283 lines
32 KiB
XML
283 lines
32 KiB
XML
<document id="1B1A0C4CEA9D9461A9310BD40C846567" ID-DOI="10.5281/zenodo.198244" ID-GBIF-Dataset="2145a6ff-d31c-45f1-bea1-af8477a51da6" ID-ISSN="1175-5326" ID-Zenodo-Dep="198244" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460290874690" checkinUser="plazi" docAuthor="Gharali, Babak, Kamali, Karim, Evenhuis, Neal, Talebi, Ali Asghar & Khalgani, Jafar" docDate="2010" docId="03B987B2FFE15E3BFF50F27CFD3954CF" docLanguage="en" docName="zt02627p019.pdf" docOrigin="Zootaxa 2627" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empidideicus matricarius Gharali & Evenhuis, sp. nov." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FF80FFCAFFEB5E36FFC7F449FFC05658" masterDocTitle="First record of the genus Empidideicus (Diptera: Bombylioidea: Mythicomyiidae) from Iran, with description of six new species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="11" updateTime="1698251303231" updateUser="plazi">
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<mods:mods id="991B2E032B68DA904FEE098429CA9BCE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="94A082B4C30CC285F5DAF237866CC97C">
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<mods:title id="A7697F6D4DE273A66BA64A6654EEFC94">First record of the genus Empidideicus (Diptera: Bombylioidea: Mythicomyiidae) from Iran, with description of six new species</mods:title>
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<mods:name id="C1F4938CDFB9B14A42767F12B40C7998" type="personal">
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<mods:namePart id="0778D1E736D06AA8E358D928D282C4EB">Gharali, Babak</mods:namePart>
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<mods:name id="C19B32A61076CACEC6925956792F73AD" type="personal">
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<mods:roleTerm id="C98FC4FD0C5A84BCDE386CD8C25D7E70">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3CA2041EA6D7BF91824D071142F96F70">Kamali, Karim</mods:namePart>
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</mods:name>
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<mods:name id="AD1A6646D0D70978CDA277D0EBD8FD62" type="personal">
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<mods:role id="808129A7F74B080260CD091543CA3048">
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<mods:roleTerm id="B6AEA7F7BB37C739713318A1B78CFF72">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E32B0C4585FFEAFE755641D4CD13139C">Evenhuis, Neal</mods:namePart>
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</mods:name>
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<mods:name id="55CD9529A483BBC33A8113C728E3D4A1" type="personal">
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<mods:roleTerm id="9E7AA9FC5079722A09A11C507160C909">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FCE0F3F7565ED689E372CC689DDD57BD">Talebi, Ali Asghar</mods:namePart>
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</mods:name>
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<mods:name id="B8F796AFB6E93BFA2800E04A6AE4ED82" type="personal">
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<mods:role id="DBBF789F7ECE9010FF0863990009504D">
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<mods:roleTerm id="FED64784AC9446C6F1D5624974A89EF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A275378875F7296E1ACD81741F3CA5B9">Khalgani, Jafar</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C263EA6AB06D6F18FB23E8DFC7F3BD4E">text</mods:typeOfResource>
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<mods:title id="69DD36549DAC8E2FC2B8B6CA3B79E8D0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="63F720B568915CAF1A59B1369C752E7F">
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<mods:date id="5C31711D497C8E0DBD09DB49491B9E53">2010</mods:date>
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<mods:detail id="99609FDEC7DD5FCD4281E7D5BC79EDAB" type="volume">
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<mods:number id="01A634151D21713292A1C1D22DCB545F">2627</mods:number>
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<mods:classification id="81506B3253ADB8630E119CFC5FDFB39E">journal article</mods:classification>
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<mods:identifier id="366894B4965270B8AC2B6DF4B8A94003" type="DOI">10.5281/zenodo.198244</mods:identifier>
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<mods:identifier id="2814BE51F7E1E02B3F83CAC92942BB11" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EEA8B012E640F3DBC5A32E39D462C267" type="Zenodo-Dep">198244</mods:identifier>
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<treatment id="03B987B2FFE15E3BFF50F27CFD3954CF" ID-DOI="http://doi.org/10.5281/zenodo.6209831" ID-GBIF-Taxon="119398635" ID-Zenodo-Dep="6209831" LSID="urn:lsid:plazi:treatment:03B987B2FFE15E3BFF50F27CFD3954CF" httpUri="http://treatment.plazi.org/id/03B987B2FFE15E3BFF50F27CFD3954CF" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<subSubSection id="C30A652FFFE15E3CFF50F27CFCF0510A" pageId="10" pageNumber="11" type="nomenclature">
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||
<paragraph id="8BAF36A4FFE15E3CFF50F27CFCFE5017" blockId="10.[151,830,1589,1650]" box="[151,830,1589,1615]" pageId="10" pageNumber="11">
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||
<heading id="D0E781C8FFE15E3CFF50F27CFCFE5017" bold="true" box="[151,830,1589,1615]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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||
<emphasis id="B964EAB6FFE15E3CFF50F27CFCFE5017" bold="true" box="[151,830,1589,1615]" pageId="10" pageNumber="11">
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<taxonomicName id="4C104D27FFE15E3CFF50F27CFD155017" authority="Gharali & Evenhuis" authorityName="Gharali & Evenhuis" box="[151,725,1589,1615]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="matricarius" status="sp. nov.">
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<emphasis id="B964EAB6FFE15E3CFF50F27CFE0E5017" bold="true" box="[151,462,1589,1615]" italics="true" pageId="10" pageNumber="11">Empidideicus matricarius</emphasis>
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Gharali & Evenhuis
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25757CDFFE15E3CFD26F27CFCFE5017" box="[737,830,1589,1615]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAF36A4FFE15E3CFF50F211FEBA502A" blockId="10.[151,830,1589,1650]" box="[151,378,1624,1650]" pageId="10" pageNumber="11">
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||
(
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<figureCitation id="132B2A21FFE15E3CFF58F211FF30502A" box="[159,240,1624,1650]" captionStart="FIGURE 5" captionStartId="11.[151,259,1963,1987]" captionTargetBox="[177,1417,193,1913]" captionTargetId="figure@11.[165,1422,166,1942]" captionTargetPageId="11" captionText="FIGURE 5. Empidideicus matricarius: a—antennae lateral view, b—wing, c—phallic complex dorsal view, depandrium anterodorsal view, e—phallic complex lateral view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198248/files/figure.png" pageId="10" pageNumber="11">Figs. 5</figureCitation>
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a–g, 7d, 8e)
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</paragraph>
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<paragraph id="8BAF36A4FFE15E3CFF50F2EBFCF0510A" blockId="10.[151,1437,1698,2024]" pageId="10" pageNumber="11">
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<emphasis id="B964EAB6FFE15E3CFF50F2EBFE6C50E4" bold="true" box="[151,428,1698,1724]" pageId="10" pageNumber="11">Specimens examined:</emphasis>
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<typeStatus id="54AB8806FFE15E3CFE72F2EAFDE550E5" box="[437,549,1699,1725]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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female (dried by HMDS) and
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<specimenCount id="9D16FD2DFFE15E3CFC5AF2EAFC3250E5" box="[925,1010,1699,1725]" pageId="10" pageNumber="11" type="male">3 male</specimenCount>
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,
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<specimenCount id="9D16FD2DFFE15E3CFC38F2EAFBB750E5" box="[1023,1143,1699,1725]" pageId="10" pageNumber="11" type="female">10 female</specimenCount>
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<typeStatus id="54AB8806FFE15E3CFB47F2EAFB3450E5" box="[1152,1268,1699,1725]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
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(preserved in alcohol),
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<collectingCountry id="F3077634FFE15E3CFECCF281FE9A50BA" box="[267,346,1736,1762]" name="Iran" pageId="10" pageNumber="11">IRAN</collectingCountry>
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: research field, Faculty of Agriculture, Tarbiat Modares University, Tehran,
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<date id="FFAE1064FFE15E3CFAC3F281FA5850BA" box="[1284,1432,1736,1762]" pageId="10" pageNumber="11" value="2008-07-07">7 July 2008</date>
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, swept on
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<taxonomicName id="4C104D27FFE15E3CFECDF2A7FE2F515F" box="[266,495,1774,1799]" class="Magnoliopsida" family="Asteraceae" genus="Matricaria" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="recutita">
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<emphasis id="B964EAB6FFE15E3CFECDF2A7FE2F515F" box="[266,495,1774,1799]" italics="true" pageId="10" pageNumber="11">Matricaria recutita</emphasis>
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</taxonomicName>
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, leg. B. Gharali (all in TMUC),
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<specimenCount id="9D16FD2DFFE15E3CFCB1F2A7FC185150" box="[886,984,1774,1800]" pageId="10" pageNumber="11" type="male">12 male</specimenCount>
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,
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<specimenCount id="9D16FD2DFFE15E3CFC22F2A7FB8D5150" box="[997,1101,1774,1800]" pageId="10" pageNumber="11" type="female">7 female</specimenCount>
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<typeStatus id="54AB8806FFE15E3CFB93F2A7FB065150" box="[1108,1222,1774,1800]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
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same as
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<typeStatus id="54AB8806FFE15E3CFAF3F2A7FA5C5150" box="[1332,1436,1774,1800]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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(BPBM);
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<specimenCount id="9D16FD2DFFE15E3CFECBF35AFE405175" box="[268,384,1811,1837]" pageId="10" pageNumber="11" type="female">10 female</specimenCount>
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<typeStatus id="54AB8806FFE15E3CFE40F35AFE375175" box="[391,503,1811,1837]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
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same as
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<typeStatus id="54AB8806FFE15E3CFDA4F35AFD0A5175" box="[611,714,1811,1837]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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(ZMHB);
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<specimenCount id="9D16FD2DFFE15E3CFC8EF35AFC665175" box="[841,934,1811,1837]" pageId="10" pageNumber="11" type="male">2 males</specimenCount>
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,
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<specimenCount id="9D16FD2DFFE15E3CFC75F35AFBF25175" box="[946,1074,1811,1837]" pageId="10" pageNumber="11" type="female">10 females</specimenCount>
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same locality data as
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<typeStatus id="54AB8806FFE15E3CFAF2F35AFA5B5175" box="[1333,1435,1811,1837]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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except
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<date id="FFAE1064FFE15E3CFF2CF371FE47510A" box="[235,391,1848,1874]" pageId="10" pageNumber="11" value="2009-06-25">25 June 2009</date>
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(personal collection of first author).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A652FFFE15E3CFF01F314FD5F51B0" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BAF36A4FFE15E3CFF01F314FD5F51B0" blockId="10.[151,1437,1698,2024]" pageId="10" pageNumber="11">
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<emphasis id="B964EAB6FFE15E3CFF01F314FEFF512F" bold="true" box="[198,319,1885,1911]" pageId="10" pageNumber="11">Diagnosis</emphasis>
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.
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<taxonomicName id="4C104D27FFE15E3CFE8AF317FE36512F" box="[333,502,1886,1911]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="matricarius">
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<emphasis id="B964EAB6FFE15E3CFE8AF317FE36512F" box="[333,502,1886,1911]" italics="true" pageId="10" pageNumber="11">E. matricarius</emphasis>
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</taxonomicName>
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is separated from congeners in
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<collectingCountry id="F3077634FFE15E3CFCA0F317FC565120" box="[871,918,1886,1912]" name="Iran" pageId="10" pageNumber="11">Iran</collectingCountry>
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by the combination of following characters: Occiput yellow laterally; mesonotum with brown or blackish brown pattern, prescutellar area and sctuellum yellow; second flagellomere much shorter than first flagellomere; invagination of reservoir shallow and furca without sclerotisation around genital orifice.
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</paragraph>
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</subSubSection>
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<caption id="DF6F662CFFE05E3DFF50F3E2FAEE51BF" httpUri="https://zenodo.org/record/198248/files/figure.png" pageId="11" pageNumber="12" targetBox="[177,1417,193,1913]" targetPageId="11">
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<paragraph id="8BAF36A4FFE05E3DFF50F3E2FAEE51BF" blockId="11.[151,1436,1963,2023]" pageId="11" pageNumber="12">
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<emphasis id="B964EAB6FFE05E3DFF50F3E2FEE0519B" bold="true" box="[151,288,1963,1987]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
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<taxonomicName id="4C104D27FFE05E3DFEEFF3E5FD8F519B" box="[296,591,1964,1987]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="matricarius">
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<emphasis id="B964EAB6FFE05E3DFEEFF3E5FD8F519B" box="[296,591,1964,1987]" italics="true" pageId="11" pageNumber="12">Empidideicus matricarius</emphasis>
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</taxonomicName>
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: a—antennae lateral view, b—wing, c—phallic complex dorsal view, depandrium anterodorsal view, e—phallic complex lateral view, f—gonocoxa dorsal view, g—female genitalia.
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</paragraph>
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</caption>
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<subSubSection id="C30A652FFFE75E3AFF02F4DEFCC25016" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BAF36A4FFE75E3AFF02F4DEFC3656EA" blockId="12.[151,1437,151,2025]" box="[197,1014,151,178]" pageId="12" pageNumber="13">
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<emphasis id="B964EAB6FFE75E3AFF02F4DEFE9D56E9" bold="true" box="[197,349,151,177]" pageId="12" pageNumber="13">Description.</emphasis>
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Female. Length
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<quantity id="4CE89B41FFE75E3AFDE4F4D1FDA756EA" box="[547,615,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
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(Female)—
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<quantity id="4CE89B41FFE75E3AFD3FF4D1FC8F56EA" box="[760,847,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
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(Male) (n=5).
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</paragraph>
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<paragraph id="8BAF36A4FFE75E3AFF02F4F4FC17545A" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">
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Head (
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<figureCitation id="132B2A21FFE75E3AFED3F4F4FE9C568F" box="[276,348,189,215]" captionStart="FIGURE 7" captionStartId="15.[151,256,1963,1987]" captionTargetBox="[175,1410,194,1941]" captionTargetId="figure@15.[175,1410,194,1942]" captionTargetPageId="15" captionText="FIGURE 7. Lateral view of Empidideicus adults (females): a— E. amicus, b— E. ebellicus, c— E. greatheadi, d— E. matricarius, e— E. legulicoxa, f— E. persicus." httpUri="https://zenodo.org/record/198250/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
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d). Slightly higher than long; eye dichoptic, separated at vertex by 2 times distance between lateral ocelli; ocellar triangle black; vertex black, black color narrowly joined to upper eye margin; occiput black medially, yellow laterally; postgena yellow, frons slightly depressed medially with blackish brown longitudinal stripe from anterior ocellus to frontal depression; face, mentum, and oral margin yellow; antennae (
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<figureCitation id="132B2A21FFE75E3AFED4F51BFE9B5734" box="[275,347,338,364]" captionStart="FIGURE 4" captionStartId="10.[151,254,1479,1503]" captionText="FIGURE 4. Empidideicus legulicoxa: lateral view of thorax (= pleura)." pageId="12" pageNumber="13">Fig. 4</figureCitation>
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a) set in deep pocket; two basal segments of antennae yellow; scape short, subtrapezoidal; pedicel spheroid-ellipsoid, wider than long; first flagellomere dark brown, ovoid, length about 1.3 times width,; second flagellomere faintly brown, 0.7 times length of first flagellomere, apical stylus transparent, minute, 0.3 times length of second flagellomere, proboscis 2.5 times of head length, labrum black, sclerotised, stiff, pointed apically, length about half of proboscis; palpi not evident.
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</paragraph>
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<paragraph id="8BAF36A4FFE75E3AFF02F644FE0D555F" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">
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Thorax (
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<figureCitation id="132B2A21FFE75E3AFEF7F644FEBB547F" box="[304,379,525,551]" captionStart="FIGURE 8" captionStartId="16.[151,256,1295,1319]" captionTargetBox="[153,1436,189,1267]" captionTargetId="figure@16.[153,1438,189,1268]" captionTargetPageId="16" captionText="FIGURE 8. Dorsal view of Empidideicus mesonotum (females): a, b— E. amicus, c— E. ebellicus, d— E. greatheadi, e— E. matricarius, f— E. persicus." httpUri="https://zenodo.org/record/198251/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
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e). Mesonotum yellow to brownish yellow, with short moderately dense pubescence, dorsum with admedian whitish longitudinal vittae and pattern of brown and dark brown pattern medially and dorsolaterally; length of median longitudinal vittae ¾ length of mesonotum; humeral callus, notopleural area, postalar callus and rectangular prescutellar area yellowish, posterior margin of mesonotum faintly brownish. Scutellum yellow, pleura yellow with black color on following: anepisternum as crescent mark anteroventrally, ventral margin of anepimeron, lower three-fourth of katepisternum, and meron; halter stem and knob yellowish white.
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</paragraph>
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<paragraph id="8BAF36A4FFE75E3AFF02F75BFEFC550A" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">Legs. Yellow except coxa III with brown mark dorsally, two last tarsal segments brown, claws black, pulvilli white.</paragraph>
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<paragraph id="8BAF36A4FFE75E3AFF02F714FC655231" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">
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Wing (
|
||
<figureCitation id="132B2A21FFE75E3AFED2F714FE9A552F" box="[277,346,861,887]" captionStart="FIGURE 5" captionStartId="11.[151,259,1963,1987]" captionTargetBox="[177,1417,193,1913]" captionTargetId="figure@11.[165,1422,166,1942]" captionTargetPageId="11" captionText="FIGURE 5. Empidideicus matricarius: a—antennae lateral view, b—wing, c—phallic complex dorsal view, depandrium anterodorsal view, e—phallic complex lateral view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198248/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||
b). Hyaline, length (from base to end of R4+5) 2.6 times width (from end of R1 to CuA2); veins brown; costa, Sc, R4+5 and CuA2 well sclerotised; veins M1, M2, M1+2 and CuA1 sclerotised; vein separating basal cells, the base of Rs and A1 evanescent; costa ends slightly beyond end of R4+5; Sc incomplete; R4+5 fairly straight to wing margin, meeting costa well beyond level of CuA1; M1 separated from M2 at an acute angle, curved to wing margin; M2 straight, slightly curved apically, 1.8 times as long as M1+2; cell dm open apically; A1 straight to wing margin; anal cell open in wing margin by width subequal to r-m crossvein; fringe of hair shortest at wing tip becoming longer and more sparse toward base.
|
||
</paragraph>
|
||
<paragraph id="8BAF36A4FFE75E3AFF02F03DFC4E5281" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">Abdomen. Tergites I–VI yellow with transverse brown bands anteriorly, bands narrowing toward tergite VI; tergites II–V with scattered brown spots; remainder of tergites yellow; lateral margins of all tergites narrowly brown; sternites yellow except last sternite pale brown.</paragraph>
|
||
<paragraph id="8BAF36A4FFE75E3AFF02F0ADFD57505B" blockId="12.[151,1437,151,2025]" pageId="12" pageNumber="13">
|
||
Genitalia (
|
||
<figureCitation id="132B2A21FFE75E3AFE87F0ADFE4752A6" box="[320,391,1252,1278]" captionStart="FIGURE 5" captionStartId="11.[151,259,1963,1987]" captionTargetBox="[177,1417,193,1913]" captionTargetId="figure@11.[165,1422,166,1942]" captionTargetPageId="11" captionText="FIGURE 5. Empidideicus matricarius: a—antennae lateral view, b—wing, c—phallic complex dorsal view, depandrium anterodorsal view, e—phallic complex lateral view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198248/files/figure.png" pageId="12" pageNumber="13">
|
||
Fig.
|
||
<quantity id="4CE89B41FFE75E3AFEB0F0ADFE4752A6" box="[375,391,1252,1278]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="12" pageNumber="13" unit="g" value="5.0">5</quantity>
|
||
</figureCitation>
|
||
g). Spermathecal reservoir subglobular, slightly wider than high, with shallow cylindrical invagination, floor of invagination slightly puffed up, length of invagination 0.42 times reservoir height, width of invagination about 0.4 diameter of reservoir; apical spermathecal duct long, thin, length 1.3 times as long as sperm pump, slightly sclerotised apically; sperm pump long, thin and striated, about 1.9 times reservoir height, with shallow apical valve; basal duct short, slightly sclerotised, about half reservoir height; common spermathecal duct wide, membranous, thin, subequal to apical duct in length; furca U-shaped, much sclerotised apically, lateral arm apically broadened abruptly as two large and wide plates with deeply concave outer margin leaving two narrow processes.
|
||
</paragraph>
|
||
<paragraph id="8BAF36A4FFE75E3AFF02F246FE235071" blockId="12.[151,1437,151,2025]" box="[197,483,1551,1577]" pageId="12" pageNumber="13">Male. Similar to female.</paragraph>
|
||
<paragraph id="8BAF36A4FFE75E3AFF02F27DFCC25016" blockId="12.[151,1437,151,2025]" box="[197,770,1588,1614]" pageId="12" pageNumber="13">
|
||
Male genitalia (
|
||
<figureCitation id="132B2A21FFE75E3AFEBAF27DFE0D5016" box="[381,461,1588,1614]" captionStart="FIGURE 5" captionStartId="11.[151,259,1963,1987]" captionTargetBox="[177,1417,193,1913]" captionTargetId="figure@11.[165,1422,166,1942]" captionTargetPageId="11" captionText="FIGURE 5. Empidideicus matricarius: a—antennae lateral view, b—wing, c—phallic complex dorsal view, depandrium anterodorsal view, e—phallic complex lateral view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198248/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
|
||
c–f). Similar to
|
||
<taxonomicName id="4C104D27FFE75E3AFD4FF27DFD3B5015" box="[648,763,1588,1613]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="amicus">
|
||
<emphasis id="B964EAB6FFE75E3AFD4FF27DFD3B5015" box="[648,763,1588,1613]" italics="true" pageId="12" pageNumber="13">E. amicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A652FFFE75E3BFF02F210FB6B5784" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8BAF36A4FFE75E3BFF02F210FB6B5784" blockId="12.[151,1437,151,2025]" lastBlockId="13.[151,1436,152,663]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="B964EAB6FFE75E3AFF02F210FE97502B" bold="true" box="[197,343,1625,1651]" pageId="12" pageNumber="13">Comments.</emphasis>
|
||
<taxonomicName id="4C104D27FFE75E3AFEA7F210FD5D502A" box="[352,669,1625,1650]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="matricarius">
|
||
<emphasis id="B964EAB6FFE75E3AFEA7F210FD5D502A" box="[352,669,1625,1650]" italics="true" pageId="12" pageNumber="13">Empidideicus matricarius</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C104D27FFE75E3AFCB8F210FC38502A" box="[895,1016,1625,1650]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="amicus">
|
||
<emphasis id="B964EAB6FFE75E3AFCB8F210FC38502A" box="[895,1016,1625,1650]" italics="true" pageId="12" pageNumber="13">E. amicus</emphasis>
|
||
</taxonomicName>
|
||
morphologically and belongs to a species complex here we tentatively call the
|
||
<taxonomicName id="4C104D27FFE75E3AFD71F236FCCB50C0" box="[694,779,1663,1688]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="amicus">
|
||
<emphasis id="B964EAB6FFE75E3AFD71F236FCCB50C0" box="[694,779,1663,1688]" italics="true" pageId="12" pageNumber="13">amicus</emphasis>
|
||
</taxonomicName>
|
||
complex. The
|
||
<taxonomicName id="4C104D27FFE75E3AFC05F236FBD750C0" box="[962,1047,1663,1688]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="amicus">
|
||
<emphasis id="B964EAB6FFE75E3AFC05F236FBD750C0" box="[962,1047,1663,1688]" italics="true" pageId="12" pageNumber="13">amicus</emphasis>
|
||
</taxonomicName>
|
||
complex comprises at least two Iranian species whose mesonotal dorsum are yellow with a brown pattern and have the occiput yellow laterally. The male genitalia of both species are very similar and have some shared features, which are not seen in other species. The gonostyli in both species are deeply inserted in the median part of the gonocoxa instead of its apical portion; and the apical portion is less sclerotised and rounded, also the aedeagal bulb is very large and spheroid with only a short narrow tip. In the female genitalia both have a long and membranous common spermathecal duct that are seen in a few other species. They have few variable morphological differences and for the correct identification of species, the female genitalia should be checked.
|
||
<bibRefCitation id="EF814B55FFE75E3AFAF1F3CDFF29519B" author="Theodor" pageId="12" pageNumber="13" refString="Theodor, O. (1983) The genitalia of Bombyliidae (Diptera). Israel Academy of Sciences and Humanities, Jerusalem, 275 pp." type="book" year="1983">Theodor (1983)</bibRefCitation>
|
||
, in describing the female genitalia of
|
||
<taxonomicName id="4C104D27FFE75E3AFD77F3E0FC92519A" box="[688,850,1961,1986]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964EAB6FFE75E3AFD77F3E0FC92519A" box="[688,850,1961,1986]" italics="true" pageId="12" pageNumber="13">Empidideicus</emphasis>
|
||
</taxonomicName>
|
||
species, illustrated two unknown species that he said were closely related to
|
||
<taxonomicName id="4C104D27FFE75E3AFE24F386FDA951B0" box="[483,617,1999,2024]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mariouti">
|
||
<emphasis id="B964EAB6FFE75E3AFE24F386FDA951B0" box="[483,617,1999,2024]" italics="true" pageId="12" pageNumber="13">E. mariouti</emphasis>
|
||
</taxonomicName>
|
||
in external morphology but had distinct female genitalia. He labeled his figures of them as “
|
||
<emphasis id="B964EAB6FFE65E3BFE6FF4D1FE0256E9" box="[424,450,152,177]" italics="true" pageId="13" pageNumber="14">E.</emphasis>
|
||
sp. 1” and “
|
||
<emphasis id="B964EAB6FFE65E3BFD93F4D1FDAD56E9" box="[596,621,152,177]" italics="true" pageId="13" pageNumber="14">E.</emphasis>
|
||
sp. 2”. The genitalia of “
|
||
<emphasis id="B964EAB6FFE65E3BFC51F4D1FC6F56E9" box="[918,943,152,177]" italics="true" pageId="13" pageNumber="14">E.</emphasis>
|
||
sp. 1” appears the same as
|
||
<taxonomicName id="4C104D27FFE65E3BFB35F4D1FA5C56E9" box="[1266,1436,152,177]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="matricarius">
|
||
<emphasis id="B964EAB6FFE65E3BFB35F4D1FA5C56E9" box="[1266,1436,152,177]" italics="true" pageId="13" pageNumber="14">E. matricarius</emphasis>
|
||
</taxonomicName>
|
||
except the common spermathecal duct is absent in his figure. This difference may not be reliable because of the reasons discussed above for
|
||
<taxonomicName id="4C104D27FFE65E3BFDE1F4ABFD7656A3" box="[550,694,226,251]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ebellicus">
|
||
<emphasis id="B964EAB6FFE65E3BFDE1F4ABFD7656A3" box="[550,694,226,251]" italics="true" pageId="13" pageNumber="14">E. ebellicus</emphasis>
|
||
</taxonomicName>
|
||
. Recently,
|
||
<bibRefCitation id="EF814B55FFE65E3BFC85F4ABFBB956A4" box="[834,1145,226,252]" pageId="13" pageNumber="14" refString="El-Hawagry, M. S. A., Abdel-Dayem, M. S. & Hassan, S. A. (2003) An annotated key to the species of family Mythicomyiidae (Diptera) from Egypt. Bulletin of the Entomological Society of Egypt, 80, 77 - 84." type="journal article">
|
||
El-Hawagry
|
||
<emphasis id="B964EAB6FFE65E3BFC27F4ABFBD756A3" box="[992,1047,226,251]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
|
||
. (2003)
|
||
</bibRefCitation>
|
||
illustrated a part of the female genitalia of
|
||
<taxonomicName id="4C104D27FFE65E3BFEBBF541FDC35779" box="[380,515,264,289]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mariouti">
|
||
<emphasis id="B964EAB6FFE65E3BFEBBF541FDC35779" box="[380,515,264,289]" italics="true" pageId="13" pageNumber="14">E. mariouti</emphasis>
|
||
</taxonomicName>
|
||
(the spermathecal reservoir and a part of apical tube). His illustration appears to be the same as Theodor’s figure of “
|
||
<emphasis id="B964EAB6FFE65E3BFDA0F564FD41571E" box="[615,641,301,326]" italics="true" pageId="13" pageNumber="14">E.</emphasis>
|
||
sp. 1”. Unfortunately, he did not depict the furca, a very important character to distinguish species, because he had only a single female specimen and this part was damaged during preparation of genitalia (El Hawagry, pers. comm.). We believe El Hawagry’s specimen and Theodor’s “
|
||
<emphasis id="B964EAB6FFE65E3BFF63F5D4FF7E57EE" box="[164,190,413,438]" italics="true" pageId="13" pageNumber="14">E.</emphasis>
|
||
sp.1” are the same our species and different from
|
||
<taxonomicName id="4C104D27FFE65E3BFCDEF5D4FC6157EE" box="[793,929,413,438]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mariouti">
|
||
<emphasis id="B964EAB6FFE65E3BFCDEF5D4FC6157EE" box="[793,929,413,438]" italics="true" pageId="13" pageNumber="14">E. mariouti</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<taxonomicName id="4C104D27FFE65E3BFC21F5D4FBFA57EE" box="[998,1082,413,438]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="amicus">
|
||
<emphasis id="B964EAB6FFE65E3BFC21F5D4FBFA57EE" box="[998,1082,413,438]" italics="true" pageId="13" pageNumber="14">amicus</emphasis>
|
||
</taxonomicName>
|
||
complex may also include
|
||
<taxonomicName id="4C104D27FFE65E3BFA44F5D4FF3C5783" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mariouti">
|
||
<emphasis id="B964EAB6FFE65E3BFA44F5D4FF3C5783" italics="true" pageId="13" pageNumber="14">E. mariouti</emphasis>
|
||
</taxonomicName>
|
||
but in order to confirm this, the
|
||
<typeStatus id="54AB8806FFE65E3BFDBEF58BFD6C5784" box="[633,684,450,476]" pageId="13" pageNumber="14">type</typeStatus>
|
||
and its female genitalia should be checked.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A652FFFE65E3BFF02F5AEFCF7547F" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BAF36A4FFE65E3BFF02F5AEFCF7547F" blockId="13.[151,1436,152,663]" pageId="13" pageNumber="14">
|
||
<emphasis id="B964EAB6FFE65E3BFF02F5AEFE985459" bold="true" box="[197,344,487,513]" pageId="13" pageNumber="14">Etymology.</emphasis>
|
||
The specific epithet derives from the genus of the plant
|
||
<collectingCountry id="F3077634FFE65E3BFBDDF5A1FBBC545A" box="[1050,1148,488,514]" name="Germany" pageId="13" pageNumber="14">German</collectingCountry>
|
||
chamomile
|
||
<taxonomicName id="4C104D27FFE65E3BFAD2F5A1FE4F547F" class="Magnoliopsida" family="Asteraceae" genus="Matricaria" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="recutita">
|
||
<emphasis id="B964EAB6FFE65E3BFAD2F5A1FF31547E" italics="true" pageId="13" pageNumber="14">Matricaria recutita</emphasis>
|
||
(Asteraceae)
|
||
</taxonomicName>
|
||
, from which specimens were swept.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A652FFFE65E3BFF02F67BFD3954CF" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BAF36A4FFE65E3BFF02F67BFBB35414" blockId="13.[151,1436,152,663]" box="[197,1139,562,588]" pageId="13" pageNumber="14">
|
||
<emphasis id="B964EAB6FFE65E3BFF02F67BFEA55414" bold="true" box="[197,357,562,588]" pageId="13" pageNumber="14">Distribution.</emphasis>
|
||
This species is currently known only from
|
||
<collectingCountry id="F3077634FFE65E3BFCA4F67BFC525414" box="[867,914,562,588]" name="Iran" pageId="13" pageNumber="14">Iran</collectingCountry>
|
||
(Tehran province).
|
||
</paragraph>
|
||
<paragraph id="8BAF36A4FFE65E3BFF02F61EFD3954CF" blockId="13.[151,1436,152,663]" pageId="13" pageNumber="14">
|
||
<emphasis id="B964EAB6FFE65E3BFF02F61EFE915429" bold="true" box="[197,337,599,625]" pageId="13" pageNumber="14">Variability.</emphasis>
|
||
Some specimens are paler and do not have brown spots on the abdomen or anepisternum and the coxae can also be yellow without a brown mark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |