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<document id="1DE40ECB8C4374D6C5C5224F6EA11FAC" ID="10.11646/zootaxa.4150.3.2" ID-DOI="10.11646/zootaxa.4150.3.2" ID-GBIF-Dataset="8347a890-caad-4a81-8a02-8ec476083ffb" ID-ISSN="1175-5326" ID-Zenodo-Dep="272101" ID-ZooBank="08E63512-49DB-495C-83FC-CDF206A516CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470983924873" checkinUser="plazi" docAuthor="Gordon, Cat A., Hood, Ali R. &amp; Ellis, Jim R." docDate="2016" docId="03C2E354FFC9FFC2FF49F9DEFA865E05" docLanguage="en" docName="zootaxa.4150.3.2.pdf" docOrigin="Zootaxa 4150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucoraja naevus Muller &amp; Henle 1841" docType="treatment" docVersion="9" lastPageNumber="263" masterDocId="FFFB9B2CFFCEFFCAFFDEFFB5FFF25D59" masterDocTitle="Descriptions and revised key to the eggcases of the skates (Rajiformes: Rajidae) and catsharks (Carcharhiniformes: Scyliorhinidae) of the British Isles" masterLastPageNumber="280" masterPageNumber="255" pageNumber="262" updateTime="1698381824525" updateUser="plazi">
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<mods:title id="97165CBA8E605FFEA39F7C5CC11278BF">Descriptions and revised key to the eggcases of the skates (Rajiformes: Rajidae) and catsharks (Carcharhiniformes: Scyliorhinidae) of the British Isles</mods:title>
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<mods:namePart id="B639D854D0F8BC2BBE4B6EA56B7BA55D">Gordon, Cat A.</mods:namePart>
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<treatment id="03C2E354FFC9FFC2FF49F9DEFA865E05" ID-DOI="http://doi.org/10.5281/zenodo.5631360" ID-GBIF-Taxon="124329263" ID-Zenodo-Dep="5631360" LSID="urn:lsid:plazi:treatment:03C2E354FFC9FFC2FF49F9DEFA865E05" httpUri="http://treatment.plazi.org/id/03C2E354FFC9FFC2FF49F9DEFA865E05" lastPageId="8" lastPageNumber="263" pageId="7" pageNumber="262">
<subSubSection id="C37101C9FFC9FFCDFF49F9DEFD6B5BDC" box="[151,665,1643,1669]" pageId="7" pageNumber="262" type="nomenclature">
<paragraph id="8BD45242FFC9FFCDFF49F9DEFD6B5BDC" blockId="7.[151,665,1643,1669]" box="[151,665,1643,1669]" pageId="7" pageNumber="262">
<heading id="D09CE52EFFC9FFCDFF49F9DEFD6B5BDC" bold="true" box="[151,665,1643,1669]" fontSize="11" level="1" pageId="7" pageNumber="262" reason="1">
<taxonomicName id="4C6B29C1FFC9FFCDFF49F9DEFD6B5BDC" ID-CoL="725XK" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[151,665,1643,1669]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="262" phylum="Chordata" rank="species" species="naevus">
<emphasis id="B91F8E50FFC9FFCDFF49F9DEFD6B5BDC" bold="true" box="[151,665,1643,1669]" pageId="7" pageNumber="262">
<emphasis id="B91F8E50FFC9FFCDFF49F9DEFE9D5BDC" bold="true" box="[151,367,1643,1669]" italics="true" pageId="7" pageNumber="262">Leucoraja naevus</emphasis>
(Müller &amp; Henle, 1841)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37101C9FFC9FFCDFF49F907FB105A02" pageId="7" pageNumber="262" type="distribution">
<paragraph id="8BD45242FFC9FFCDFF49F907FB105A02" blockId="7.[151,1436,1714,2027]" pageId="7" pageNumber="262">
<emphasis id="B91F8E50FFC9FFCDFF49F907FEC25B92" bold="true" box="[151,304,1714,1739]" pageId="7" pageNumber="262">Distribution.</emphasis>
Cuckoo ray (
<taxonomicName id="4C6B29C1FFC9FFCDFE12F906FDCB5B93" box="[460,569,1715,1738]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="262" phylum="Chordata" rank="species" species="naevus">
<emphasis id="B91F8E50FFC9FFCDFE12F906FDCB5B93" box="[460,569,1715,1738]" italics="true" pageId="7" pageNumber="262">L. naevus</emphasis>
</taxonomicName>
) is relatively widespread in the Northeast Atlantic, occurring from
<collectingCountry id="F37C12D2FFC9FFCDFAE9F906FA6E5B93" box="[1335,1436,1715,1738]" name="Morocco" pageId="7" pageNumber="262">Morocco</collectingCountry>
to
<collectingCountry id="F37C12D2FFC9FFCDFF6AF962FEFD5BB6" box="[180,271,1751,1775]" name="Norway" pageId="7" pageNumber="262">Norway</collectingCountry>
, including the Mediterranean Sea (
<bibRefCitation id="EFFA2FB3FFC9FFCDFD44F960FC3A5BB7" author="Stehmann" box="[666,968,1749,1774]" pageId="7" pageNumber="262" refString="Stehmann, M. F. W. &amp; Burkel D. L. (1984). Rajidae. In: Whitehead P. J. P., Bauchot, M. - L., Hureaun, J. - C., Nielsen, J. &amp; Tortonese, E. (Eds) Fishes of the North-eastern Atlantic and the Mediterranean. Vol. I. pp. 163 - 196. UNESCO, Paris, 163 - 196." type="book chapter" year="1984">Stehmann &amp; Bürkel, 1984</bibRefCitation>
;
<bibRefCitation id="EFFA2FB3FFC9FFCDFC0AF963FB5F5BB7" author="Serena" box="[980,1197,1750,1774]" pageId="7" pageNumber="262" refString="Serena, F., Mancusi, C. &amp; Barone, M. (2010) Field identification guide to the skates (Rajidae) of the Mediterranean Sea. Guidelines for data collection and analysis. Biologia Marina Mediterranea, 17 (Supplement 2), 204 pp." type="book" year="2010">
Serena
<emphasis id="B91F8E50FFC9FFCDFBF6F962FB9A5BB7" box="[1064,1128,1750,1774]" italics="true" pageId="7" pageNumber="262">et al.,</emphasis>
2010
</bibRefCitation>
). Juveniles are found further offshore than the young of
<taxonomicName id="4C6B29C1FFC9FFCDFDC6F94EFDBF5A4B" box="[536,589,1787,1810]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="262" phylum="Chordata" rank="genus">
<emphasis id="B91F8E50FFC9FFCDFDC6F94EFDBF5A4B" box="[536,589,1787,1810]" italics="true" pageId="7" pageNumber="262">Raja</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="EFFA2FB3FFC9FFCDFD4EF94FFCBC5A4A" author="Ellis" box="[656,846,1786,1811]" pageId="7" pageNumber="262" refString="Ellis, J. R., Cruz-Martinez, A., Rackham, B. D. &amp; Rogers, S. I. (2005) The distribution of chondrichthyan fishes around the British Isles and implications for conservation. Journal of Northwest Atlantic Fishery Science, 35, 195 - 213. http: // dx. doi. org / 10.2960 / J. v 35. m 485" type="journal article" year="2005">
Ellis
<emphasis id="B91F8E50FFC9FFCDFD12F94EFCF95A4B" box="[716,779,1786,1810]" italics="true" pageId="7" pageNumber="262">et al.,</emphasis>
2005
</bibRefCitation>
). Records submitted to the Great Eggcase Hunt were collected predominantly from southwest
<collectingCountry id="F37C12D2FFC9FFCDFDBEF8A8FD325A6F" box="[608,704,1821,1846]" name="United Kingdom" pageId="7" pageNumber="262">England</collectingCountry>
, although verified reports have been made along the southern and western coasts of
<collectingCountry id="F37C12D2FFC9FFCDFE81F8F7FE4E5A02" box="[351,444,1858,1883]" name="United Kingdom" pageId="7" pageNumber="262">England</collectingCountry>
and
<collectingCountry id="F37C12D2FFC9FFCDFE2CF8F7FDCB5A02" box="[498,569,1858,1883]" name="United Kingdom" pageId="7" pageNumber="262">Wales</collectingCountry>
, with scattered reports from
<collectingCountry id="F37C12D2FFC9FFCDFCA4F8F7FC2E5A02" box="[890,988,1858,1883]" name="United Kingdom" pageId="7" pageNumber="262">Scotland</collectingCountry>
and the Northern Isles.
</paragraph>
</subSubSection>
<subSubSection id="C37101C9FFC9FFCDFF19F8D0FAE95AB2" pageId="7" pageNumber="262" type="materials_examined">
<paragraph id="8BD45242FFC9FFCDFF19F8D0FAE95AB2" blockId="7.[151,1436,1714,2027]" pageId="7" pageNumber="262">
<materialsCitation id="3B03581FFFC9FFCDFF19F8D0FC9B5AB2" ID-GBIF-Occurrence="1291551000" collectionCode="BMNH" latitude="50.9" location="Specimens" municipality="western English Channel" pageId="7" pageNumber="262" specimenCount="1">
<emphasis id="B91F8E50FFC9FFCDFF19F8D0FE405A27" bold="true" box="[199,434,1893,1918]" pageId="7" pageNumber="262">Material examined.</emphasis>
Ninety-four specimens were examined overall, including those recorded from the Great Eggcase Hunt (n = 52) and trawl surveys (n = 39), with three museum specimens also examined (
<collectionCode id="ED7ACA87FFC9FFCDFA98F83FFA6E5AFB" box="[1350,1436,1930,1954]" pageId="7" pageNumber="262">BMNH</collectionCode>
1927.1.31.13).
<location id="8EB40499FFC9FFCDFE8AF818FE3F5A9F" LSID="urn:lsid:plazi:treatment:03C2E354FFC9FFC2FF49F9DEFA865E05:8EB40499FFC9FFCDFE8AF818FE3F5A9F" box="[340,461,1965,1990]" latitude="50.9" municipality="western English Channel" name="Specimens" pageId="7" pageNumber="262">Specimens</location>
from trawl surveys were captured primarily by beam trawl in the
<collectingMunicipality id="6BB0C838FFC9FFCDFB3AF81AFF065AB2" pageId="7" pageNumber="262">western English Channel</collectingMunicipality>
and Celtic Sea, over the area from 49.1
<geoCoordinate id="EE5F3485FFC9FFCDFD1EF867FCE25AB2" box="[704,784,2002,2027]" direction="north" orientation="latitude" pageId="7" pageNumber="262" precision="5555" value="50.9">50.9°N</geoCoordinate>
and 3.9
</materialsCitation>
6.7°W, and in waters
<quantity id="4C93FFA7FFC9FFCDFBB9F867FB255AB3" box="[1127,1239,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="1.01" metricValueMax="1.18" metricValueMin="0.84" pageId="7" pageNumber="262" unit="m" value="101.0" valueMax="118.0" valueMin="84.0">84118 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="C37101C9FFC6FFC2FF19FF22FD425CF4" pageId="8" pageNumber="263" type="description">
<paragraph id="8BD45242FFC6FFC2FF19FF22FD425CF4" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="263">
<emphasis id="B91F8E50FFC6FFC2FF19FF22FEAB5DE9" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="263">Description.</emphasis>
The eggcase of
<taxonomicName id="4C6B29C1FFC6FFC2FDC1FF2CFD7D5DE9" box="[543,655,153,176]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="naevus">
<emphasis id="B91F8E50FFC6FFC2FDC1FF2CFD7D5DE9" box="[543,655,153,176]" italics="true" pageId="8" pageNumber="263">L. naevus</emphasis>
</taxonomicName>
is small (eggcase length 50.0 ±
<quantity id="4C93FFA7FFC6FFC2FBCDFF2DFB8C5DF6" box="[1043,1150,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="8" pageNumber="263" unit="mm" value="13.2">13.2 mm</quantity>
;
<quantity id="4C93FFA7FFC6FFC2FB53FF2DFAC75DE9" box="[1165,1333,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.84" metricValueMax="6.52" metricValueMin="3.16" pageId="8" pageNumber="263" unit="mm" value="48.4" valueMax="65.2" valueMin="31.6">31.665.2 mm</quantity>
) but has extended horns (
<figureCitation id="13504EC7FFC6FFC2FE83FF09FE375D8D" box="[349,453,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1078,1100]" captionTargetBox="[311,1269,197,1049]" captionTargetId="figure@4.[300,1287,193,1056]" captionTargetPageId="4" captionText="FIGURE 2. Eggcases of (a) Amblyraja radiata (large specimen from Borth, Cardigan Bay), (b) A. radiata (typical specimen from North Sea) and (c) Leucoraja naevus." httpUri="https://zenodo.org/record/272103/files/figure.png" pageId="8" pageNumber="263">Figure 2</figureCitation>
c), and the range of total eggcase length is
<quantity id="4C93FFA7FFC6FFC2FC02FF09FB565D8D" box="[988,1188,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.7735" metricValueMax="1.9789999999999999" metricValueMin="1.568" pageId="8" pageNumber="263" unit="mm" value="177.35" valueMax="197.9" valueMin="156.8">156.8197.9 mm</quantity>
. The three historical specimens examined closely matched the size range recorded in contemporary samples. Both dorsal and ventral surfaces are convex, with obvious longitudinal striations on both surfaces. The eggcase is without keels and the dorsal and ventral surfaces meet seamlessly at the margin. The anterior apron is pronounced and convex whereas the posterior apron is sometimes lacking. The anterior horns are long (on average representing 48% of the entire eggcase length) and in complete specimens curve inwards and often overlap. The posterior horns are shorter than the capsule length and often terminate in hooks.
</paragraph>
</subSubSection>
<subSubSection id="C37101C9FFC6FFC2FF19FE02FA865E05" pageId="8" pageNumber="263" type="discussion">
<paragraph id="8BD45242FFC6FFC2FF19FE02FC385F39" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="263">
<emphasis id="B91F8E50FFC6FFC2FF19FE02FEC95C89" bold="true" box="[199,315,439,464]" pageId="8" pageNumber="263">Remarks.</emphasis>
As the anterior horns are so long and delicate, they are often broken in strandline specimens, which can sometimes cause confusion with
<taxonomicName id="4C6B29C1FFC6FFC2FDE6FE68FC8B5CAD" authority="Fowler, 1910" authorityName="Fowler" authorityYear="1910" box="[568,889,476,501]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="montagui">
<emphasis id="B91F8E50FFC6FFC2FDE6FE68FD2C5CAD" box="[568,734,477,500]" italics="true" pageId="8" pageNumber="263">Raja montagui</emphasis>
Fowler, 1910
</taxonomicName>
when attempting identification. However, as the capsule is biconvex and has a more rounded appearance, it is still distinct from the other species found in European inshore waters. The mean eggcase width in the present study was within the range of previous reports (
<tableCitation id="C6E967F9FFC6FFC2FAD3FD91FA925F65" box="[1293,1376,548,573]" captionStart="TABLE 1" captionStartId="10.[119,204,553,575]" captionTargetBox="[119,1868,631,1415]" captionTargetPageId="10" captionText="TABLE 1. (Continueđ)" httpUri="http://table.plazi.org/id/DF1402CAFFC4FFC0FFA9FD9CFE975F66" pageId="8" pageNumber="263" tableUuid="DF1402CAFFC4FFC0FFA9FD9CFE975F66">Table 1</tableCitation>
), but mean eggcase length was slightly shorter than recorded in earlier studies.
</paragraph>
<paragraph id="8BD45242FFC6FFC2FF19FDD8FA865E05" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="263">
No comparative material for the related sandy ray
<taxonomicName id="4C6B29C1FFC6FFC2FCF9FDD8FB355FDD" authority="Couch, 1838" authorityName="Couch" authorityYear="1838" box="[807,1223,620,645]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="circularis">
<emphasis id="B91F8E50FFC6FFC2FCF9FDD8FBE15FDD" box="[807,1043,620,644]" italics="true" pageId="8" pageNumber="263">Leucoraja circularis</emphasis>
(Couch, 1838)
</taxonomicName>
and shagreen ray
<taxonomicName id="4C6B29C1FFC6FFC2FF49FD25FDB55FF1" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,583,655,680]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="fullonica">
<emphasis id="B91F8E50FFC6FFC2FF49FD25FE8B5FF1" box="[151,377,656,680]" italics="true" pageId="8" pageNumber="263">Leucoraja fullonica</emphasis>
(Linnaeus, 1758)
</taxonomicName>
were available during this study. Some of the earlier descriptions of the eggcases of what was purported to be
<taxonomicName id="4C6B29C1FFC6FFC2FD82FD00FD1B5F95" box="[604,745,692,716]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="circularis">
<emphasis id="B91F8E50FFC6FFC2FD82FD00FD1B5F95" box="[604,745,692,716]" italics="true" pageId="8" pageNumber="263">L. circularis</emphasis>
</taxonomicName>
should be treated with care, as cuckoo ray was sometimes referred to as
<taxonomicName id="4C6B29C1FFC6FFC2FEEAFD6DFE2F5FA9" box="[308,477,728,752]" class="Elasmobranchii" family="Rajidae" genus="Raia" higherTaxonomySource="CoL" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="circularis">
<emphasis id="B91F8E50FFC6FFC2FEEAFD6DFE2F5FA9" box="[308,477,728,752]" italics="true" pageId="8" pageNumber="263">Raia circularis</emphasis>
</taxonomicName>
in early accounts, as noted by
<bibRefCitation id="EFFA2FB3FFC6FFC2FCE3FD62FC225FA9" author="Clark" box="[829,976,727,752]" pageId="8" pageNumber="263" refString="Clark, R. S. (1922) Rays and Skates (Raiae) No. I: Egg capsules and young. Journal of the Marine Biological Association of the United Kingdom, 12, 577 - 643. http: // dx. doi. org / 10.1017 / S 002531540000967 X" type="journal article" year="1922">Clark (1922)</bibRefCitation>
. Both
<taxonomicName id="4C6B29C1FFC6FFC2FBC5FD6CFB575FA9" box="[1051,1189,728,752]" class="Elasmobranchii" family="Rajidae" genus="Raia" higherTaxonomySource="CoL" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="circularis">
<emphasis id="B91F8E50FFC6FFC2FBC5FD6CFB575FA9" box="[1051,1189,728,752]" italics="true" pageId="8" pageNumber="263">L. circularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B29C1FFC6FFC2FB03FD6CFA925FA9" box="[1245,1376,728,752]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="fullonica">
<emphasis id="B91F8E50FFC6FFC2FB03FD6CFA925FA9" box="[1245,1376,728,752]" italics="true" pageId="8" pageNumber="263">L. fullonica</emphasis>
</taxonomicName>
have eggcases of similar shape to
<taxonomicName id="4C6B29C1FFC6FFC2FE06FD48FDB15E4D" box="[472,579,765,788]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="naevus">
<emphasis id="B91F8E50FFC6FFC2FE06FD48FDB15E4D" box="[472,579,765,788]" italics="true" pageId="8" pageNumber="263">L. naevus</emphasis>
</taxonomicName>
, with preliminary descriptions given by
<bibRefCitation id="EFFA2FB3FFC6FFC2FBD4FD49FB2F5E4D" author="Desbrosses" box="[1034,1245,764,789]" pageId="8" pageNumber="263" refString="Desbrosses, P. (1931) Note sur l'oeuf de la raie chardon Raja fullonica L. Bulletin de la Societe Zoologique de France, 1931, 15 - 18." type="journal article" year="1931">Desbrosses (1931)</bibRefCitation>
. The eggcases of these two species are both larger than the cuckoo ray, ranging from
<quantity id="4C93FFA7FFC6FFC2FC70FC95FBD85E6E" box="[942,1066,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="9.2" metricValueMin="8.3" pageId="8" pageNumber="263" unit="mm" value="87.5" valueMax="92.0" valueMin="83.0">8392 mm</quantity>
long and
<quantity id="4C93FFA7FFC6FFC2FB7FFCAAFAEF5E6E" box="[1185,1309,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="8" pageNumber="263" unit="mm" value="49.0" valueMax="53.0" valueMin="45.0">4553 mm</quantity>
wide in
<taxonomicName id="4C6B29C1FFC6FFC2FA5BFC94FEF15E05" class="Elasmobranchii" family="Rajidae" genus="Raia" higherTaxonomySource="CoL" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="circularis">
<emphasis id="B91F8E50FFC6FFC2FA5BFC94FEF15E05" italics="true" pageId="8" pageNumber="263">L. circularis</emphasis>
</taxonomicName>
and
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long and
<quantity id="4C93FFA7FFC6FFC2FDFFFCF1FD685E05" box="[545,666,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="6.7" metricValueMin="4.5" pageId="8" pageNumber="263" unit="mm" value="56.0" valueMax="67.0" valueMin="45.0">4567 mm</quantity>
wide in
<taxonomicName id="4C6B29C1FFC6FFC2FD25FCF0FC8E5E05" box="[763,892,836,860]" class="Elasmobranchii" family="Rajidae" genus="Leucoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="263" phylum="Chordata" rank="species" species="fullonica">
<emphasis id="B91F8E50FFC6FFC2FD25FCF0FC8E5E05" box="[763,892,836,860]" italics="true" pageId="8" pageNumber="263">L. fullonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFA2FB3FFC6FFC2FC54FCF1FBA15E05" author="Desbrosses" box="[906,1107,836,861]" pageId="8" pageNumber="263" refString="Desbrosses, P. (1931) Note sur l'oeuf de la raie chardon Raja fullonica L. Bulletin de la Societe Zoologique de France, 1931, 15 - 18." type="journal article" year="1931">Desbrosses, 1931</bibRefCitation>
;
<bibRefCitation id="EFFA2FB3FFC6FFC2FB81FCF1FAF45E04" author="Wheeler" box="[1119,1286,836,861]" pageId="8" pageNumber="263" refString="Wheeler, A. (1969) The fishes of the British Isles and North-west Europe. Michigan State University Press, 613 pp." type="book" year="1969">Wheeler, 1969</bibRefCitation>
;
<tableCitation id="C6E967F9FFC6FFC2FACFFCF1FA915E05" box="[1297,1379,836,861]" captionStart="TABLE 1" captionStartId="10.[119,204,553,575]" captionTargetBox="[119,1868,631,1415]" captionTargetPageId="10" captionText="TABLE 1. (Continueđ)" httpUri="http://table.plazi.org/id/DF1402CAFFC4FFC0FFA9FD9CFE975F66" pageId="8" pageNumber="263" tableUuid="DF1402CAFFC4FFC0FFA9FD9CFE975F66">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>