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<document id="9A5A83AE5618F4490759C0B7B974FEA7" ID-CLB-Dataset="25875" ID-DOI="10.11646/zootaxa.4661.2.5" ID-GBIF-Dataset="eda0d38a-f1b2-43cb-bb9b-7b8296e4f60f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3379495" ID-ZooBank="71122FBE-0022-4D1F-B200-3946D770CAE8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566978690026" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2019" docId="03CEF84AFFB5FFEBFF6753B036D639F6" docLanguage="en" docName="zootaxa.4661.2.5.pdf" docOrigin="Zootaxa 4661 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Haudrhytidaspis Ingrisch 2019, gen. n." docType="treatment" docVersion="10" lastPageNumber="369" masterDocId="FFF78032FFACFFF1FFF05745321C387D" masterDocTitle="Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae)" masterLastPageNumber="370" masterPageNumber="343" pageNumber="368" updateTime="1698753004269" updateUser="ExternalLinkService">
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<mods:title id="CAC6F218D2203AA983FAA9AFD36796CF">Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae)</mods:title>
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<mods:namePart id="6052179A6A0E091157BFAEF4F79D0E43">Ingrisch, Sigfrid</mods:namePart>
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<mods:date id="C8B15C652371C8FCA4FF244D4C7307ED">2019</mods:date>
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<treatment id="03CEF84AFFB5FFEBFF6753B036D639F6" ID-DOI="http://doi.org/10.5281/zenodo.5935393" ID-GBIF-Taxon="159293151" ID-Zenodo-Dep="5935393" LSID="urn:lsid:plazi:treatment:03CEF84AFFB5FFEBFF6753B036D639F6" httpUri="http://treatment.plazi.org/id/03CEF84AFFB5FFEBFF6753B036D639F6" lastPageId="26" lastPageNumber="369" pageId="25" pageNumber="368">
<subSubSection id="C37D1AD7FFB5FFE8FF6753B033A53D72" box="[151,441,1269,1296]" pageId="25" pageNumber="368" type="nomenclature">
<paragraph id="8BD8495CFFB5FFE8FF6753B033A53D72" blockId="25.[151,441,1269,1296]" box="[151,441,1269,1296]" pageId="25" pageNumber="368">
<heading id="D090FE30FFB5FFE8FF6753B033A53D72" bold="true" box="[151,441,1269,1296]" fontSize="11" level="1" pageId="25" pageNumber="368" reason="1">
<emphasis id="B913954EFFB5FFE8FF6753B033A53D72" bold="true" box="[151,441,1269,1296]" pageId="25" pageNumber="368">
<taxonomicName id="4C6732DFFFB5FFE8FF6753B033433D6D" ID-CoL="8PV3X" authority="Ingrisch, 2019" authorityName="Ingrisch" authorityYear="2019" box="[151,351,1269,1296]" class="Insecta" family="Tettigoniidae" genus="Haudrhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B913954EFFB5FFE8FF6753B033433D6D" bold="true" box="[151,351,1269,1296]" italics="true" pageId="25" pageNumber="368">Haudrhytidaspis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2202835FFB5FFE8FE9753B033A53D72" box="[359,441,1269,1295]" pageId="25" pageNumber="368" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37D1AD7FFB5FFE8FF67527F310E3D2C" box="[151,786,1336,1362]" pageId="25" pageNumber="368" type="type_taxon">
<paragraph id="8BD8495CFFB5FFE8FF67527F310E3D2C" blockId="25.[151,786,1336,1362]" box="[151,786,1336,1362]" pageId="25" pageNumber="368">
<typeStatus id="54DCF7FEFFB5FFE8FF67527F32D53D2F" box="[151,201,1338,1362]" pageId="25" pageNumber="368">Type</typeStatus>
species:
<taxonomicName id="4C6732DFFFB5FFE8FED6527C30443D2C" ID-CoL="8PJQ7" authority="Karny, 1911" authorityName="Karny" authorityYear="1911" box="[294,600,1336,1362]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B913954EFFB5FFE8FED6527C33C43D2C" box="[294,472,1337,1361]" italics="true" pageId="25" pageNumber="368">Rhytidaspis fusca</emphasis>
<bibRefCitation id="EFF634ADFFB5FFE8FE2F527F30443D2C" author="Karny, H. H." box="[479,600,1336,1362]" pageId="25" pageNumber="368" pagination="334 - 347" refId="ref18510" refString="Karny, H. H. (1911) Descriptiones Conocephalidarum novarum. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 61, 334 - 347." type="journal article" year="1911">Karny, 1911</bibRefCitation>
</taxonomicName>
—here designated
</paragraph>
</subSubSection>
<subSubSection id="C37D1AD7FFB5FFEBFF67523D36D639F6" lastPageId="26" lastPageNumber="369" pageId="25" pageNumber="368" type="diagnosis">
<paragraph id="8BD8495CFFB5FFE8FF67523D372E3F8A" blockId="25.[151,1437,1400,2039]" pageId="25" pageNumber="368">
<emphasis id="B913954EFFB5FFE8FF67523D330E3DEE" bold="true" box="[151,274,1400,1427]" pageId="25" pageNumber="368">Diagnosis.</emphasis>
The new genus differs from
<taxonomicName id="4C6732DFFFB5FFE8FD95523C30F43DEE" authorityName="Redtenbacher" authorityYear="1891" box="[613,744,1401,1427]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FD95523C30F43DEE" box="[613,744,1401,1427]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
by the following characters: The fastigium verticis is rather short, hardly surpassing scapus, triangular and dorsoventrally compressed instead of long and laterally compressed with narrow conical apical area; the face is robust and wide with the surface finely rugose with strong, irregular rugae and small impressed dots and gradually passes into genae (
<figureCitation id="135C55D9FFB5FFE8FC7152A031C03D82" box="[897,988,1509,1535]" captionStart="FIGURE 2" captionStartId="6.[151,250,1836,1861]" captionTargetBox="[151,1436,206,1765]" captionTargetId="figure@6.[151,1436,206,1765]" captionTargetPageId="6" captionText="FIGURE 2. Face of A–B, Rhytidaspis variata meja ssp. n. (A, male Tafelberg; B, female Dorey); C–E, R. arfak sp. n. (C, male Koffo; D, male Siwi; E, female Siwi); F, R. variata brevis ssp. n. male (Freeport); G, R. variata variata ssp. n. (male Nabire); H–J, R. picta Redt., 1891 (H, male Fakfak; I, female Arguni; J, male Bundi); K–M, R. ornata sp. n. (K, male South N.G.; L, male Mindiptana; M, male Tiel); N, R. camela sp. n. (male Sorong); O, R. nigropunctata sp. n. (male Mt. Nok); P, R. genyem sp. n. (female Genyem); Q, Haudrhytidaspis fusca (Karny, 1911) comb. nov. (female type). Scales 10 mm. The arrows point at the sub-ocular carina, present in Rhytidaspis, absent in Haudrhytidaspis." httpUri="https://zenodo.org/record/3379499/files/figure.png" pageId="25" pageNumber="368">Fig. 2Q</figureCitation>
), while In
<taxonomicName id="4C6732DFFFB5FFE8FBA952A036C03D82" authorityName="Redtenbacher" authorityYear="1891" box="[1113,1244,1509,1535]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FBA952A036C03D82" box="[1113,1244,1509,1535]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
it appears rather narrow, elongate with the surface covered with large impressed dots and the face is separated from the genae by strong sub-ocular carinae. The fore tibiae are markedly angular with the surface below the tympana coarsely rugose instead of nearly subsmooth and the hind femora carry thorns only on the external ventral margin instead of on both ventral margins as in
<taxonomicName id="4C6732DFFFB5FFE8FE32513030593EF2" authorityName="Redtenbacher" authorityYear="1891" box="[450,581,1653,1679]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FE32513030593EF2" box="[450,581,1653,1679]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
. The pronotum in
<taxonomicName id="4C6732DFFFB5FFE8FCE6513031CC3EF2" authority="Ingrisch, 2019" authorityName="Ingrisch" authorityYear="2019" box="[790,976,1653,1679]" class="Insecta" family="Tettigoniidae" genus="Haudrhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B913954EFFB5FFE8FCE6513031CC3EF2" box="[790,976,1653,1679]" italics="true" pageId="25" pageNumber="368">Haudrhytidaspis</emphasis>
</taxonomicName>
is short with only one transverse furrow, has the ventral margin convex and the humeral sinus only weakly indicated (
<figureCitation id="135C55D9FFB5FFE8FBF451DC36773ECE" box="[1028,1131,1689,1715]" captionStart="FIGURE 11" captionStartId="22.[151,250,1796,1821]" captionTargetBox="[179,1408,166,1769]" captionTargetId="figure@22.[178,1410,166,1776]" captionTargetPageId="22" captionText="FIGURE 11. Female subgenital plate in ventral (A–H, X), lateral (I–O, W) and oblique apical view (P–V); thorax in lateral view (Y); apex of abdomen with ovipositor and hind leg in lateral view (Z); faint serration of ventral ovipositor valves before tip (Z insert).—Rhytidaspis arfak sp. n. from Koffo (A, I, P) and Siwi (E, M, T, W); R. picta Redt., 1891 Arguni (B, N, S); R. variata meja ssp. n. Pami (C, J, Q) and Dorey (F); R. ornata sp. n. South N.G. (D) and Kloofbivak (K, R); R. genyem sp. n. (G, L, V); R. variata ssp.Aindua (H, O, U); Haudrhytidaspis fusca (Karny, 1911) (X–Z).—Abbreviations: ls lateral area, vs ventral area of subgenital plate, ov base of ventral ovipositor valves. The black arrows in B and N mark the up-curved projections of the ventral surface, the white arrow in L the widened base, in Y the tegmen. Scales 1 mm; X–Z not to scale." httpUri="https://zenodo.org/record/3379521/files/figure.png" pageId="25" pageNumber="368">Fig. 11Y</figureCitation>
), while in
<taxonomicName id="4C6732DFFFB5FFE8FB1651DC37753ECE" authorityName="Redtenbacher" authorityYear="1891" box="[1254,1385,1689,1715]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FB1651DC37753ECE" box="[1254,1385,1689,1715]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
it is elongate, with concave ventral margin, two transverse furrows, and a distinct humeral sinus. The female subgenital plate is short in the new genus, hardly covering the base of the ovipositor, funnel-shaped with gradually up-curved lateral margins and at the apical margin there is only a weak incision in middle (
<figureCitation id="135C55D9FFB5FFE8FBE5504036673F62" box="[1045,1147,1797,1823]" captionStart="FIGURE 11" captionStartId="22.[151,250,1796,1821]" captionTargetBox="[179,1408,166,1769]" captionTargetId="figure@22.[178,1410,166,1776]" captionTargetPageId="22" captionText="FIGURE 11. Female subgenital plate in ventral (A–H, X), lateral (I–O, W) and oblique apical view (P–V); thorax in lateral view (Y); apex of abdomen with ovipositor and hind leg in lateral view (Z); faint serration of ventral ovipositor valves before tip (Z insert).—Rhytidaspis arfak sp. n. from Koffo (A, I, P) and Siwi (E, M, T, W); R. picta Redt., 1891 Arguni (B, N, S); R. variata meja ssp. n. Pami (C, J, Q) and Dorey (F); R. ornata sp. n. South N.G. (D) and Kloofbivak (K, R); R. genyem sp. n. (G, L, V); R. variata ssp.Aindua (H, O, U); Haudrhytidaspis fusca (Karny, 1911) (X–Z).—Abbreviations: ls lateral area, vs ventral area of subgenital plate, ov base of ventral ovipositor valves. The black arrows in B and N mark the up-curved projections of the ventral surface, the white arrow in L the widened base, in Y the tegmen. Scales 1 mm; X–Z not to scale." httpUri="https://zenodo.org/record/3379521/files/figure.png" pageId="25" pageNumber="368">
Fig.
<date id="FFD96F9CFFB5FFE8FBB9504036673F62" box="[1097,1147,1797,1823]" pageId="25" pageNumber="368" value="2011-10">11X</date>
</figureCitation>
), while in
<taxonomicName id="4C6732DFFFB5FFE8FB01504037683F62" authorityName="Redtenbacher" authorityYear="1891" box="[1265,1396,1797,1823]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FB01504037683F62" box="[1265,1396,1797,1823]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
the subgenital plate has the ventral surface elongate triangular with bifid tip while the lateral areas are almost vertically up-bent and for a large part membranous. The very base of the ventral ovipositor valves starts at base with a narrow transverse bulge (
<figureCitation id="135C55D9FFB5FFE8FEAD503433DD3FF6" box="[349,449,1905,1931]" captionStart="FIGURE 11" captionStartId="22.[151,250,1796,1821]" captionTargetBox="[179,1408,166,1769]" captionTargetId="figure@22.[178,1410,166,1776]" captionTargetPageId="22" captionText="FIGURE 11. Female subgenital plate in ventral (A–H, X), lateral (I–O, W) and oblique apical view (P–V); thorax in lateral view (Y); apex of abdomen with ovipositor and hind leg in lateral view (Z); faint serration of ventral ovipositor valves before tip (Z insert).—Rhytidaspis arfak sp. n. from Koffo (A, I, P) and Siwi (E, M, T, W); R. picta Redt., 1891 Arguni (B, N, S); R. variata meja ssp. n. Pami (C, J, Q) and Dorey (F); R. ornata sp. n. South N.G. (D) and Kloofbivak (K, R); R. genyem sp. n. (G, L, V); R. variata ssp.Aindua (H, O, U); Haudrhytidaspis fusca (Karny, 1911) (X–Z).—Abbreviations: ls lateral area, vs ventral area of subgenital plate, ov base of ventral ovipositor valves. The black arrows in B and N mark the up-curved projections of the ventral surface, the white arrow in L the widened base, in Y the tegmen. Scales 1 mm; X–Z not to scale." httpUri="https://zenodo.org/record/3379521/files/figure.png" pageId="25" pageNumber="368">
Fig.
<date id="FFD96F9CFFB5FFE8FE61503433DD3FF6" box="[401,449,1905,1931]" pageId="25" pageNumber="368" value="2011-10">11X</date>
</figureCitation>
) while in
<taxonomicName id="4C6732DFFFB5FFE8FDC1503430A83FF6" authorityName="Redtenbacher" authorityYear="1891" box="[561,692,1905,1931]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FDC1503430A83FF6" box="[561,692,1905,1931]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
there are long, swollen extensions at base of the ventral ovipositor valves. The ovipositor in the new genus is slightly but regularly curved throughout its length and the ventral valves are in apical area of ventral valves faintly serrated (
<figureCitation id="135C55D9FFB5FFE8FD2850FC31233FAE" box="[728,831,1977,2003]" captionStart="FIGURE 11" captionStartId="22.[151,250,1796,1821]" captionTargetBox="[179,1408,166,1769]" captionTargetId="figure@22.[178,1410,166,1776]" captionTargetPageId="22" captionText="FIGURE 11. Female subgenital plate in ventral (A–H, X), lateral (I–O, W) and oblique apical view (P–V); thorax in lateral view (Y); apex of abdomen with ovipositor and hind leg in lateral view (Z); faint serration of ventral ovipositor valves before tip (Z insert).—Rhytidaspis arfak sp. n. from Koffo (A, I, P) and Siwi (E, M, T, W); R. picta Redt., 1891 Arguni (B, N, S); R. variata meja ssp. n. Pami (C, J, Q) and Dorey (F); R. ornata sp. n. South N.G. (D) and Kloofbivak (K, R); R. genyem sp. n. (G, L, V); R. variata ssp.Aindua (H, O, U); Haudrhytidaspis fusca (Karny, 1911) (X–Z).—Abbreviations: ls lateral area, vs ventral area of subgenital plate, ov base of ventral ovipositor valves. The black arrows in B and N mark the up-curved projections of the ventral surface, the white arrow in L the widened base, in Y the tegmen. Scales 1 mm; X–Z not to scale." httpUri="https://zenodo.org/record/3379521/files/figure.png" pageId="25" pageNumber="368">
Fig.
<date id="FFD96F9CFFB5FFE8FCFC50FC31233FAE" box="[780,831,1977,2003]" pageId="25" pageNumber="368" value="2011-10">11X</date>
</figureCitation>
), while in
<taxonomicName id="4C6732DFFFB5FFE8FC4550FC36243FAE" authorityName="Redtenbacher" authorityYear="1891" box="[949,1080,1977,2003]" class="Insecta" family="Tettigoniidae" genus="Rhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB5FFE8FC4550FC36243FAE" box="[949,1080,1977,2003]" italics="true" pageId="25" pageNumber="368">Rhytidaspis</emphasis>
</taxonomicName>
the ovipositor is sub-straight in about basal half and slightly curved in apical half and the margin of the ventral valves is smooth until tip.
</paragraph>
<paragraph id="8BD8495CFFB6FFEBFF3757DC36D639F6" blockId="26.[151,1436,153,395]" pageId="26" pageNumber="369">
The outline of the robust head of
<taxonomicName id="4C6732DFFFB6FFEBFDCF57DC30E538CE" authority="Ingrisch, 2019" authorityName="Ingrisch" authorityYear="2019" box="[575,761,153,179]" class="Insecta" family="Tettigoniidae" genus="Haudrhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B913954EFFB6FFEBFDCF57DC30E538CE" box="[575,761,153,179]" italics="true" pageId="26" pageNumber="369">Haudrhytidaspis</emphasis>
</taxonomicName>
with a short fastigium verticis, the sculpture of the face, the short meso- and metasternal lobes, and the angular fore tibiae with short spines resemble some species of the genus
<taxonomicName id="4C6732DFFFB6FFEBFF6757A433D73886" authority="Blanchard, 1853" authorityName="Blanchard" authorityYear="1853" box="[151,459,224,251]" class="Insecta" family="Tettigoniidae" genus="Salomona" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B913954EFFB6FFEBFF6757A4331B3886" box="[151,263,225,251]" italics="true" pageId="26" pageNumber="369">Salomona</emphasis>
Blanchard, 1853
</taxonomicName>
.
<taxonomicName id="4C6732DFFFB6FFEBFE2657A4308C3886" authority="Ingrisch, 2019" authorityName="Ingrisch" authorityYear="2019" box="[470,656,225,251]" class="Insecta" family="Tettigoniidae" genus="Haudrhytidaspis" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B913954EFFB6FFEBFE2657A4308C3886" box="[470,656,225,251]" italics="true" pageId="26" pageNumber="369">Haudrhytidaspis</emphasis>
</taxonomicName>
differs however strikingly by the low, rounded pronotum with only a faint indication of a humeral sinus and a single interrupted transverse sulcus, the sub-straight, only faintly curved ovipositor and the strongly reduced wings speak against any affinity with that genus. The short rectangular female subgenital plate that hardly covers the base of the ovipositor is also of diagnostic value. The combination of characters listed before differentiates the new genus also from other genera of Australasian
<taxonomicName id="4C6732DFFFB6FFEBFBBC563436DA39F6" box="[1100,1222,369,395]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="369" phylum="Arthropoda" rank="tribe" tribe="Agraeciini">Agraeciini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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