbreviations: lr: lateral ridge; t: triangular tooth. Scale bar: 0.1 mm." captionText-1="FIGURE 4. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), left opisthomere (posterior gonopod). A. Posteromesal view. B. Mesal view. C. Anterior view. D. Anterolateral view. E. Lateral view. F. Posterolateral view. G. Distal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; m: mesomeral process; o: opisthomere; s: solenomere; sg: sperm groove; v: velum. Scale bar: 0.1 mm." captionText-2="FIGURE 5. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), right gonopods. A. Right gonopods, lateral view. B. Right gonopods, mesal view. C. Right promere, posterior view. D. Right gonopods, anterior view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8142346" figureDoi-1="http://doi.org/10.5281/zenodo.8142348" figureDoi-2="http://doi.org/10.5281/zenodo.8142350" httpUri-0="https://zenodo.org/record/8142346/files/figure.png" httpUri-1="https://zenodo.org/record/8142348/files/figure.png" httpUri-2="https://zenodo.org/record/8142350/files/figure.png" pageId="7" pageNumber="463">35</figureCitation>
). Stout and rather robust. Anterior gonopods or promeres (
<emphasis id="B908EA95FFC2FF9FA58A83FD3CFFBECF" bold="true" box="[1199,1215,943,969]" pageId="7" pageNumber="463">p</emphasis>
in
<figureCitation id="13472A02FFC2FF9FA5CD83FD3D25BECF" box="[1256,1381,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Figs 2F, G</figureCitation>
,
<figureCitation id="13472A02FFC2FF9FA45683FC38ECBEEB" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1931]" captionTargetBox="[169,1423,189,1877]" captionTargetId="figure-15@6.[158,1430,181,1883]" captionTargetPageId="6" captionText="FIGURE 5. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), right gonopods. A. Right gonopods, lateral view. B. Right gonopods, mesal view. C. Right promere, posterior view. D. Right gonopods, anterior view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8142350" httpUri="https://zenodo.org/record/8142350/files/figure.png" pageId="7" pageNumber="463">5A, B</figureCitation>
;
<figureCitation id="13472A02FFC2FF9FA19F83813953BEEB" box="[186,275,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1360,181,1851]" captionTargetId="figure-19@3.[226,1360,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM). A. Gnathochilarium, ventral view. B. Right leg 1, lateral view. C. Distal part of right leg 2, posterior view. D. Penis, posterior view. E. Right promere, posterior view. F. Right gonopods, lateral view. G. Right gonopods, mesal view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142344" httpUri="https://zenodo.org/record/8142344/files/figure.png" pageId="7" pageNumber="463">Figs 2E</figureCitation>
,
<figureCitation id="13472A02FFC2FF9FA0058381396FBEEB" box="[288,303,979,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[185,1401,181,1919]" captionTargetId="figure-15@4.[185,1401,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), left promere (anterior gonopod). A. Posterior view. B. Lateral view. C. Anterolateral view. D. Anterior view. E. Anteromesal view. F. Mesal view. Abbreviations: lr: lateral ridge; t: triangular tooth. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8142346" httpUri="https://zenodo.org/record/8142346/files/figure.png" pageId="7" pageNumber="463">3</figureCitation>
,
<figureCitation id="13472A02FFC2FF9FA018838039C1BEEB" box="[317,385,978,1005]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1931]" captionTargetBox="[169,1423,189,1877]" captionTargetId="figure-15@6.[158,1430,181,1883]" captionTargetPageId="6" captionText="FIGURE 5. Karadenizia merti gen. nov., sp. nov., ♁ holotype (ABBM), right gonopods. A. Right gonopods, lateral view. B. Right gonopods, mesal view. C. Right promere, posterior view. D. Right gonopods, anterior view. Abbreviations: am: accessory membrane; l: subtriangular lobe; lr: lateral ridge; m: mesomeral process; o: opisthomere; p: promere; s: solenomere; sg: sperm groove; t: triangular tooth; v: velum. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8142350" httpUri="https://zenodo.org/record/8142350/files/figure.png" pageId="7" pageNumber="463">5C, D</figureCitation>
) spoon-shaped; lateral ridge (
<emphasis id="B908EA95FFC2FF9FA3F183813AA8BEEB" bold="true" box="[724,744,979,1005]" pageId="7" pageNumber="463">lr</emphasis>
), with 5 or 6 setae, extending continuously to half the height of promere, then continuing discontinuously to 2/3 of height and moving towards mesal side; mesal margin with noticeable folds in basal and central part, distally with a triangular tooth (
<emphasis id="B908EA95FFC2FF9FA2D784493BBBB933" bold="true" box="[1010,1019,1051,1077]" pageId="7" pageNumber="463">t</emphasis>
); distal half with diagonal posterior shallow cavity to accommodate mesomeral process. Posterior gonopods same height as promere, with two parts also of same height: a mesomeral process (
<emphasis id="B908EA95FFC2FF9FA3AB84313AE5B97B" bold="true" box="[654,677,1123,1149]" pageId="7" pageNumber="463">m</emphasis>
) and an opisthomere (
<emphasis id="B908EA95FFC2FF9FA28884313BFBB97B" bold="true" box="[941,955,1123,1149]" pageId="7" pageNumber="463">o</emphasis>
). Mesomeral process strongly developed; anteriorly originating from base of posterior gonopod, laterally fused to opisthomere at half its height; anterolaterally very wide; distal half slender, digitiform, significantly curved mesad and slightly laterad; mesally with accessory membrane (
<emphasis id="B908EA95FFC2FF9FA03A849D3904B9EF" bold="true" box="[287,324,1231,1257]" pageId="7" pageNumber="463">am</emphasis>
) forming a pocket to connect mesomeral process and opisthomere from mesal side. Opisthomere slightly sigmoid in mesal and lateral view; sperm groove (
<emphasis id="B908EA95FFC2FF9FA21084A13B0EB80B" bold="true" box="[821,846,1267,1293]" pageId="7" pageNumber="463">sg</emphasis>
) beginning mesally from mesal pocket,, extending over entire height of opisthomere and terminating in distal part, just proximally to beaklike solenomere (
<emphasis id="B908EA95FFC2FF9FA46885453D18B837" bold="true" box="[1357,1368,1303,1329]" pageId="7" pageNumber="463">s</emphasis>
) with spoutlike anterior part. Anteriorly, below solenomere, a strongly developed fimbriate velum (
<emphasis id="B908EA95FFC2FF9FA58B85693CFCB853" bold="true" box="[1198,1212,1339,1365]" pageId="7" pageNumber="463">v</emphasis>
) or anterior lamella is formed. The entire distal part of opisthomere in mesal and lateral view resembles head of a singing songbird (with open beak). Additionally, mesal side of opisthomere has a rib or ridge ending in a small subtriangular lobe (
<emphasis id="B908EA95FFC2FF9FA46985D13D14B89B" bold="true" box="[1356,1364,1411,1437]" pageId="7" pageNumber="463">l</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFC2FF90A1E285F53D43BAE3" lastPageId="8" lastPageNumber="464" pageId="7" pageNumber="463" type="discussion">
<paragraph id="8BC33687FFC2FF9FA1E285F53C7EBB9F" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E285F5397BB8C7" bold="true" box="[199,315,1447,1473]" pageId="7" pageNumber="463">Remarks.</emphasis>
Although this species is characterised by a particular colour pattern, it is not unique in this sense, as there are other millipedes with a similar pattern (see in the discussion). Interestingly, a very similar looking animal (
<figureCitation id="13472A02FFC2FF9FA1BA85BD38A3BB0F" box="[159,227,1519,1545]" captionStart="FIGURE 6" captionStartId="8.[151,250,866,890]" captionTargetBox="[151,1436,181,842]" captionTargetId="figure-15@8.[151,1436,181,842]" captionTargetPageId="8" captionText="FIGURE 6. An unidentified millipede from a cave near Trabzon, similar in appearance to Karadenizia merti gen. nov., sp. nov., in situ. Photo by Mert Elverici." figureDoi="http://doi.org/10.5281/zenodo.8142352" httpUri="https://zenodo.org/record/8142352/files/figure.png" pageId="7" pageNumber="463">Fig. 6</figureCitation>
) was found in the cave near
<collectingRegion id="49B8F865FFC2FF9FA30385BD3AC5BB0F" box="[550,645,1519,1545]" country="Turkey" name="Trabzon" pageId="7" pageNumber="463">Trabzon</collectingRegion>
, also on the southern coast of the Black Sea, but ca
<quantity id="4C849B62FFC2FF9FA5EB85BD3D63BB0F" box="[1230,1315,1519,1545]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="463" unit="km" value="600.0">600 km</quantity>
to the east of where
<taxonomicName id="4C7C4D04FFC2FF9FA0278641398EBB2A" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[258,462,1555,1581]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="7" pageNumber="463" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFC2FF9FA0278641398EBB2A" box="[258,462,1555,1581]" italics="true" pageId="7" pageNumber="463">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFC2FF9FA0F386413AE8BB2B" bold="true" box="[470,680,1555,1581]" pageId="7" pageNumber="463">
<taxonomicNameLabel id="A23B57EEFFC2FF9FA0F386413AE8BB2B" box="[470,680,1555,1581]" pageId="7" pageNumber="463" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
was found. Unfortunately, the only specimen collected is a female. Although both animals have a moniliform body and orange anterior and posterior body ends, the female from near
<collectingRegion id="49B8F865FFC2FF9FA1B2860938B3BB73" box="[151,243,1627,1653]" country="Turkey" name="Trabzon" pageId="7" pageNumber="463">Trabzon</collectingRegion>
differs from the species described here by the presence of an elliptical ocular field, by conspicuously shorter metazonal setae and by the presence of a black transverse band on the head.
</paragraph>
<paragraph id="8BC33687FFC2FF9FA1E286F03C9EBA93" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="463">
<emphasis id="B908EA95FFC2FF9FA1E286F03981BBBB" bold="true" box="[199,449,1698,1725]" pageId="7" pageNumber="463">Locality and ecology.</emphasis>
The Erçek Cave (
<figureCitation id="13472A02FFC2FF9FA3AC86F13A8CBBBB" box="[649,716,1699,1725]" captionStart="FIGURE 7" captionStartId="8.[151,250,1780,1804]" captionTargetBox="[151,1437,986,1757]" captionTargetId="figure-46@8.[151,1437,986,1757]" captionTargetPageId="8" captionText="FIGURE 7. Entrance to the Erçek Cave, the type locality of Karadenizia merti gen. nov., sp. nov.. Photo by Ertuğrul Kulaksızoğlu." figureDoi="http://doi.org/10.5281/zenodo.8142354" httpUri="https://zenodo.org/record/8142354/files/figure.png" pageId="7" pageNumber="463">Fig. 7</figureCitation>
) is a horizontal cave with a planimetric length of 890 metres and a depth of -26 metres. The entrance of the cave lies in a north-western direction, is rectangular, 14 x
<quantity id="4C849B62FFC2FF9FA43486953D0BBBE7" box="[1297,1355,1735,1761]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="463" unit="m" value="14.0">14 m</quantity>
in size and forms a large habitat in the twilight zone. The cave bifurcates shortly after the entrance. The left part is mainly active, while the right part (including the entrance) is semi-active or fossil and forms a large terrestrial habitat. The active (left) part of the cave has the shape of a narrow, weakly flowing water stream with many meanders, flowing through a huge massif of red marble, with the floor almost completely submerged. The cave is also characterised by beautiful cave speleothems, especially stalactites in different colours (Mert Elverici, pers. comm.)
</paragraph>
<caption id="DF03660FFFCDFF90A1B283303993BE98" ID-DOI="http://doi.org/10.5281/zenodo.8142352" ID-Zenodo-Dep="8142352" httpUri="https://zenodo.org/record/8142352/files/figure.png" pageId="8" pageNumber="464" startId="8.[151,250,866,890]" targetBox="[151,1436,181,842]" targetPageId="8" targetType="figure">
<paragraph id="8BC33687FFCDFF90A1B283303993BE98" blockId="8.[151,1436,866,926]" pageId="8" pageNumber="464">
<emphasis id="B908EA95FFCDFF90A1B283303952BE7C" bold="true" box="[151,274,866,890]" pageId="8" pageNumber="464">FIGURE 6.</emphasis>
An unidentified millipede from a cave near Trabzon, similar in appearance to
<taxonomicName id="4C7C4D04FFCDFF90A53A83303C96BE7C" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[1055,1238,866,890]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="8" pageNumber="464" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFCDFF90A53A83303C96BE7C" box="[1055,1238,866,890]" italics="true" pageId="8" pageNumber="464">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFCDFF90A5F983303DD5BE7C" bold="true" box="[1244,1429,866,890]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A23B57EEFFCDFF90A5F983303DD5BE7C" box="[1244,1429,866,890]" pageId="8" pageNumber="464" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B908EA95FFCDFF90A1B283D53896BE98" box="[151,214,903,926]" italics="true" pageId="8" pageNumber="464">in situ</emphasis>
. Photo by Mert Elverici.
</paragraph>
</caption>
<caption id="DF03660FFFCDFF90A1B286A63966BA37" ID-DOI="http://doi.org/10.5281/zenodo.8142354" ID-Zenodo-Dep="8142354" httpUri="https://zenodo.org/record/8142354/files/figure.png" pageId="8" pageNumber="464" startId="8.[151,250,1780,1804]" targetBox="[151,1437,986,1757]" targetPageId="8" targetType="figure">
<paragraph id="8BC33687FFCDFF90A1B286A63966BA37" blockId="8.[151,1436,1780,1841]" pageId="8" pageNumber="464">
<emphasis id="B908EA95FFCDFF90A1B286A63958BA0B" bold="true" box="[151,280,1780,1805]" pageId="8" pageNumber="464">FIGURE 7.</emphasis>
Entrance to the Erçek Cave, the type locality of
<taxonomicName id="4C7C4D04FFCDFF90A21386A73BB4BA0A" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[822,1012,1781,1805]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="8" pageNumber="464" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFCDFF90A21386A73BB4BA0A" box="[822,1012,1781,1805]" italics="true" pageId="8" pageNumber="464">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFCDFF90A52586A73C8BBA0B" bold="true" box="[1024,1227,1781,1805]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A23B57EEFFCDFF90A52586A73C8BBA0B" box="[1024,1227,1781,1805]" pageId="8" pageNumber="464" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
. Photo by Ertuğrul Kulaksızoğlu.
</paragraph>
</caption>
<paragraph id="8BC33687FFCDFF90A1E2870D3BD2BAC7" blockId="8.[151,1436,1887,2021]" pageId="8" pageNumber="464">
The
<typeStatus id="54C78825FFCDFF90A1D9870D391EBA7F" box="[252,350,1887,1913]" pageId="8" pageNumber="464">holotype</typeStatus>
was found crawling on the ground in the twilight zone, while the
<typeStatus id="54C78825FFCDFF90A57F870D3CFABA7F" box="[1114,1210,1887,1913]" pageId="8" pageNumber="464">paratype</typeStatus>
juvenile was found clinging to the red marble wall in the active part of the cave. Besides this new genus, the cave is also inhabited by a possibly new species of the genus
<taxonomicName id="4C7C4D04FFCDFF90A33C87F53B0DBAC7" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[537,845,1959,1985]" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B908EA95FFCDFF90A33C87F53ADFBAC7" box="[537,671,1959,1985]" italics="true" pageId="8" pageNumber="464">Polydesmus</emphasis>
<bibRefCitation id="EFED4B76FFCDFF90A38387F53B0DBAC7" author="Latreille, P. A." box="[678,845,1959,1985]" pageId="8" pageNumber="464" refId="ref5720" refString="Latreille, P. A. (1802) Histoire naturelle, generale et particuliere des crustaces et des insectes. Bd. 3. de L'imprimerie de F. Dufart, Paris, 467 pp. https: // doi. org / 10.5962 / bhl. title. 15764" type="book" year="1802">Latreille, 1802</bibRefCitation>
</taxonomicName>
/1803.
</paragraph>
<paragraph id="8BC33687FFCDFF90A1E287993D43BAE3" blockId="8.[151,1436,1887,2021]" box="[199,1283,1995,2021]" pageId="8" pageNumber="464">
<taxonomicName id="4C7C4D04FFCDFF90A1E2879939D4BAE3" authority="Antić, 2023" authorityName="Antić" authorityYear="2023" box="[199,404,1995,2021]" class="Diplopoda" family="Julidae" genus="Karadenizia" kingdom="Animalia" order="Julida" pageId="8" pageNumber="464" phylum="Arthropoda" rank="species" species="merti" status="gen. nov., sp. nov.">
<emphasis id="B908EA95FFCDFF90A1E2879939D4BAE3" box="[199,404,1995,2021]" italics="true" pageId="8" pageNumber="464">Karadenizia merti</emphasis>
</taxonomicName>
<emphasis id="B908EA95FFCDFF90A0BE87993A2BBAE3" bold="true" box="[411,619,1995,2021]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A23B57EEFFCDFF90A0BE87993A2BBAE3" box="[411,619,1995,2021]" pageId="8" pageNumber="464" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
is probably an epigean taxon without troglobiomorphisms.
</paragraph>
</subSubSection>
<subSubSection id="C366650CFFCCFF91A1E280C43ABDBDD3" pageId="9" pageNumber="465" type="distribution">
<paragraph id="8BC33687FFCCFF91A1E280C43ABDBDD3" blockId="9.[151,1436,150,213]" pageId="9" pageNumber="465">
<emphasis id="B908EA95FFCCFF91A1E280C43921BDB6" bold="true" box="[199,353,150,176]" pageId="9" pageNumber="465">Distribution.</emphasis>
A Turkish endemic known so far only from its
<typeStatus id="54C78825FFCCFF91A25E80C53BEBBDB7" box="[891,939,151,177]" pageId="9" pageNumber="465">type</typeStatus>
locality, Erçek cave, near Erçek village, the southern coast of the Black Sea (
<collectingRegion id="49B8F865FFCCFF91A32380E93AB1BDD3" box="[518,753,187,213]" country="Turkey" name="Zonguldak" pageId="9" pageNumber="465">Zonguldak Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>