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<document id="D136F8D64C240173C32D7DCBA658719E" ID-DOI="10.11646/zootaxa.5424.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821192" ID-ZooBank="9F446F11-2997-43AE-BD75-1AA60049D321" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1710500715128" checkinUser="plazi" docAuthor="Du, Ruolan, Yang, Yuxia, Yang, Xingke &amp; Liu, Haoyu" docDate="2024" docId="03D387DEFF90FF98CBF3FB190371FDFF" docLanguage="en" docName="zootaxa.5424.3.4.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrolycus subapicis Y. Yang, Du &amp; Liu 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="364" masterDocId="FFEAFFA6FF94FF9ECB64FF8C0150FFE6" masterDocTitle="A taxonomic study on the nominate subgenus Macrolycus Waterhouse, 1878 from China (Coleoptera, Lycidae)" masterLastPageNumber="366" masterPageNumber="358" pageNumber="362" updateTime="1710892380381" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="048E9C554BCA4EF37BB044C05BB7F6C6">A taxonomic study on the nominate subgenus Macrolycus Waterhouse, 1878 from China (Coleoptera, Lycidae)</mods:title>
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<mods:namePart id="78727BAE6E10C4EF35593C5712D871C6">Du, Ruolan</mods:namePart>
<mods:affiliation id="C1CBABD723C23D76B7B6112511A13115">Key Laboratory of Zoological Systematics and Application, School of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding 071002, Hebei Province, China</mods:affiliation>
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<mods:namePart id="E8D1E8DCE6B942A6C8B47FBC974234B5">Yang, Yuxia</mods:namePart>
<mods:affiliation id="1A7CA83FEA4458A260538F086D8BB1AC">Key Laboratory of Zoological Systematics and Application, School of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding 071002, Hebei Province, China</mods:affiliation>
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<mods:namePart id="CEE8B0118467CA4F5E3523D7E860BA8A">Yang, Xingke</mods:namePart>
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<mods:namePart id="1CFFF12AB8CE43150CA2CA0D51931E38">Liu, Haoyu</mods:namePart>
<mods:affiliation id="C96E027ED6084CE7EB56223B2623C4A0">Key Laboratory of Zoological Systematics and Application, School of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding 071002, Hebei Province, China</mods:affiliation>
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<subSubSection id="C3606543FF90FF9ACBF3FB1903A7FB56" box="[151,759,1173,1200]" pageId="4" pageNumber="362" type="nomenclature">
<paragraph id="8BC536C8FF90FF9ACBF3FB1903A7FB56" blockId="4.[151,759,1173,1236]" box="[151,759,1173,1200]" pageId="4" pageNumber="362">
<heading id="D08D81A4FF90FF9ACBF3FB1903A7FB56" bold="true" box="[151,759,1173,1200]" fontSize="11" level="1" pageId="4" pageNumber="362" reason="1">
<emphasis id="B90EEADAFF90FF9ACBF3FB1903A7FB56" bold="true" box="[151,759,1173,1200]" pageId="4" pageNumber="362">
<taxonomicName id="4C7A4D4BFF90FF9ACBF3FB1903DDFB49" authority="Y. Yang, Du &amp; Liu" authorityName="Y. Yang, Du &amp; Liu" authorityYear="2024" box="[151,653,1173,1200]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="subapicis" status="sp. nov.">
<emphasis id="B90EEADAFF90FF9ACBF3FB1900CCFB56" bold="true" box="[151,412,1173,1200]" italics="true" pageId="4" pageNumber="362">Macrolycus subapicis</emphasis>
Y. Yang, Du &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A23D57A1FF90FF9AC9FEFB1A03A7FB56" box="[666,759,1174,1200]" pageId="4" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3606543FF90FF9ACBF3FB360019FB32" box="[151,329,1210,1236]" pageId="4" pageNumber="362" type="description">
<paragraph id="8BC536C8FF90FF9ACBF3FB360019FB32" blockId="4.[151,759,1173,1236]" box="[151,329,1210,1236]" pageId="4" pageNumber="362">
(
<figureCitation id="13412A4DFF90FF9ACBFBFB3601ACFB32" box="[159,252,1210,1236]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="4" pageNumber="362">Figs 2D</figureCitation>
,
<figureCitation id="13412A4DFF90FF9ACA6CFB360011FB32" box="[264,321,1210,1236]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="4" pageNumber="362">3JL</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3606543FF90FF9ACBF3FA8D0594FA12" pageId="4" pageNumber="362" type="diagnosis">
<paragraph id="8BC536C8FF90FF9ACBF3FA8D0047FAFD" blockId="4.[151,1437,1281,1992]" box="[151,279,1281,1307]" pageId="4" pageNumber="362">
<heading id="D08D81A4FF90FF9ACBF3FA8D0047FAFD" bold="true" box="[151,279,1281,1307]" fontSize="11" level="1" pageId="4" pageNumber="362" reason="1">
<emphasis id="B90EEADAFF90FF9ACBF3FA8D0047FAFD" bold="true" box="[151,279,1281,1307]" pageId="4" pageNumber="362">Diagnosis.</emphasis>
</heading>
</paragraph>
<paragraph id="8BC536C8FF90FF9ACBF3FAAA0594FA12" blockId="4.[151,1437,1281,1992]" pageId="4" pageNumber="362">
The species resembles
<taxonomicName id="4C7A4D4BFF90FF9ACAFDFAAB03B9FAA6" authority="Kazantsev, 2001" authorityName="Kazantsev" authorityYear="2001" box="[409,745,1318,1344]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90EEADAFF90FF9ACAFDFAAB0379FAA6" box="[409,553,1319,1344]" italics="true" pageId="4" pageNumber="362">M. yunnanus</emphasis>
<bibRefCitation id="EFEB4B39FF90FF9AC954FAAA03B9FAA6" author="Kazantsev, S. V." box="[560,745,1318,1344]" pageId="4" pageNumber="362" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">
Kazantsev,
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in appearance, but can be distinguished from the latter by the lamella of antennomere III 1.1 times as long as the antennomere itself (
<figureCitation id="13412A4DFF90FF9AC8D3FAC60542FA82" box="[951,1042,1354,1380]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="4" pageNumber="362">Fig. 2D</figureCitation>
); subapical part of phallus strongly inflated laterally in ventral view, 3.5 times as wide as basal part (
<figureCitation id="13412A4DFF90FF9AC811FAE2029DFA6E" box="[885,973,1390,1416]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="4" pageNumber="362">Fig. 3K</figureCitation>
) and strongly inflated ventrally in lateral view (
<figureCitation id="13412A4DFF90FF9ACBB8FA1E0063FA4A" box="[220,307,1426,1452]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="4" pageNumber="362">Fig. 3L</figureCitation>
). In comparison, the lamella of antennomere III of
<taxonomicName id="4C7A4D4BFF90FF9AC80FFA1F02ABFA4A" authorityName="Kazantsev" authorityYear="2001" box="[875,1019,1427,1452]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90EEADAFF90FF9AC80FFA1F02ABFA4A" box="[875,1019,1427,1452]" italics="true" pageId="4" pageNumber="362">M. yunnanus</emphasis>
</taxonomicName>
0.8 times as long as the antennomere itself (
<bibRefCitation id="EFEB4B39FF90FF9ACBBAFA3A00C8FA36" author="Kazantsev, S. V." box="[222,408,1462,1488]" pageId="4" pageNumber="362" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 1); subapical part of phallus feebly inflated laterally in ventral view, 2.5 times as wide as basal part (
<bibRefCitation id="EFEB4B39FF90FF9ACA54FA5600BAFA12" author="Kazantsev, S. V." box="[304,490,1498,1524]" pageId="4" pageNumber="362" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 2) and flat ventrally in lateral view (
<bibRefCitation id="EFEB4B39FF90FF9AC8D1FA56053EFA12" author="Kazantsev, S. V." box="[949,1134,1498,1524]" pageId="4" pageNumber="362" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 3).
</paragraph>
</subSubSection>
<subSubSection id="C3606543FF90FF9ACBA3FA72027AF9DA" pageId="4" pageNumber="362" type="etymology">
<paragraph id="8BC536C8FF90FF9ACBA3FA72027AF9DA" blockId="4.[151,1437,1281,1992]" pageId="4" pageNumber="362">
<emphasis id="B90EEADAFF90FF9ACBA3FA72001EF9FE" bold="true" box="[199,334,1534,1560]" pageId="4" pageNumber="362">Etymology.</emphasis>
The specific name is derived from the Latin
<emphasis id="B90EEADAFF90FF9AC821FA720225F9FE" box="[837,885,1534,1560]" italics="true" pageId="4" pageNumber="362">sub-</emphasis>
(under, below, beneath) and
<emphasis id="B90EEADAFF90FF9ACFDDFA7305BEF9FE" box="[1209,1262,1535,1560]" italics="true" pageId="4" pageNumber="362">apex</emphasis>
(summit, peak, tip), referring to its characteristic subapical part of phallus.
</paragraph>
</subSubSection>
<subSubSection id="C3606543FF90FF9ACBA3F9CA0191F962" pageId="4" pageNumber="362" type="materials_examined">
<paragraph id="8BC536C8FF90FF9ACBA3F9CA0191F962" blockId="4.[151,1437,1281,1992]" pageId="4" pageNumber="362">
<emphasis id="B90EEADAFF90FF9ACBA3F9CA00A4F986" bold="true" box="[199,500,1605,1632]" pageId="4" pageNumber="362">
Type material.
<typeStatus id="54C1886AFF90FF9ACAE4F9CA00BFF986" box="[384,495,1606,1632]" pageId="4" pageNumber="362" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F36D7658FF90FF9ACA99F9CA0304F986" box="[509,596,1606,1632]" name="China" pageId="4" pageNumber="362">CHINA</collectingCountry>
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(MHBU
<emphasis id="B90EEADAFF90FF9AC988F9C903A5F9B9" bold="true" box="[748,757,1605,1631]" pageId="4" pageNumber="362">)</emphasis>
,
<collectingRegion id="49BEF82AFF90FF9AC862F9CA020FF986" box="[774,863,1606,1632]" country="China" name="Yunnan" pageId="4" pageNumber="362">Yunnan</collectingRegion>
, Nujiang, Biyuhe,
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,
<date id="FFC41008FF90FF9ACFBFF9CA0413F986" box="[1243,1347,1606,1632]" pageId="4" pageNumber="362" value="2023-07">VII.2023</date>
, Z. Pan leg.
</paragraph>
</subSubSection>
<subSubSection id="C3606543FF90FF98CBA3F90100F2FE13" lastPageId="6" lastPageNumber="364" pageId="4" pageNumber="362" type="description">
<paragraph id="8BC536C8FF90FF9ACBA3F90102A7F94E" blockId="4.[151,1437,1281,1992]" box="[199,1015,1677,1704]" pageId="4" pageNumber="362">
<emphasis id="B90EEADAFF90FF9ACBA3F9010001F941" bold="true" box="[199,337,1677,1703]" pageId="4" pageNumber="362">Description</emphasis>
.
<emphasis id="B90EEADAFF90FF9ACA04F90200CDF94E" bold="true" box="[352,413,1678,1704]" pageId="4" pageNumber="362">Male</emphasis>
(
<figureCitation id="13412A4DFF90FF9ACAC8F9020356F94E" box="[428,518,1678,1704]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="4" pageNumber="362">Fig. 2D</figureCitation>
). Length 12.0 mm, width at humeri
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.
</paragraph>
<paragraph id="8BC536C8FF90FF9ACBA3F93E0414F92A" blockId="4.[151,1437,1281,1992]" box="[199,1348,1714,1740]" pageId="4" pageNumber="362">Body black. Pronotum and elytra orange red. Surface covered with decumbent orange red pubescence.</paragraph>
<paragraph id="8BC536C8FF90FF9ACBA3F95A0057F8BA" blockId="4.[151,1437,1281,1992]" pageId="4" pageNumber="362">
Eyes small, interocular distance about twice greater than the diameter of an eye.Antennae flabellate, overlapping basal 2/3 length of elytra when inclined, antennomeres IIIXI lamellate and slender, lamellae pointed at apices, lamella of III 1.1 times as long as the antennomere itself, lamella of VIII longest, 2.6 times longer than joint itself (
<figureCitation id="13412A4DFF90FF9ACBFBF8CE01AAF8BA" box="[159,250,1858,1884]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="4" pageNumber="362">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8BC536C8FF90FF9ACBA3F8EA0274F82E" blockId="4.[151,1437,1281,1992]" pageId="4" pageNumber="362">
Pronotum trapezoidal, 1.1 times wider than long. Anterior margin widely rounded and projecting anteriad, lateral margins sinuate and posterior margin bisinuate; anterior angles rounded, posterior angles sharp and projecting laterally. Scutellum trapezoidal, straight at apex (
<figureCitation id="13412A4DFF90FF9AC9D8F8220247F82E" box="[700,791,1966,1992]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="4" pageNumber="362">Fig. 2D</figureCitation>
).
</paragraph>
<caption id="DF056640FF91FF9BCBFCF81B03CFF811" ID-DOI="http://doi.org/10.5281/zenodo.10821200" ID-Zenodo-Dep="10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="5" pageNumber="363" startId="5.[152,255,1943,1967]" targetBox="[216,1373,185,1909]" targetPageId="5" targetType="figure">
<paragraph id="8BC536C8FF91FF9BCBFCF81B03CFF811" blockId="5.[151,1437,1943,2039]" pageId="5" pageNumber="363">
<emphasis id="B90EEADAFF91FF9BCBFCF81B0047F849" bold="true" box="[152,279,1943,1967]" pageId="5" pageNumber="363">FIGURE 3.</emphasis>
Aedeagi of
<taxonomicName id="4C7A4D4BFF91FF9BCAEAF81B0305F849" box="[398,597,1943,1967]" pageId="5" pageNumber="363">
<emphasis id="B90EEADAFF91FF9BCAEAF81B0355F849" box="[398,517,1943,1967]" italics="true" pageId="5" pageNumber="363">Macrolycus</emphasis>
species
</taxonomicName>
, dorsal views: AC.
<taxonomicName id="4C7A4D4BFF91FF9BC845F814050BF849" authority="Kazantsev, 2002" authorityName="Kazantsev" authorityYear="2002" box="[801,1115,1943,1967]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="oreophilus">
<emphasis id="B90EEADAFF91FF9BC845F81402FCF849" box="[801,940,1943,1967]" italics="true" pageId="5" pageNumber="363">M. oreophilus</emphasis>
<bibRefCitation id="EFEB4B39FF91FF9BC8D6F814050BF849" author="Kazantsev, S. V." box="[946,1115,1943,1967]" pageId="5" pageNumber="363" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev, 2002</bibRefCitation>
</taxonomicName>
; DF.
<taxonomicName id="4C7A4D4BFF91FF9BCFFEF814019DF835" authority="Kazantsev, 2002" authorityName="Kazantsev" authorityYear="2002" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="B90EEADAFF91FF9BCFFEF8140479F849" box="[1178,1321,1944,1967]" italics="true" pageId="5" pageNumber="363">M. gansuensis</emphasis>
<bibRefCitation id="EFEB4B39FF91FF9BCE4AF814019DF835" author="Kazantsev, S. V." pageId="5" pageNumber="363" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev, 2002</bibRefCitation>
</taxonomicName>
; GI.
<taxonomicName id="4C7A4D4BFF91FF9BCA68F830031AF835" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[268,586,1979,2003]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="bowringi">
<emphasis id="B90EEADAFF91FF9BCA68F83000DAF835" box="[268,394,1979,2003]" italics="true" pageId="5" pageNumber="363">M. bowringi</emphasis>
<bibRefCitation id="EFEB4B39FF91FF9BCAF4F837031AF835" author="Waterhouse, C. O." box="[400,586,1979,2003]" pageId="5" pageNumber="363" pagination="95 - 118" refId="ref6154" refString="Waterhouse, C. O. (1878) On the different forms occurring in the Coleopterous family Lycidae, with descriptions of new genera and species. Transactions of the Entomological Society of London, 26, 95 - 118. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01944. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
</taxonomicName>
; JL.
<taxonomicName id="4C7A4D4BFF91FF9BC9E3F8300259F835" authority="Du &amp; Yang &amp; Yang &amp; Liu, 2024" authorityName="Du &amp; Yang &amp; Yang &amp; Liu" authorityYear="2024" box="[647,777,1979,2003]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="subapicis" status="sp. nov.">
<emphasis id="B90EEADAFF91FF9BC9E3F8300259F835" box="[647,777,1979,2003]" italics="true" pageId="5" pageNumber="363">M. subapicis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57A1FF91FF9BC86BF830020FF835" box="[783,863,1980,2003]" pageId="5" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
(holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BC536C8FF92FF98CBA3FF1B00ECFF33" blockId="6.[151,1437,151,537]" pageId="6" pageNumber="364">
Elytra leaf-like, 3.1 times longer than humeral width. Costa I as strong as II and IV, costa III weak but visible in whole length (
<figureCitation id="13412A4DFF92FF98CA31FF3700FFFF33" box="[341,431,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="6" pageNumber="364">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8BC536C8FF92FF98CBA3FF5303BEFE37" blockId="6.[151,1437,151,537]" pageId="6" pageNumber="364">
Aedeagus (
<figureCitation id="13412A4DFF92FF98CA22FF5300E9FF1F" box="[326,441,223,249]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3JL</figureCitation>
) moderately sclerotized. Parameres moderately developed and spoon-shaped, about 2/7 length of phallus, with lateral margins converging apically in ventral view (
<figureCitation id="13412A4DFF92FF98CF6DFE8F0533FEFB" box="[1033,1123,259,285]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3K</figureCitation>
) and widely emarginate in lateral view (
<figureCitation id="13412A4DFF92FF98CA55FEAB00DBFEA7" box="[305,395,295,321]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3L</figureCitation>
). Phallus abruptly expanded at basal 1/4 portion, then parallel-sided in dorsal and ventral views (
<figureCitation id="13412A4DFF92FF98CB8DFEC70032FE83" box="[233,354,331,357]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3J, K</figureCitation>
), subapical part strongly and asymmetrically inflated laterally, about 3.5 times as wide as basal part, with a fusiform ventral-cavity, apical part parallel-sided, apex about 0.32 times as wide as subapical part; basal part bisinuate at basal part and bent dorsally near middle in lateral view (
<figureCitation id="13412A4DFF92FF98C8B6FE1F0578FE4B" box="[978,1064,403,429]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3L</figureCitation>
), subapical part strongly inflated ventrally, apical part progressively constricted distad.
</paragraph>
<paragraph id="8BC536C8FF92FF98CBA3FE5700F2FE13" blockId="6.[151,1437,151,537]" box="[199,418,475,501]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CBA3FE570075FE13" bold="true" box="[199,293,475,501]" pageId="6" pageNumber="364">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3606543FF92FF98CBA3FE720371FDFF" box="[199,545,510,537]" pageId="6" pageNumber="364" type="distribution">
<paragraph id="8BC536C8FF92FF98CBA3FE720371FDFF" blockId="6.[151,1437,151,537]" box="[199,545,510,537]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CBA3FE720031FDFE" bold="true" box="[199,353,510,536]" pageId="6" pageNumber="364">Distribution.</emphasis>
<collectingCountry id="F36D7658FF92FF98CA03FE7300FAFDFF" box="[359,426,511,537]" name="China" pageId="6" pageNumber="364">China</collectingCountry>
(
<collectingRegion id="49BEF82AFF92FF98CADDFE730345FDFF" box="[441,533,511,537]" country="China" name="Yunnan" pageId="6" pageNumber="364">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>