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<document id="28D60290F00D106E5F14E42E3C1B28E9" ID-ISSN="0024-4082" ID-ZooBank="A0C63287-87B3-42E0-BC0E-CACDE62A8137" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630092563143" checkinUser="felipe" docAuthor="Ge, Deyan, Feijó, Anderson, Abramov, Alexei V., Wen, Zhixin, Liu, Zhengjia, Cheng, Jilong, Xia, Lin, Lu, Liang &amp; Yang, Qisen" docDate="2021" docId="03C28792E560FFBEFCF2F9C1FC5157C4" docLanguage="en" docName="zlaa040.pdf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Niviventer fulvescens" docType="treatment" docVersion="2" masterDocId="FFFBFFEAE56DFFB0FF8CFF8AFFA2573E" masterDocTitle="Molecular phylogeny and morphological diversity of the Niviventer fulvescens species complex with emphasis on species from China" updateTime="1699142589269" updateUser="plazi">
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<mods:title id="89D182D41ED50A42DDC9AAEBF5B6DD99">Molecular phylogeny and morphological diversity of the Niviventer fulvescens species complex with emphasis on species from China</mods:title>
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<mods:namePart id="734C5166BC1FF3F77C8B417C9C0B7D94">Ge, Deyan</mods:namePart>
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<mods:namePart id="927DBD53403B142C36EA6291A42D8678">Feijó, Anderson</mods:namePart>
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<mods:namePart id="E3FF6C6598C3D15476BBB8601A88912C">Abramov, Alexei V.</mods:namePart>
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<mods:namePart id="0C51310730978EBF5E5FDAC0BC4573E7">Wen, Zhixin</mods:namePart>
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<mods:namePart id="EDAA4D4243BE3C70B130804D6EC4BD18">Liu, Zhengjia</mods:namePart>
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<mods:namePart id="EABD5EC4283837EE38E6E884FC198F60">Cheng, Jilong</mods:namePart>
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<mods:namePart id="568373DD0BDCFE132806FEEB94835E81">Xia, Lin</mods:namePart>
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<mods:namePart id="9108C985A7342DE0F949B73194E60D51">Lu, Liang</mods:namePart>
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<mods:namePart id="5134B25C65535C504C06224E1245ECA9">Yang, Qisen</mods:namePart>
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<mods:title id="9B9226AB80AF39293D9DBCA7AA411B5B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6D0E20A4963D8764C4631570B58362B7">2021</mods:date>
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<subSubSection id="C371650FE560FFBDFCF2F9C1FA9E515D" box="[894,1340,1610,1635]" pageId="13" type="nomenclature">
<paragraph id="8BD43684E560FFBDFCF2F9C1FA9E515D" blockId="13.[894,1340,1610,1635]" box="[894,1340,1610,1635]" pageId="13">
<heading id="D09C81E8E560FFBDFCF2F9C1FA9E515D" box="[894,1340,1610,1635]" centered="true" fontSize="9" level="2" pageId="13" reason="2">
<taxonomicName id="4C6B4D07E560FFBDFCF2F9C1FB02515C" ID-CoL="47K8S" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[894,1184,1611,1634]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="13" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B91FEA96E560FFBDFCF2F9C1FB02515C" box="[894,1184,1611,1634]" italics="true" pageId="13">NIVIVENTER FULVESCENS</emphasis>
</taxonomicName>
(GRAY, 1847)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C371650FE560FFBDFB95F90CFAD651FA" pageId="13" type="description">
<paragraph id="8BD43684E560FFBDFB95F90CFB0251A1" blockId="13.[1049,1184,1670,1695]" box="[1049,1184,1670,1695]" pageId="13">
(
<figureCitation id="13502A01E560FFBDFBADF90CFBDF51A1" box="[1057,1149,1670,1695]" captionStart="Figure 5" captionStartId="10.[163,244,1146,1168]" captionTargetBox="[165,1443,197,1105]" captionTargetId="figure-369@10.[163,1446,195,1107]" captionTargetPageId="10" captionText="Figure 5. Comparison of the skin and skull of N. fengi with closely related species. Niviventer fengi, A1, A2, IOZCAS JL17111; A3, A4, IOZCASJL17059, HBL = 146. Niviventer fulvescens, B1, B2, IOZCAS JD113; B3, B4, JL17057, HBL = 120. Niviventer huang, C1, C2, NHM 98.11.1.16; C3, C4, IOZCAS HN2017242, HBL = 125. Niviventer mekongis, D1, D2, NHM 26.11.17.13; D3, D4, ZIN102439, HBL = 134. Niviventer cremoriventer, E1, E2, KMIOZCAS; E3, E4. KMIOZCAS, HBL = 107." pageId="13">FIGS 5B</figureCitation>
,
<figureCitation id="13502A01E560FFBDFB06F90CFB3A51A1" box="[1162,1176,1670,1695]" captionStart="Figure 6" captionStartId="12.[163,242,1512,1534]" captionTargetBox="[324,1281,196,1471]" captionTargetId="figure-110@12.[323,1283,195,1473]" captionTargetPageId="12" captionText="Figure 6. Diagnostic external traits for the differentiation of N. fengi, N. huang, N. fulvescens and N. mekongis. A, dorsal hairs. B, ventral tail. Note that N. fengi has a hairier tail and darker skin without spines." pageId="13">6</figureCitation>
)
</paragraph>
<paragraph id="8BD43684E560FFBDFCA5F924FAD651FA" blockId="13.[809,1396,1710,1732]" box="[809,1396,1710,1732]" pageId="13">Indomalayan niviventer, chestnut white-bellied rat.</paragraph>
</subSubSection>
<subSubSection id="C371650FE560FFBDFCA5F969FAA85029" pageId="13" type="reference_group">
<paragraph id="8BD43684E560FFBDFCA5F969FB7151C6" blockId="13.[809,1290,1762,1815]" box="[809,1235,1762,1784]" pageId="13">
<taxonomicName id="4C6B4D07E560FFBDFCA5F969FBDF51C6" authority="Grey, 1847: 18" authorityName="Grey" authorityPageNumber="18" authorityYear="1847" box="[809,1149,1762,1784]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="13" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B91FEA96E560FFBDFCA5F969FC7151C9" box="[809,979,1762,1784]" italics="true" pageId="13">Mus fulvescens</emphasis>
Grey, 1847: 18
</taxonomicName>
.
<collectingCountry id="F37C7614E560FFBDFB0BF968FB6D51C6" box="[1159,1231,1762,1784]" name="Nepal" pageId="13">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8BD43684E560FFBDFCA5F888FAA85029" blockId="13.[809,1290,1762,1815]" box="[809,1290,1793,1815]" pageId="13">
<taxonomicName id="4C6B4D07E560FFBDFCA5F888FB175028" ID-CoL="44ND7" authority="Hodgson, 1849: 203" authorityName="Hodgson" authorityPageNumber="203" authorityYear="1849" box="[809,1205,1793,1815]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="13" phylum="Chordata" rank="species" species="caudatior">
<emphasis id="B91FEA96E560FFBDFCA5F888FC6D5028" box="[809,975,1793,1815]" italics="true" pageId="13">Mus caudatior</emphasis>
Hodgson, 1849: 203
</taxonomicName>
.
<collectingCountry id="F37C7614E560FFBDFB33F88BFAA55029" box="[1215,1287,1793,1815]" name="Nepal" pageId="13">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C371650FE560FFBEFCA5F8BFFC5157C4" lastPageId="14" pageId="13" type="materials_examined">
<paragraph id="8BD43684E560FFBDFCA5F8BFFA335054" blockId="13.[809,1425,1845,1898]" pageId="13">
<emphasis id="B91FEA96E560FFBDFCA5F8BFFC035074" box="[809,929,1845,1866]" italics="true" pageId="13">
<typeStatus id="54D08826E560FFBDFCA5F8BFFC385074" box="[809,922,1845,1866]" pageId="13" type="holotype">Holotype</typeStatus>
:
</emphasis>
NHM 45.1.8.376. Collected by B. H. Hodgson. The
<typeStatus id="54D08826E560FFBDFC41F8DEFB8D5054" box="[973,1071,1876,1898]" pageId="13" type="holotype">holotype</typeStatus>
is badly damaged. The animal
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFF4FFE5957C4" blockId="14.[163,778,197,250]" pageId="14">is small and immature, with the dorsal and ventral coat missing in several places.</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFEA9FE325606" blockId="14.[163,400,290,312]" box="[163,400,290,312]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FFEA9FE9A5609" box="[163,312,290,312]" italics="true" pageId="14">Type locality:</emphasis>
<collectingCountry id="F37C7614E563FFBEFEC9FEA8FE2F5606" box="[325,397,290,312]" name="Nepal" pageId="14">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFEE8FD12543B" blockId="14.[163,779,353,774]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FFEE8FEFA5649" box="[163,344,354,375]" italics="true" pageId="14">Measurements:</emphasis>
No measurements were available for the
<typeStatus id="54D08826E563FFBEFE8AFE0AFED756A8" box="[262,373,384,406]" pageId="14" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED7AAE41E563FFBEFE0AFE0AFE1656AB" box="[390,436,384,405]" country="United Kingdom" name="Bristol Museum" pageId="14">BM</collectionCode>
, EMs and CMs (mean ± 1 SD, range,
<collectionCode id="ED7AAE41E563FFBEFEBDFE15FEF9568A" box="[305,347,415,436]" country="United Kingdom" name="Bristol Museum" pageId="14">BM</collectionCode>
in g, measurements in mm) from molecular vouchers and museum specimens:
<collectionCode id="ED7AAE41E563FFBEFF2FFE56FF6E56CF" box="[163,204,476,497]" country="United Kingdom" name="Bristol Museum" pageId="14">BM</collectionCode>
= 66.82 ± 14.04 (4091,
<emphasis id="B91FEA96E563FFBEFE67FE57FE5D56CC" box="[491,511,477,498]" italics="true" pageId="14">N</emphasis>
= 37). EMs of
<specimenCount id="9D6DFD0DE563FFBEFD26FE56FCA856CC" box="[682,778,476,498]" pageId="14" type="adult">37 adult</specimenCount>
specimens: HBL = 131.60 ± 11.18, TL = 187.74 ± 14.94, HFL = 27.22 ± 2.01 and EL = 20 ± 2.50. CMs from 29 intact adult specimens: TLC = 15.15 ± 1.73, NL = 12.92 ± 1.22, GWS = 5.40 ± 0.44, SDO = 5.65 ± 0.30, ZB = 15.09 ± 1.24, GMB = 13.28 ± 0.61,
<collectionCode id="ED7AAE41E563FFBEFDC5FDFCFDC855B5" box="[585,618,630,651]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" type="Museum">PL</collectionCode>
= 17.27 ± 0.95, IFL = 5.79 ± 0.59, WIF = 2.52 ± 0.20, GPB = 6.68 ± 0.35, LTB = 5.84 ± 0.52,
<collectionCode id="ED7AAE41E563FFBEFE41FD39FD8B55F6" box="[461,553,691,712]" pageId="14">ULMM</collectionCode>
= 5.76 ± 0.30,
<collectionCode id="ED7AAE41E563FFBEFF2FFD58FF5955D9" box="[163,251,722,743]" pageId="14">ULMD</collectionCode>
= 8.86 ± 0.68,
<collectionCode id="ED7AAE41E563FFBEFE62FD58FDBA55D9" box="[494,536,722,743]" country="France" name="Musee de Lectoure" pageId="14">ML</collectionCode>
= 16.70 ± 1.13,
<collectionCode id="ED7AAE41E563FFBEFF2FFD7AFF50543B" box="[163,242,752,773]" pageId="14">LLMM</collectionCode>
= 5.66 ± 0.33 and
<collectionCode id="ED7AAE41E563FFBEFE48FD7AFDAD543B" box="[452,527,752,773]" pageId="14">LLMD</collectionCode>
= 4.61 ± 0.49.
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFCA5FDCF549E" blockId="14.[163,779,814,928]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FFCA5FDBA547D" box="[163,536,814,836]" italics="true" pageId="14">
Diagnosis:
<taxonomicName id="4C6B4D07E563FFBEFEABFCA5FDBA547D" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[295,536,814,836]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="fulvescens">Niviventer fulvescens</taxonomicName>
</emphasis>
can be distinguished from other species within the
<collectionCode id="ED7AAE41E563FFBEFE71FCC7FDE1545C" box="[509,579,845,866]" pageId="14">NFSC</collectionCode>
by its longer tail and ears. It could be easily mistaken for
<taxonomicName id="4C6B4D07E563FFBEFDE2FCE6FD7A54BF" baseAuthorityName="BONHOTE" baseAuthorityYear="1905" box="[622,728,876,897]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="huang">
<emphasis id="B91FEA96E563FFBEFDE2FCE6FD2054BF" box="[622,642,876,897]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFD03FCE6FD7A54BF" box="[655,728,876,897]" italics="true" pageId="14">huang</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C6B4D07E563FFBEFF2FFC01FE9454A1" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[163,310,906,928]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B91FEA96E563FFBEFF2FFC01FF15549E" box="[163,183,907,928]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFF48FC00FE9454A1" box="[196,310,906,927]" italics="true" pageId="14">fulvescens</emphasis>
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inhabits higher elevations.
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFC43FE2B5305" blockId="14.[163,778,969,1083]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FFC43FEE854E0" box="[163,330,969,990]" italics="true" pageId="14">Distribution:</emphasis>
<collectingCountry id="F37C7614E563FFBEFED7FC43FE0B54E1" box="[347,425,969,991]" name="Nepal" pageId="14">Nepal</collectingCountry>
, south-western
<collectingCountry id="F37C7614E563FFBEFD0EFC43FD6D54E1" box="[642,719,969,991]" name="China" pageId="14">China</collectingCountry>
and northern Indochina. In
<collectingCountry id="F37C7614E563FFBEFE41FC62FDB554C0" box="[461,535,1000,1022]" name="China" pageId="14">China</collectingCountry>
it is distributed in higher mountains from south-western
<collectingRegion id="49AFF866E563FFBEFD06FB8DFD4B5322" box="[650,745,1031,1052]" country="China" name="Yunnan" pageId="14">Yunnan</collectingRegion>
to south-eastern
<collectingRegion id="49AFF866E563FFBEFECAFBAFFE275305" box="[326,389,1061,1083]" country="China" name="Xizang" pageId="14">Tibet</collectingRegion>
.
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FFBEEFEB351BD" blockId="14.[163,779,1124,1667]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FFBEEFD9B5347" box="[163,569,1124,1145]" italics="true" pageId="14">
Comments:
<taxonomicName id="4C6B4D07E563FFBEFEBAFBEEFD9B5347" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[310,569,1124,1145]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="fulvescens">Niviventer fulvescens</taxonomicName>
</emphasis>
is taxonomically the most ambiguous species of the
<collectionCode id="ED7AAE41E563FFBEFDC7FB09FD3753A6" box="[587,661,1155,1176]" pageId="14">NFSC</collectionCode>
. Previous studies largely overestimated the distribution range of this species (
<bibRefCitation id="EFFA4B75E563FFBEFEE1FB4AFE5F53EB" author="Allen GM" box="[365,509,1216,1238]" pageId="14" refId="ref12938" refString="Allen GM. 1938. The mammals of China and Mongolia. New York: American Museum of Natural History." type="book" year="1938">Allen, 1938</bibRefCitation>
;
<bibRefCitation id="EFFA4B75E563FFBEFD9CFB4AFF7F53CA" author="Musser GG &amp; Carleton MD" pageId="14" refId="ref14380" refString="Musser GG, Carleton MD. 2005. Superfamily Muroidea. In: Wilson DE, Reeder DM, eds. Mammal species of the world, a taxonomic and geographic reference. Baltimore: Johns Hopkins University Press." type="book" year="2005">Musser &amp; Carleton, 2005</bibRefCitation>
). According to molecular voucher specimens, it inhabits middle- to high-elevation mountains in south-western Yunnan and south-eastern Tibet. The present analyses identifies two major genetic lineages within
<taxonomicName id="4C6B4D07E563FFBEFF70FAD0FE3E5250" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[252,412,1369,1391]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B91FEA96E563FFBEFF70FAD0FEB25251" box="[252,272,1370,1391]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFEADFAD3FE3E5250" box="[289,412,1369,1390]" italics="true" pageId="14">fulvescens</emphasis>
</taxonomicName>
(
<figureCitation id="13502A01E563FFBEFE22FAD3FE515250" box="[430,499,1369,1391]" captionStart="Figure 1" captionStartId="2.[163,242,1817,1839]" captionTargetBox="[322,1282,873,1774]" captionTargetId="figure-384@2.[319,1285,870,1777]" captionTargetPageId="2" captionText="Figure 1. Distribution of molecular voucher specimens showing the spatial partitions between species within the NFSC. The holotype localities of the taxa studied in the present work are indicated on the map using colours corresponding to the molecular voucher specimens." pageId="14">Fig. 1</figureCitation>
); the first lineage was confined to south-eastern Tibet along the Himalayan Mountains. The second lineage was distributed in south-western Yunnan, western Sichuan and northern
<collectingCountry id="F37C7614E563FFBEFE98FA5EFEDE52D4" box="[276,380,1492,1514]" name="Vietnam" pageId="14">Vietnam</collectingCountry>
. Pelage coloration differs slightly between these two genetic lineages. The lineage from south-eastern Tibet has a darker coloration than those from Yunnan. Spines are more prominent in individuals from Yunnan than in those from the other locations.
</paragraph>
<paragraph id="8BD43684E563FFBEFF2FF926FC5157C4" blockId="14.[163,780,1708,1883]" lastBlockId="14.[827,1442,197,250]" pageId="14">
<emphasis id="B91FEA96E563FFBEFF2FF926FEDE51FF" box="[163,380,1708,1729]" italics="true" pageId="14">Sympatric species:</emphasis>
The distribution of
<taxonomicName id="4C6B4D07E563FFBEFDFDF926FCA951FF" baseAuthorityName="Smith &amp; Xie" baseAuthorityYear="2008" box="[625,779,1708,1729]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="fulvescens">
<emphasis id="B91FEA96E563FFBEFDFDF926FD2751FF" box="[625,645,1708,1729]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFD18F926FCA951FF" box="[660,779,1708,1729]" italics="true" pageId="14">fulvescens</emphasis>
</taxonomicName>
largely overlaps with those of
<taxonomicName id="4C6B4D07E563FFBEFD9AF941FD1251E1" baseAuthorityName="Thomas" baseAuthorityYear="1911" box="[534,688,1738,1760]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="andersoni">
<emphasis id="B91FEA96E563FFBEFD9AF941FD8851DE" box="[534,554,1739,1760]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFDB6F940FD1251E1" box="[570,688,1738,1759]" italics="true" pageId="14">andersoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D07E563FFBEFD7CF941FEBA51C1" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="niviventer">
<emphasis id="B91FEA96E563FFBEFD7CF941FCA651DE" box="[752,772,1739,1760]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFF2FF960FEBA51C1" box="[163,280,1770,1791]" italics="true" pageId="14">niviventer</emphasis>
</taxonomicName>
. The last two species also inhabit middleto high-elevation mountains in southern
<collectingCountry id="F37C7614E563FFBEFD34F882FCA75020" box="[696,773,1800,1822]" name="China" pageId="14">China</collectingCountry>
.
<taxonomicName id="4C6B4D07E563FFBEFF2FF8ADFE3B5005" baseAuthorityName="Thomas" baseAuthorityYear="1911" box="[163,409,1830,1852]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="andersoni">
<emphasis id="B91FEA96E563FFBEFF2FF8ADFE3B5005" box="[163,409,1830,1852]" italics="true" pageId="14">Niviventer andersoni</emphasis>
</taxonomicName>
is distinguishable by its larger size and bicoloured tail (the one-third near the tip is white) and
<taxonomicName id="4C6B4D07E563FFBEFC37FF4CFBEC57E5" box="[955,1102,197,219]" class="Mammalia" family="Muridae" genus="Niviventer" kingdom="Animalia" order="Rodentia" pageId="14" phylum="Chordata" rank="species" species="niviventer">
<emphasis id="B91FEA96E563FFBEFC37FF4CFC6D57E5" box="[955,975,198,219]" italics="true" pageId="14">N</emphasis>
.
<emphasis id="B91FEA96E563FFBEFC50FF4CFBEC57E5" box="[988,1102,198,219]" italics="true" pageId="14">niviventer</emphasis>
</taxonomicName>
belongs to the
<collectionCode id="ED7AAE41E563FFBEFB75FF4FFAE157E4" box="[1273,1347,197,218]" pageId="14">NNSC</collectionCode>
and has a darker pelage.
</paragraph>
</subSubSection>
</treatment>
</document>