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<mods:title id="24AE524BC28F6F2CCF2627B4DD6176AA">On the taxonomy of Philyra sagittifera (Alcock, 1896) and P. concinnus Ghani &amp; Tirmizi, 1995 (Decapoda, Brachyura, Leucosiidae), with description of a new genus from the Indian Ocean</mods:title>
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<mods:affiliation id="36608A9D73C558104C63AB2ACE3291EE">Marine Biodiversity and Ecology Lab, Department of Zoology, The Maharaja Sayajirao University of Baroda, Vadodara, Gujarat, India. &amp; Animal Taxonomy and Ecology Laboratory, Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India.</mods:affiliation>
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<mods:affiliation id="DD3FF64BAD827526DF1DDDC34922B586">Center of Excellence in Phylogeny of Living Organisms, School of Biology, University of Tehran, Tehran, Iran. rnaderloo @ ut. ac. ir; https: // orcid. org / 0000 - 0002 - 5020 - 0962</mods:affiliation>
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<emphasis id="B976EACAFFFEFF835C858637FCEDF9B3" bold="true" box="[151,395,1570,1597]" italics="true" pageId="4" pageNumber="434">Tirmilyra sagittifera</emphasis>
(
<bibRefCitation id="EF934B29FFFEFF835D898637FF5BF9B3" author="Alcock, A." box="[411,573,1570,1596]" pageId="4" pageNumber="434" pagination="1 - 296" refId="ref6079" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2 The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, India, 65, 1 - 296. https: // doi. org / 10.5962 / bhl. title. 16033" type="journal article" year="1896">Alcock, 1896</bibRefCitation>
)
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(
<figureCitation id="13392A5DFFFEFF835C8D8652FC40F9EE" box="[159,294,1607,1633]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="4" pageNumber="434">Figs. 1A–D</figureCitation>
, 
<figureCitation id="13392A5DFFFEFF835D238652FC10F9EE" box="[305,374,1607,1633]" captionStart="FIGURE 2" captionStartId="3.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1590]" captionTargetId="figure-22@3.[151,1436,181,1590]" captionTargetPageId="3" captionText="FIGURE 2. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) dorsal view; (B) ventral view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254912" httpUri="https://zenodo.org/record/8254912/files/figure.png" pageId="4" pageNumber="434">2A, B</figureCitation>
, 
<figureCitation id="13392A5DFFFEFF835D908652FCA1F9EE" box="[386,455,1607,1633]" captionStart="FIGURE 3" captionStartId="6.[151,250,1328,1352]" captionTargetBox="[160,1427,678,1295]" captionTargetId="figure-271@6.[151,1436,669,1304]" captionTargetPageId="6" captionText="FIGURE 3. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) maxilliped 3; (B) pleon. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) maxilliped 3; (D) pleon." figureDoi="http://doi.org/10.5281/zenodo.8254914" httpUri="https://zenodo.org/record/8254914/files/figure.png" pageId="4" pageNumber="434">3A, B</figureCitation>
, 
<figureCitation id="13392A5DFFFEFF835DC18652FF7EF9EE" box="[467,536,1607,1633]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="4" pageNumber="434">4A–D</figureCitation>
)
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<treatmentCitation id="0AA310C9FFFEFF835C85869EFF6CF92C" author="Alcock, A." box="[151,522,1675,1699]" page="186" pageId="4" pageNumber="434" year="1896">
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<emphasis id="B976EACAFFFEFF835C85869EFC2FF92C" box="[151,329,1675,1699]" italics="true" pageId="4" pageNumber="434">Ebalia sagittifera</emphasis>
<bibRefCitation id="EF934B29FFFEFF835D5C869EFF6CF92C" author="Alcock, A." box="[334,522,1675,1699]" pageId="4" pageNumber="434" pagination="1 - 296" refId="ref6079" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2 The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, India, 65, 1 - 296. https: // doi. org / 10.5962 / bhl. title. 16033" type="journal article" year="1896">Alcock, 1896: 186</bibRefCitation>
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(in key), 188, 189— 
<bibRefCitation id="EF934B29FFFEFF835ECF869EFE83F92C" author="Alcock, A. &amp; Anderson, A. R. S." box="[733,997,1675,1699]" pageId="4" pageNumber="434" refId="ref6131" refString="Alcock, A. &amp; Anderson, A. R. S. (1897) s. n. In: Illustrations of the zoology of the Royal Indian Marine Surveying Steamer Investigator: Crustacea. Part V. Office of the Superintendent of Government Printing, Calcutta, pls. XXVIII - XXXII. https: // doi. org / 10.5962 / bhl. title. 64230" type="book" year="1897">Alcock &amp; Anderson 1897</bibRefCitation>
: pl 29, fig 9— 
<treatmentCitation id="0AA310C9FFFEFF83586A869EF86AF92C" author="Apel, M." box="[1144,1292,1675,1699]" page="50" pageId="4" pageNumber="434" year="2001">
<bibRefCitation id="EF934B29FFFEFF83586A869EF86AF92C" author="Apel, M." box="[1144,1292,1675,1699]" pageId="4" pageNumber="434" refId="ref6201" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguidea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes. Biologie und Informatik. Advisor: Prof. Dr. K. - D. Entian. PhD thesis, Frankfurt am Main, Johann Wolfgang Goethe-Universitat, Frankfurt, 260 pp." type="book" year="2001">Apel 2001: 50</bibRefCitation>
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.
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<treatmentCitationGroup id="AB1211F6FFFEFF835C8586BFFE8CF94E" box="[151,1002,1706,1730]" pageId="4" pageNumber="434">
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<taxonomicName id="4C024D5BFFFEFF835C8586BFFC30F94D" ID-CoL="7NXVH" authority="Sagittifera" authorityName="Leach" authorityYear="1817" box="[151,342,1706,1730]" class="Malacostraca" family="Leucosiidae" genus="Ebalia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="434" phylum="Arthropoda" rank="genus">Ebalia? Sagittifera</taxonomicName>
—
</emphasis>
<treatmentCitation id="0AA310C9FFFEFF835D7D86BFFF42F94D" author="Titgen, R." box="[367,548,1706,1730]" page="113" pageId="4" pageNumber="434" year="1982">
<bibRefCitation id="EF934B29FFFEFF835D7D86BFFF42F94D" author="Titgen, R." box="[367,548,1706,1730]" pageId="4" pageNumber="434" refId="ref7199" refString="Titgen, R. (1982) The systematics and ecology of the Decapods of Dubai, and their zoogeographic relationships to the Persian Gulf and the Western Indian Ocean. Ph. D. thesis, Texas A &amp; M University, College Station, Texas, 292 pp." type="book" year="1982">Titgen, 1982: 113</bibRefCitation>
</treatmentCitation>
, 247 (in list)— 
<treatmentCitation id="0AA310C9FFFEFF835EAF86BFFE11F94D" author="Cooper, R. T." box="[701,887,1706,1730]" page="173" pageId="4" pageNumber="434" year="1997">
<bibRefCitation id="EF934B29FFFEFF835EAF86BFFE11F94D" author="Cooper, R. T." box="[701,887,1706,1730]" pageId="4" pageNumber="434" pagination="155 - 179" refId="ref6359" refString="Cooper, R. T. (1997) Mangal-associated Brachyura (Ocypodidae, Grapsidae, Portunidae, Majidae, Xanthidae and Leucosiidae) from the north-eastern coastal Islands of Abu Dhabi, United Arab Emirates. Crustaceana, 70 (2), 155 - 179. https: // doi. org / 10.1163 / 156854097 X 00807" type="journal article" year="1997">Cooper 1997: 173</bibRefCitation>
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, 174, fig 7.
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</paragraph>
<paragraph id="8BBD36D8FFFEFF835C8586DCF879F96F" blockId="4.[151,1437,1675,1853]" box="[151,1311,1736,1761]" pageId="4" pageNumber="434">
<treatmentCitationGroup id="AB1211F6FFFEFF835C8586DCF879F96F" box="[151,1311,1736,1761]" pageId="4" pageNumber="434">
<emphasis id="B976EACAFFFEFF835C8586DCFCE1F96E" box="[151,391,1737,1761]" italics="true" pageId="4" pageNumber="434">
<taxonomicName id="4C024D5BFFFEFF835C8586DCFC08F96E" ID-CoL="76YKT" authorityName="Miers" authorityYear="1884" box="[151,366,1737,1761]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="rectangularis">Philyra rectangularis</taxonomicName>
—
</emphasis>
<treatmentCitation id="0AA310C9FFFEFF835D9A86DCFF05F96E" author="Stephensen, K." box="[392,611,1736,1761]" page="87" pageId="4" pageNumber="434" year="1946">
<bibRefCitation id="EF934B29FFFEFF835D9A86DCFF05F96E" author="Stephensen, K." box="[392,611,1736,1761]" pageId="4" pageNumber="434" pagination="57 - 237" refId="ref7062" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf with an appendix: The male pleopoda of the Brachyura. Danish Scientific Investigations in Iran, 4, 57 - 237." type="journal article" year="1946">Stephensen, 1946: 87</bibRefCitation>
</treatmentCitation>
, 88, fig 15a, b— 
<treatmentCitation id="0AA310C9FFFEFF835F1F86DCFEA6F96E" author="Guinot, D." box="[781,960,1736,1761]" page="249" pageId="4" pageNumber="434" year="1967">
<bibRefCitation id="EF934B29FFFEFF835F1F86DCFEA6F96E" author="Guinot, D." box="[781,960,1736,1761]" pageId="4" pageNumber="434" pagination="238 - 352" refId="ref6610" refString="Guinot, D. (1967) La faune carcinologique (Crustacea Brachyura) de l'Ocean Indien Occidental et de la Mer Rouge. Catalogue, Remarques Biogeographiques et Bibliographie. Memoirs de l'Institut Fondamental d'Afrique Noir, 77, 238 - 352." type="journal article" year="1967">Guinot 1967: 249</bibRefCitation>
</treatmentCitation>
(in list)— 
<treatmentCitation id="0AA310C9FFFEFF83583886DCF87DF96F" author="Basson, P. W. &amp; Burchard, J. A. &amp; Hardy, J. T. &amp; Price, A. R. G." box="[1066,1307,1736,1761]" page="250" pageId="4" pageNumber="434" year="1977">
<bibRefCitation id="EF934B29FFFEFF83583886DCF87DF96F" author="Basson, P. W. &amp; Burchard, J. A. &amp; Hardy, J. T. &amp; Price, A. R. G." box="[1066,1307,1736,1761]" pageId="4" pageNumber="434" refId="ref6259" refString="Basson, P. W., Burchard, J. A., Hardy, J. T. &amp; Price, A. R. G. (1977) Biotopes of the Western Arabian Gulf: marine life and environments of Saudi Arabia. Department of Loss Prevention and Environmental Affairs, ARAMCO, Dhahran, 284 pp." type="book" year="1977">
Basson 
<emphasis id="B976EACAFFFEFF83586B86DCF9C0F96E" box="[1145,1190,1737,1761]" italics="true" pageId="4" pageNumber="434">et al</emphasis>
. 1977: 250
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBD36D8FFFEFF835C8586F2F9F7F970" blockId="4.[151,1437,1675,1853]" box="[151,1169,1767,1791]" pageId="4" pageNumber="434">
<treatmentCitationGroup id="AB1211F6FFFEFF835C8586F2F9F7F970" box="[151,1169,1767,1791]" pageId="4" pageNumber="434">
<emphasis id="B976EACAFFFEFF835C8586F2FC01F970" box="[151,359,1767,1791]" italics="true" pageId="4" pageNumber="434">
<taxonomicName id="4C024D5BFFFEFF835C8586F2FC2BF970" ID-CoL="76YLY" authorityName="Ghani &amp; Tirmizi" authorityYear="1995" box="[151,333,1767,1791]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="concinnus">Philyra concinnus</taxonomicName>
—
</emphasis>
<treatmentCitation id="0AA310C9FFFEFF835D7586F2FFDEF970" author="Ghani, N. &amp; Tirmizi, N. M." box="[359,696,1767,1791]" page="143" pageId="4" pageNumber="434" year="1995">
<bibRefCitation id="EF934B29FFFEFF835D7586F2FFDEF970" author="Ghani, N. &amp; Tirmizi, N. M." box="[359,696,1767,1791]" pageId="4" pageNumber="434" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi, 1995: 143–147</bibRefCitation>
</treatmentCitation>
, fig 2a–e (in part, material from Persian Gulf).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBD36D8FFFEFF835C858713FCF9F8B2" blockId="4.[151,1437,1675,1853]" pageId="4" pageNumber="434">
<treatmentCitationGroup id="AB1211F6FFFEFF835C858713FCF9F8B2" pageId="4" pageNumber="434">
<emphasis id="B976EACAFFFEFF835C858713FC0DF891" box="[151,363,1798,1822]" italics="true" pageId="4" pageNumber="434">
<taxonomicName id="4C024D5BFFFEFF835C858713FC37F891" ID-CoL="4GDVC" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[151,337,1798,1822]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="sagittifera">Philyra sagittifera</taxonomicName>
—
</emphasis>
<treatmentCitation id="0AA310C9FFFEFF835D798713FF4DF891" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[363,555,1798,1822]" page="93" pageId="4" pageNumber="434" year="2008">
<bibRefCitation id="EF934B29FFFEFF835D798713FF4DF891" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[363,555,1798,1822]" pageId="4" pageNumber="434" pagination="1 - 286" refId="ref6941" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="B976EACAFFFEFF835D818712FCA4F891" box="[403,450,1798,1822]" italics="true" pageId="4" pageNumber="434">et al</emphasis>
. 2008: 93
</bibRefCitation>
</treatmentCitation>
— 
<treatmentCitation id="0AA310C9FFFEFF835E548713FF8AF891" author="Galil, B. S." box="[582,748,1798,1822]" page="281" pageId="4" pageNumber="434" year="2009">
<bibRefCitation id="EF934B29FFFEFF835E548713FF8AF891" author="Galil, B. S." box="[582,748,1798,1822]" pageId="4" pageNumber="434" pagination="279 - 320" refId="ref6475" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009: 281</bibRefCitation>
</treatmentCitation>
, tab 1— 
<treatmentCitation id="0AA310C9FFFEFF835F568713F911F891" author="Naderloo, R. &amp; Turkay, M." box="[836,1143,1798,1822]" page="32" pageId="4" pageNumber="434" year="2012">
<bibRefCitation id="EF934B29FFFEFF835F568713F911F891" author="Naderloo, R. &amp; Turkay, M." box="[836,1143,1798,1822]" pageId="4" pageNumber="434" pagination="1 - 67" refId="ref6831" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 32</bibRefCitation>
</treatmentCitation>
— 
<treatmentCitation id="0AA310C9FFFEFF8358838713F805F891" author="Naderloo, R." box="[1169,1379,1798,1822]" page="112" pageId="4" pageNumber="434" year="2017">
<bibRefCitation id="EF934B29FFFEFF8358838713F805F891" author="Naderloo, R." box="[1169,1379,1798,1822]" pageId="4" pageNumber="434" refId="ref6733" refString="Naderloo, R. (2017) Atlas of Crabs of the Persian Gulf. Springer International Publishing, Cham, 444 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017: 112</bibRefCitation>
</treatmentCitation>
, figs. 14.35d, 14.38, 14.41.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3186553FFFEFF825C85877EF911FEEA" lastPageId="5" lastPageNumber="435" pageId="4" pageNumber="434" type="materials_examined">
<paragraph id="8BBD36D8FFFEFF835C85877EF84AF826" blockId="4.[151,1437,1898,2034]" pageId="4" pageNumber="434">
<emphasis id="B976EACAFFFEFF835C85877EFC5BF80B" bold="true" box="[151,317,1898,1925]" pageId="4" pageNumber="434">Type material</emphasis>
. 
<materialsCitation id="3B6A3C85FFFEFF835D59877EFC34F826" ID-GBIF-Occurrence="4176773315" collectorName="F. W. Townsend." country="Pakistan" location="Pakistan" pageId="4" pageNumber="434" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54B9887AFFFEFF835D59877EFCDDF80A" box="[331,443,1899,1925]" pageId="4" pageNumber="434" type="lectotype">Lectotype</typeStatus>
(herein selected): ZSI C910/10, male (CL 5.0 mm, CW 
<quantity id="4CFA9B3DFFFEFF83583F877EF9E0F80A" box="[1069,1158,1899,1926]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="434" unit="mm" value="4.5">4.5 mm</quantity>
), Karachi, 
<collectingCountry id="F3157648FFFEFF8358ED877EF807F80A" box="[1279,1377,1899,1925]" name="Pakistan" pageId="4" pageNumber="434">Pakistan</collectingCountry>
, coll. 
<collectorName id="26F7530EFFFEFF835C85879AFC34F826" box="[151,338,1935,1961]" pageId="4" pageNumber="434">F. W. Townsend.</collectorName>
</materialsCitation>
<typeStatus id="54B9887AFFFEFF835D4A879AFC93F826" box="[344,501,1935,1961]" pageId="4" pageNumber="434" type="paralectotype">Paralectotype</typeStatus>
: ZSI C910/10, female (CL 6.0 mm, CW 
<quantity id="4CFA9B3DFFFEFF835FAE879AF970F826" box="[956,1046,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="434" unit="mm" value="5.7">5.7 mm</quantity>
), same data as lectotype.
</paragraph>
<paragraph id="8BBD36D8FFFEFF825CD587A7F911FEEA" blockId="4.[151,1437,1898,2034]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="435" pageId="4" pageNumber="434">
<emphasis id="B976EACAFFFEFF835CD587A7FFCCF842" bold="true" box="[199,682,1970,1997]" pageId="4" pageNumber="434">
Other material examined. 
<emphasis id="B976EACAFFFEFF835E1887A6FFC7F842" bold="true" box="[522,673,1970,1997]" italics="true" pageId="4" pageNumber="434">Persian Gulf</emphasis>
:
</emphasis>
<materialsCitation id="3B6A3C85FFFEFF835EA687A6FFE3F87E" ID-GBIF-Occurrence="4176773312" collectingDate="1993-03-03" collectionCode="ZUMC" collectorName="G. Thorson" country="Iran" elevation="6" location="Bushehr" pageId="4" pageNumber="434" specimenCode="CRU885" specimenCount="1" specimenCount-male="1" stateProvince="Bushehr">
<collectionCode id="ED13AE1DFFFEFF835EA687A6FE63F842" box="[692,773,1971,1997]" pageId="4" pageNumber="434">ZUMC</collectionCode>
<specimenCode id="DBA49EA3FFFEFF835F1C87A6FE13F841" box="[782,885,1971,1998]" collectionCode="CRU" pageId="4" pageNumber="434">CRU885</specimenCode>
, 
<specimenCount id="9D04FD51FFFEFF835F9087A6FEB6F842" box="[898,976,1971,1997]" pageId="4" pageNumber="434" type="male">1 male</specimenCount>
(CL 
<quantity id="4CFA9B3DFFFEFF83581D87A6F90CF842" box="[1039,1130,1971,1997]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="434" unit="mm" value="4.2">4.2 mm</quantity>
, CW 
<quantity id="4CFA9B3DFFFEFF8358BC87A6F86DF842" box="[1198,1291,1971,1997]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="434" unit="mm" value="3.9">3.9 mm</quantity>
), 
<elevation id="002FD1EBFFFEFF83593287A6F8FEF842" box="[1312,1432,1971,1997]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="434" unit="m" value="6.0">
<quantity id="4CFA9B3DFFFEFF83593287A6F828F842" box="[1312,1358,1971,1997]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="434" unit="m" value="6.0">6 m</quantity>
depth
</elevation>
, 
<collectingRegion id="49C6F83AFFFEFF835C8587C2FD90F87E" box="[151,246,2007,2033]" country="Iran" name="Bushehr" pageId="4" pageNumber="434">Bushehr</collectingRegion>
, 
<collectingCountry id="F3157648FFFEFF835D1087C2FC54F87E" box="[258,306,2007,2033]" name="Iran" pageId="4" pageNumber="434">Iran</collectingCountry>
, 
<date id="FFBC1018FFFEFF835D2F87C2FCD8F87E" box="[317,446,2007,2033]" pageId="4" pageNumber="434" value="1993-03-03">
<collectingDate id="EFF8E9F0FFFEFF835D2F87C2FCD8F87E" box="[317,446,2007,2033]" pageId="4" pageNumber="434" value="1993-03-03">03.03.1993</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFEFF835E1087C2FFE3F87E" box="[514,645,2007,2033]" pageId="4" pageNumber="434">G. Thorson</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B6A3C85FFFEFF835E8387C2F8FEF87E" ID-GBIF-Occurrence="4176773309" box="[657,1432,2007,2034]" collectionCode="ZMUC" country="Iran" elevation="6" location="Bushehr" pageId="4" pageNumber="434" specimenCode="CRU-885" specimenCount="2" specimenCount-male="2" stateProvince="Bushehr">
<collectionCode id="ED13AE1DFFFEFF835E8387C2FF84F87E" box="[657,738,2007,2033]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="434">ZMUC</collectionCode>
<specimenCode id="DBA49EA3FFFEFF835EFB87C2FE3FF87D" box="[745,857,2007,2034]" collectionCode="CRU-" pageId="4" pageNumber="434">CRU-885</specimenCode>
, 
<specimenCount id="9D04FD51FFFEFF835F7687C2FED8F87E" box="[868,958,2007,2033]" pageId="4" pageNumber="434" type="male">2 males</specimenCount>
, 3 ovigerous females, 
<elevation id="002FD1EBFFFEFF8358A887C2F84BF87E" box="[1210,1325,2007,2033]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="434" unit="m" value="6.0">
<quantity id="4CFA9B3DFFFEFF8358A887C2F983F87E" box="[1210,1253,2007,2033]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="434" unit="m" value="6.0">6 m</quantity>
depth
</elevation>
, 
<collectingRegion id="49C6F83AFFFEFF83592B87C2F8FEF87E" box="[1337,1432,2007,2033]" country="Iran" name="Bushehr" pageId="4" pageNumber="434">Bushehr</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF825C858082F997FF3E" ID-GBIF-Occurrence="4176773308" box="[151,1265,151,177]" collectingDate="1993-03-03" collectionCode="ZUTC" collectorName="G. Thorson" country="Iran" location="Bandar-Abbas" pageId="5" pageNumber="435" specimenCode="ZUTC 1339" specimenCount="1" specimenCount-juvenile="1">
<collectingCountry id="F3157648FFFFFF825C858082FDA1FF3E" box="[151,199,151,177]" name="Iran" pageId="5" pageNumber="435">Iran</collectingCountry>
, 
<date id="FFBC1018FFFFFF825CC18082FC32FF3E" box="[211,340,151,177]" pageId="5" pageNumber="435" value="1993-03-03">
<collectingDate id="EFF8E9F0FFFFFF825CC18082FC32FF3E" box="[211,340,151,177]" pageId="5" pageNumber="435" value="1993-03-03">03.03.1993</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFFFF825D8B8082FF7BFF3E" box="[409,541,151,177]" pageId="5" pageNumber="435">G. Thorson</collectorName>
; 
<specimenCode id="DBA49EA3FFFFFF825E3B8082FFD3FF3E" box="[553,693,151,177]" collectionCode="ZUTC" pageId="5" pageNumber="435">ZUTC 1339</specimenCode>
, 
<specimenCount id="9D04FD51FFFFFF825ED38082FE52FF3E" box="[705,820,151,177]" pageId="5" pageNumber="435" type="juvenile">1 juvenile</specimenCount>
, muddy-sandy flat, E of 
<location id="8EDD6003FFFFFF82585F8082F997FF3E" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF82585F8082F997FF3E" box="[1101,1265,151,177]" country="Iran" name="Bandar-Abbas" pageId="5" pageNumber="435">Bandar-Abbas</location>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF8258EF8082F995FF5A" ID-GBIF-Occurrence="4176773306" collectionCode="ZUTC" collectorName="Reza Naderloo &amp; Abbas Kazemi" country="Iran" latitude="27.183332" location="Bandar-Khamir" longLatPrecision="1238" longitude="56.35" pageId="5" pageNumber="435" specimenCode="ZUTC 1340" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F3157648FFFFFF8258EF8082F84BFF3E" box="[1277,1325,151,177]" name="Iran" pageId="5" pageNumber="435">Iran</collectingCountry>
, 
<geoCoordinate id="EE36501FFFFFFF82592B808DF8FEFF3C" box="[1337,1432,152,179]" degrees="27" direction="north" minutes="11" orientation="latitude" pageId="5" pageNumber="435" precision="925" value="27.183332">27°11′N</geoCoordinate>
, 
<geoCoordinate id="EE36501FFFFFFF825C8580A9FD92FF58" box="[151,244,188,215]" degrees="56" direction="east" minutes="21" orientation="longitude" pageId="5" pageNumber="435" precision="925" value="56.35">56°21′E</geoCoordinate>
, coll. 
<collectorName id="26F7530EFFFFFF825D2480AEFCB9FF5A" box="[310,479,187,213]" pageId="5" pageNumber="435">Reza Naderloo</collectorName>
, 
<collectorName id="26F7530EFFFFFF825DFA80AEFFF7FF5A" box="[488,657,187,213]" pageId="5" pageNumber="435">Abbas Kazemi</collectorName>
; 
<specimenCode id="DBA49EA3FFFFFF825E8980AEFE43FF5A" box="[667,805,187,213]" collectionCode="ZUTC" pageId="5" pageNumber="435">ZUTC 1340</specimenCode>
, 
<specimenCount id="9D04FD51FFFFFF825F3D80AEFE1AFF5A" box="[815,892,187,213]" pageId="5" pageNumber="435" type="male">1 male</specimenCount>
, muddy flat, E of 
<location id="8EDD6003FFFFFF82585080AEF995FF5A" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF82585080AEF995FF5A" box="[1090,1267,187,213]" country="Iran" latitude="27.183332" longLatPrecision="1238" longitude="56.35" name="Bandar-Khamir" pageId="5" pageNumber="435">Bandar-Khamir</location>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF8258EF80AEF90CFF76" ID-GBIF-Occurrence="4176773304" collectionCode="SMF" collectorName="Reza Naderloo" country="Iran" latitude="27.933332" location="Al-Khobar" longLatPrecision="1234" longitude="55.6" municipality="Tarut Bay" pageId="5" pageNumber="435" specimenCount="4" specimenCount-juvenile="3" specimenCount-male="1">
<collectingCountry id="F3157648FFFFFF8258EF80AEF84BFF5A" box="[1277,1325,187,213]" name="Iran" pageId="5" pageNumber="435">Iran</collectingCountry>
, 
<geoCoordinate id="EE36501FFFFFFF82592A80A9F8FEFF58" box="[1336,1432,188,215]" degrees="27" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="435" precision="925" value="27.933332">27°56′N</geoCoordinate>
, 
<geoCoordinate id="EE36501FFFFFFF825C8580F5FD92FF74" box="[151,244,224,251]" degrees="55" direction="east" minutes="36" orientation="longitude" pageId="5" pageNumber="435" precision="925" value="55.6">55°36′E</geoCoordinate>
, coll. 
<collectorName id="26F7530EFFFFFF825D2B80CAFC83FF76" box="[313,485,223,249]" pageId="5" pageNumber="435">Reza Naderloo</collectorName>
; 
<collectionCode id="ED13AE1DFFFFFF825DE280CAFF4FFF76" box="[496,553,223,249]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="435">SMF</collectionCode>
, 
<specimenCount id="9D04FD51FFFFFF825E2780CAFFE5FF76" box="[565,643,223,249]" pageId="5" pageNumber="435" type="male">1 male</specimenCount>
, 
<specimenCount id="9D04FD51FFFFFF825E8280CAFE6CFF76" box="[656,778,223,249]" pageId="5" pageNumber="435" type="juvenile">3 juveniles</specimenCount>
(), N. of 
<location id="8EDD6003FFFFFF825F6680CAFE89FF76" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF825F6680CAFE89FF76" box="[884,1007,223,249]" country="Iran" latitude="27.933332" longLatPrecision="1234" longitude="55.6" municipality="Tarut Bay" name="Al-Khobar" pageId="5" pageNumber="435">Al-Khobar</location>
, 
<collectingMunicipality id="6BD9ACA2FFFFFF825FE880CAF90CFF76" box="[1018,1130,223,249]" pageId="5" pageNumber="435">Tarut Bay</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF82586480CAF95EFE92" ID-GBIF-Occurrence="4176773310" collectingDate="1993-03-19" collectionCode="SMF" collectorName="Michael Apel" country="Saudi Arabia" location="Al-Khobar" municipality="Tarut Bay" pageId="5" pageNumber="435" specimenCount="8" specimenCount-male="8">
<collectingCountry id="F3157648FFFFFF82586480CAF86DFF76" box="[1142,1291,223,249]" name="Saudi Arabia" pageId="5" pageNumber="435">Saudi Arabia</collectingCountry>
, 
<date id="FFBC1018FFFFFF82590580CAF8F1FF76" box="[1303,1431,223,249]" pageId="5" pageNumber="435" value="1993-03-19">
<collectingDate id="EFF8E9F0FFFFFF82590580CAF8F1FF76" box="[1303,1431,223,249]" pageId="5" pageNumber="435" value="1993-03-19">19.03.1993</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFFFF825CDC8116FC0FFE92" box="[206,361,259,285]" pageId="5" pageNumber="435">Michael Apel</collectorName>
; 
<collectionCode id="ED13AE1DFFFFFF825D618116FCCAFE92" box="[371,428,259,285]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="435">SMF</collectionCode>
, 
<specimenCount id="9D04FD51FFFFFF825DA48116FF69FE92" box="[438,527,259,285]" pageId="5" pageNumber="435" type="male">8 males</specimenCount>
, 6 ovigerous females, N. of 
<location id="8EDD6003FFFFFF825F568116FED9FE92" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF825F568116FED9FE92" box="[836,959,259,285]" country="Saudi Arabia" municipality="Tarut Bay" name="Al-Khobar" pageId="5" pageNumber="435">Al-Khobar</location>
, 
<collectingMunicipality id="6BD9ACA2FFFFFF825FDB8116F95EFE92" box="[969,1080,259,285]" pageId="5" pageNumber="435">Tarut Bay</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF8258508116FF93FECE" ID-GBIF-Occurrence="4176773314" collectingDate="1993-03-25" collectionCode="SMF" collectorName="Michael Apel" country="Saudi Arabia" location="Jubail" pageId="5" pageNumber="435" specimenCount="1" specimenCount-juvenile="1">
<collectingCountry id="F3157648FFFFFF8258508116F9B0FE92" box="[1090,1238,259,285]" name="Saudi Arabia" pageId="5" pageNumber="435">Saudi Arabia</collectingCountry>
, 
<date id="FFBC1018FFFFFF8258F28117F806FE92" box="[1248,1376,258,285]" pageId="5" pageNumber="435" value="1993-03-25">
<collectingDate id="EFF8E9F0FFFFFF8258F28117F806FE92" box="[1248,1376,258,285]" pageId="5" pageNumber="435" value="1993-03-25">25.03.1993</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFFFF825C858132FC54FECE" box="[151,306,295,321]" pageId="5" pageNumber="435">Michael Apel</collectorName>
; 
<collectionCode id="ED13AE1DFFFFFF825D2F8132FC10FECE" box="[317,374,295,321]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="435">SMF</collectionCode>
, 
<specimenCount id="9D04FD51FFFFFF825D928132FC94FECE" box="[384,498,295,321]" pageId="5" pageNumber="435" type="juvenile">1 juvenile</specimenCount>
, intertidal, N. of 
<location id="8EDD6003FFFFFF825EBD8132FF93FECE" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF825EBD8132FF93FECE" box="[687,757,295,321]" country="Saudi Arabia" name="Jubail" pageId="5" pageNumber="435">Jubail</location>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF825EED8132FF0CFEEA" ID-GBIF-Occurrence="4176773316" collectingDate="1993-04-30" collectionCode="SMF" collectorName="Michael Apel &amp; Qurmah Channel, N." country="Saudi Arabia" elevation="6" location="Jubail" pageId="5" pageNumber="435" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F3157648FFFFFF825EED8132FEF5FECE" box="[767,915,295,321]" name="Saudi Arabia" pageId="5" pageNumber="435">Saudi Arabia</collectingCountry>
, 
<date id="FFBC1018FFFFFF825F8C8132F979FECE" box="[926,1055,295,321]" pageId="5" pageNumber="435" value="1993-04-30">
<collectingDate id="EFF8E9F0FFFFFF825F8C8132F979FECE" box="[926,1055,295,321]" pageId="5" pageNumber="435" value="1993-04-30">30.04.1993</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFFFF8258738132F99AFECE" box="[1121,1276,295,321]" pageId="5" pageNumber="435">Michael Apel</collectorName>
; 
<collectionCode id="ED13AE1DFFFFFF8259158132F826FECE" box="[1287,1344,295,321]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="435">SMF</collectionCode>
, 
<specimenCount id="9D04FD51FFFFFF8259588132F8FEFECE" box="[1354,1432,295,321]" pageId="5" pageNumber="435" type="male">1 male</specimenCount>
, 
<elevation id="002FD1EBFFFFFF825C85815EFC6CFEEA" box="[151,266,331,357]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="435" unit="m" value="6.0">
<quantity id="4CFA9B3DFFFFFF825C85815EFDA4FEEA" box="[151,194,331,357]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="435" unit="m" value="6.0">6 m</quantity>
depth
</elevation>
, 
<collectorName id="26F7530EFFFFFF825D07815EFC99FEEA" box="[277,511,331,357]" pageId="5" pageNumber="435">Qurmah Channel, N.</collectorName>
of 
<location id="8EDD6003FFFFFF825E36815EFF0CFEEA" LSID="urn:lsid:plazi:treatment:03AB87CEFFFEFF815C858637FFF3FD0A:8EDD6003FFFFFF825E36815EFF0CFEEA" box="[548,618,331,357]" country="Saudi Arabia" name="Jubail" pageId="5" pageNumber="435">Jubail</location>
</materialsCitation>
, 
<materialsCitation id="3B6A3C85FFFFFF825E67815EF911FEEA" ID-GBIF-Occurrence="4176773302" box="[629,1143,330,357]" collectingDate="1992-05-10" collectorName="Michael Apel." country="Saudi Arabia" location="Saudi Arabia" pageId="5" pageNumber="435" specimenCount="1">
<collectingCountry id="F3157648FFFFFF825E67815EFE6FFEEA" box="[629,777,331,357]" name="Saudi Arabia" pageId="5" pageNumber="435">Saudi Arabia</collectingCountry>
, 
<date id="FFBC1018FFFFFF825F06815FFEF2FEEA" box="[788,916,330,357]" pageId="5" pageNumber="435" value="1992-05-10">
<collectingDate id="EFF8E9F0FFFFFF825F06815FFEF2FEEA" box="[788,916,330,357]" pageId="5" pageNumber="435" value="1992-05-10">05.10.1992</collectingDate>
</date>
, coll. 
<collectorName id="26F7530EFFFFFF825FCA815EF911FEEA" box="[984,1143,331,357]" pageId="5" pageNumber="435">Michael Apel.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3186553FFFFFF825CD5817BFC33F932" pageId="5" pageNumber="435" type="description">
<paragraph id="8BBD36D8FFFFFF825CD5817BF925FC9A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
<emphasis id="B976EACAFFFFFF825CD5817BFC3FFE07" bold="true" box="[199,345,366,392]" pageId="5" pageNumber="435">Description.</emphasis>
Carapace hexagonal (
<figureCitation id="13392A5DFFFFFF825E40817AFFABFE06" box="[594,717,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
), slightly longer than broad. Dorsal surface smooth, three broad tuberculated and granular ridges, uniting to form &quot;broad-arrow&quot; point directed forward, median ridge from progastric region to intestinal region; lateral ridges extending backwards and downwards to branchial region, parallel to anterolateral margins with their ends projecting beyond postero-lateral borders in forming broad granular denticles; regions relatively distinct, with elevated gastric, cardiac, branchial, intestinal regions; hepatic region excavated, forming depression; upper, lower margins beaded, originating from middle of anterolateral margin, not merging anteriorly, broadly triangular tooth projecting on anterior half of lower margin of anterolateral border (
<figureCitation id="13392A5DFFFFFF8259068252F8EBFDEE" box="[1300,1421,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
); anterolateral margin merging with posterolateral with broad triangular tooth; anterolateral, posterolateral, posterior margins beaded; epibranchial junction with right angle; posterolateral margin slightly sinuous, convex (
<figureCitation id="13392A5DFFFFFF825907829AF8EBFD26" box="[1301,1421,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
); front bilobed; epistome visible beyond frontal margin from dorsal view; posterior margin convex, lateral sides with knob-like dorsoventrally flattened teeth with rounded tips, small median tooth on posterior margin (
<figureCitation id="13392A5DFFFFFF82590282C2F8E8FD7E" box="[1296,1422,727,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
). Anterior margin of endostome almost reaching as far forwards as inhalent channels.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD5830AFC3FFC2A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
Maxilliped 3 (
<figureCitation id="13392A5DFFFFFF825D7A830AFCAAFCB6" box="[360,460,799,825]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Figs. 1D</figureCitation>
, 
<figureCitation id="13392A5DFFFFFF825DCB830AFC9AFCB6" box="[473,508,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="5" pageNumber="435">4A</figureCitation>
) smooth, entirely covering endostome; merus with notch, 0.9× as long as ischium along inner margin; ischium 1.4× longer than wide; propodus, dactylus not visible in external view when reposed, articulating on inner surface of merus. Exopod outer margin convex, setose, much longer than wide, almost twice as long as merus.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD583BAF969FBD6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
Chelipeds (
<figureCitation id="13392A5DFFFFFF825D5583BAFCD8FC46" box="[327,446,943,969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
) equal, longer than half of carapace length, granulated. Merus triquetral in cross-section with edges raised, granular; both inner, outer margins with tuberculated ridge, tubercles larger on inner margin, patch of tubercles on upper proximal and distal end. Carpus, propodus have raised row of granules along inner edge of upper surface. Fingers slender, about two-thirds length of propodus, occlusal margins of fingers toothed, with scattered setae, median shallow canal extending to distal length in both fingers.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD58477FEBEFB2E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
P2–P5 (
<figureCitation id="13392A5DFFFFFF825D0C8476FCF4FBF2" box="[286,402,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A, B</figureCitation>
) subcylindrical; total length of all pairs almost similar, merus, carpus glabrous, merus longest; carpus, propodus almost equal in length; dactylus with tapering distal end.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD584BEFE4BFA82" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
Thoracic sternum (
<figureCitation id="13392A5DFFFFFF825DB684BEFC98FB4A" box="[420,510,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1D</figureCitation>
) tuberculated, slightly concave, tubercules prominent at margins. Sternites 1–3 completely fused without any trace of sutures; sternite 3 separated from sternite 4 by shallow groove; sternite 4–6 almost similar in width; sternite 7 completely tuberculated.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD58502F947FAF6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
Male pleon long, narrow (
<figureCitation id="13392A5DFFFFFF825DFC8502FF20FABE" box="[494,582,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1D</figureCitation>
), smooth, with three articulating parts (
<figureCitation id="13392A5DFFFFFF8258168502F93BFABE" box="[1028,1117,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1D</figureCitation>
, 
<figureCitation id="13392A5DFFFFFF8258788502F9E8FABE" box="[1130,1166,1303,1329]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="5" pageNumber="435">4B</figureCitation>
): somite 1 free, narrow; somites 2–6 fused, immovable, partial suture visible between somites 5–6, posterolateral angles with small bulge, somite 6 without denticle; telson triangular, longer than broad, with curved apex.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD58596FF94FA6A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
Female pleon ovate; somite 1 free; somites 2–6 fused, suture clearly visible between somites 2 and 3, partial suture visible between somites 3–4 and 5–6, forming domed plate almost completely covering thoracic sternum. Telson longer than broad, with curved apex (
<figureCitation id="13392A5DFFFFFF825E9E85DEFF82FA6A" box="[652,740,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1590]" captionTargetId="figure-22@3.[151,1436,181,1590]" captionTargetPageId="3" captionText="FIGURE 2. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) dorsal view; (B) ventral view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254912" httpUri="https://zenodo.org/record/8254912/files/figure.png" pageId="5" pageNumber="435">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD585FAFE83F9DE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
G1 (
<figureCitation id="13392A5DFFFFFF825CE585FAFC09F986" box="[247,367,1519,1545]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="5" pageNumber="435">Fig. 4A–D</figureCitation>
) shaft long, slender, straight, apical process long, about 0.3 times as long as whole length, slightly sinuous-shape, with tapering ending, apical lobe with setae shorter than the length of the tip. Female gonopore (
<figureCitation id="13392A5DFFFFFF82597D8606FDDFF9DE" captionStart="FIGURE 5" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[160,1422,822,1692]" captionTargetId="figure-273@8.[151,1436,813,1700]" captionTargetPageId="8" captionText="FIGURE 5. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) frontal view of cephalothorax; (B) female gonopore (right side); Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) frontal view of cephalothorax; (D) female gonopore (right side)." figureDoi="http://doi.org/10.5281/zenodo.8254922" httpUri="https://zenodo.org/record/8254922/files/figure.png" pageId="5" pageNumber="435">Fig. 5B</figureCitation>
) on inner anterior edge of sternite 5, oval in shape with narrow opening.
</paragraph>
<paragraph id="8BBD36D8FFFFFF825CD5864FFC33F932" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
<emphasis id="B976EACAFFFFFF825CD5864FFC5EF9FB" bold="true" box="[199,312,1626,1652]" pageId="5" pageNumber="435">Variation</emphasis>
. The carapace of female resembles with that of male, except for lateral sides of posterior margin having smaller knob-like dorsoventrally flattened teeth, small median tooth on posterior margin less pronounced or absent (
<figureCitation id="13392A5DFFFFFF825CFC86B6FC2EF932" box="[238,328,1699,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1590]" captionTargetId="figure-22@3.[151,1436,181,1590]" captionTargetPageId="3" captionText="FIGURE 2. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) dorsal view; (B) ventral view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254912" httpUri="https://zenodo.org/record/8254912/files/figure.png" pageId="5" pageNumber="435">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186553FFFFFF825CD586D3FCF1F8C2" pageId="5" pageNumber="435" type="distribution">
<paragraph id="8BBD36D8FFFFFF825CD586D3FCF1F8C2" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
<emphasis id="B976EACAFFFFFF825CD586D3FC07F96F" bold="true" box="[199,353,1734,1760]" pageId="5" pageNumber="435">Distribution.</emphasis>
The species is recorded so far from 
<collectingCountry id="F3157648FFFFFF825EE186D2FE34F96E" box="[755,850,1735,1761]" name="Pakistan" pageId="5" pageNumber="435">Pakistan</collectingCountry>
(
<bibRefCitation id="EF934B29FFFFFF825F4D86D2FE92F96E" author="Alcock, A." box="[863,1012,1735,1761]" pageId="5" pageNumber="435" pagination="1 - 296" refId="ref6079" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2 The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, India, 65, 1 - 296. https: // doi. org / 10.5962 / bhl. title. 16033" type="journal article" year="1896">Alcock 1896</bibRefCitation>
), the Persian Gulf [
<bibRefCitation id="EF934B29FFFFFF8258DA86D2F8FAF96E" author="Stephensen, K." box="[1224,1436,1735,1761]" pageId="5" pageNumber="435" pagination="57 - 237" refId="ref7062" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf with an appendix: The male pleopoda of the Brachyura. Danish Scientific Investigations in Iran, 4, 57 - 237." type="journal article" year="1946">Stephensen (1946)</bibRefCitation>
as 
<taxonomicName id="4C024D5BFFFFFF825CA686FEFCC2F88A" authorityName="Miers" authorityYear="1884" box="[180,420,1771,1797]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B976EACAFFFFFF825CA686FEFCC2F88A" box="[180,420,1771,1797]" italics="true" pageId="5" pageNumber="435">Philyra rectangularis</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EF934B29FFFFFF825DA386FEFFE8F88A" author="Basson, P. W. &amp; Burchard, J. A. &amp; Hardy, J. T. &amp; Price, A. R. G." box="[433,654,1771,1797]" pageId="5" pageNumber="435" refId="ref6259" refString="Basson, P. W., Burchard, J. A., Hardy, J. T. &amp; Price, A. R. G. (1977) Biotopes of the Western Arabian Gulf: marine life and environments of Saudi Arabia. Department of Loss Prevention and Environmental Affairs, ARAMCO, Dhahran, 284 pp." type="book" year="1977">
Basson 
<emphasis id="B976EACAFFFFFF825E1586FEFF51F88A" box="[519,567,1771,1797]" italics="true" pageId="5" pageNumber="435">et al</emphasis>
. (1977)
</bibRefCitation>
as 
<taxonomicName id="4C024D5BFFFFFF825EA386FEFE07F88A" authorityName="Miers" authorityYear="1884" box="[689,865,1771,1797]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B976EACAFFFFFF825EA386FEFE07F88A" box="[689,865,1771,1797]" italics="true" pageId="5" pageNumber="435">P. rectangularis</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EF934B29FFFFFF825F7C86FEF960F88A" author="Titgen, R." box="[878,1030,1771,1797]" pageId="5" pageNumber="435" refId="ref7199" refString="Titgen, R. (1982) The systematics and ecology of the Decapods of Dubai, and their zoogeographic relationships to the Persian Gulf and the Western Indian Ocean. Ph. D. thesis, Texas A &amp; M University, College Station, Texas, 292 pp." type="book" year="1982">Titgen (1982)</bibRefCitation>
as 
<taxonomicName id="4C024D5BFFFFFF82583B86FEF98AF88A" authorityName="Alcock" authorityYear="1896" box="[1065,1260,1771,1797]" class="Malacostraca" family="Leucosiidae" genus="Ebalia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="sagittifera">
<emphasis id="B976EACAFFFFFF82583B86FEF98AF88A" box="[1065,1260,1771,1797]" italics="true" pageId="5" pageNumber="435">Ebalia sagittifera</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EF934B29FFFFFF8258E886FEF8FAF88A" author="Cooper, R. T." box="[1274,1436,1771,1797]" pageId="5" pageNumber="435" pagination="155 - 179" refId="ref6359" refString="Cooper, R. T. (1997) Mangal-associated Brachyura (Ocypodidae, Grapsidae, Portunidae, Majidae, Xanthidae and Leucosiidae) from the north-eastern coastal Islands of Abu Dhabi, United Arab Emirates. Crustaceana, 70 (2), 155 - 179. https: // doi. org / 10.1163 / 156854097 X 00807" type="journal article" year="1997">Cooper (1997)</bibRefCitation>
as 
<taxonomicName id="4C024D5BFFFFFF825CA7871AFC20F8A6" authorityName="Alcock" authorityYear="1896" box="[181,326,1807,1833]" class="Malacostraca" family="Leucosiidae" genus="Ebalia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="sagittifera">
<emphasis id="B976EACAFFFFFF825CA7871AFC20F8A6" box="[181,326,1807,1833]" italics="true" pageId="5" pageNumber="435">E. sagittifera</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EF934B29FFFFFF825D41871AFCBCF8A6" author="Apel, M." box="[339,474,1807,1833]" pageId="5" pageNumber="435" refId="ref6201" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguidea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes. Biologie und Informatik. Advisor: Prof. Dr. K. - D. Entian. PhD thesis, Frankfurt am Main, Johann Wolfgang Goethe-Universitat, Frankfurt, 260 pp." type="book" year="2001">Apel (2001)</bibRefCitation>
as 
<taxonomicName id="4C024D5BFFFFFF825DEC871AFFE9F8A6" authorityName="Alcock" authorityYear="1896" box="[510,655,1807,1833]" class="Malacostraca" family="Leucosiidae" genus="Ebalia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="sagittifera">
<emphasis id="B976EACAFFFFFF825DEC871AFFE9F8A6" box="[510,655,1807,1833]" italics="true" pageId="5" pageNumber="435">E. sagittifera</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EF934B29FFFFFF825E8C871AFEB9F8A6" author="Naderloo, R. &amp; Turkay, M." box="[670,991,1807,1833]" pageId="5" pageNumber="435" pagination="1 - 67" refId="ref6831" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo and Türkay (2012)</bibRefCitation>
; Nadeloo (2017)] and the Gulf of 
<collectingCountry id="F3157648FFFFFF82594A871AF8FAF8A6" box="[1368,1436,1807,1833]" name="Oman" pageId="5" pageNumber="435">Oman</collectingCountry>
(
<bibRefCitation id="EF934B29FFFFFF825C8D8726FCEAF8C2" author="Naderloo, R. &amp; Ebrahimnejad, S. &amp; Sari, A." box="[159,396,1842,1869]" pageId="5" pageNumber="435" pagination="397 - 412" refId="ref6775" refString="Naderloo, R., Ebrahimnejad, S. &amp; Sari, A. (2015) Annotated checklist of the decapod crustaceans of the Gulf of Oman, Northwestern Indian Ocean. Zootaxa, 4028 (3), 397 - 412. https: // doi. org / 10.11646 / zootaxa. 4028.3.5" type="journal article" year="2015">
Naderloo 
<emphasis id="B976EACAFFFFFF825D028726FC27F8C2" box="[272,321,1843,1869]" italics="true" pageId="5" pageNumber="435">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186553FFFFFF815CD58742FFF3FD0A" lastPageId="6" lastPageNumber="436" pageId="5" pageNumber="435" type="discussion">
<paragraph id="8BBD36D8FFFFFF825CD58742F952F852" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="435">
<emphasis id="B976EACAFFFFFF825CD58742FC5DF8FE" bold="true" box="[199,315,1879,1905]" pageId="5" pageNumber="435">Remarks.</emphasis>
The fresh specimens collected from the Persian Gulf agree with the 
<typeStatus id="54B9887AFFFFFF8258488742F9A7F8FE" box="[1114,1217,1879,1905]" pageId="5" pageNumber="435" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="4C024D5BFFFFFF8258F98742F81AF8FE" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="2023" box="[1259,1404,1879,1905]" class="Malacostraca" family="Leucosiidae" genus="Ebalia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="sagittifera" status="comb. nov.">
<emphasis id="B976EACAFFFFFF8258F98742F81AF8FE" box="[1259,1404,1879,1905]" italics="true" pageId="5" pageNumber="435">T. sagittifera</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFFFF8259948742FDB9F81A" bold="true" pageId="5" pageNumber="435">
<taxonomicNameLabel id="A24557B1FFFFFF8259948742FDB9F81A" pageId="5" pageNumber="435" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
The male specimen collected from the Persian Gulf (
<figureCitation id="13392A5DFFFFFF825F5D876EFECEF81A" box="[847,936,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="5" pageNumber="435">Fig. 1A</figureCitation>
) differs from the 
<typeStatus id="54B9887AFFFFFF825868876EF987F81A" box="[1146,1249,1915,1941]" pageId="5" pageNumber="435" type="lectotype">lectotype</typeStatus>
male by having smaller blunt teeth on the lateral sides of posterior margin as well as prominently swollen cardiac region. These differences can be attributed to differences in the body size of the male specimens.
</paragraph>
<paragraph id="8BBD36D8FFFCFF815CD58082FFF3FD0A" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="436">
<taxonomicName id="4C024D5BFFFCFF815CD58082FCCEFF3E" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[199,424,151,177]" class="Malacostraca" family="Leucosiidae" genus="Tirmilyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="sagittifera" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815CD58082FCCEFF3E" box="[199,424,151,177]" italics="true" pageId="6" pageNumber="436">Tirmilyra sagittifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557B1FFFCFF815DBF8082FF68FF3E" box="[429,526,151,177]" pageId="6" pageNumber="436" rank="species">
n. 
<emphasis id="B976EACAFFFCFF815DD48082FF68FF3E" bold="true" box="[454,526,151,177]" pageId="6" pageNumber="436">comb.</emphasis>
</taxonomicNameLabel>
closely resembles its congener 
<taxonomicName id="4C024D5BFFFCFF815F768082FE89FF3F" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[868,1007,151,176]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815F768082FE89FF3F" box="[868,1007,151,176]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557B1FFFCFF815FE18082F930FF3E" box="[1011,1110,151,177]" pageId="6" pageNumber="436" rank="species">
n. 
<emphasis id="B976EACAFFFCFF81581E8082F930FF3E" bold="true" box="[1036,1110,151,177]" pageId="6" pageNumber="436">comb.</emphasis>
</taxonomicNameLabel>
<emphasis id="B976EACAFFFCFF8158448082F93AFF3E" bold="true" box="[1110,1116,151,177]" pageId="6" pageNumber="436">,</emphasis>
but they can be differentiated on the basis of the following characteristics: dorsal surface of carapace without denticle on the metagastric region (
<figureCitation id="13392A5DFFFCFF815C8D80CAFC58FF76" box="[159,318,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A, B, D</figureCitation>
) (versus denticle present on the metagastric region of carapace in 
<taxonomicName id="4C024D5BFFFCFF81582180CAF9A4FF77" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[1075,1218,223,248]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF81582180CAF9A4FF77" box="[1075,1218,223,248]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF8158D880CAF85FFF76" bold="true" box="[1226,1337,223,249]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF8158D880CAF854FF76" box="[1226,1330,223,249]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
,
</emphasis>
<figureCitation id="13392A5DFFFCFF81595380CAF8F1FF76" box="[1345,1431,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1E</figureCitation>
; 
<bibRefCitation id="EF934B29FFFCFF815C858116FCF5FE92" author="Ghani, N. &amp; Tirmizi, N. M." box="[151,403,258,285]" pageId="6" pageNumber="436" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi 1995</bibRefCitation>
: 
<figureCitation id="13392A5DFFFCFF815DB68116FC9AFE92" box="[420,508,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A</figureCitation>
); posterior margin slightly convex (
<figureCitation id="13392A5DFFFCFF815F838116F94BFE92" box="[913,1069,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A, B, D</figureCitation>
) (versus straight in 
<taxonomicName id="4C024D5BFFFCFF81591D8116F8FBFE93" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[1295,1437,259,284]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF81591D8116F8FBFE93" box="[1295,1437,259,284]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815C858132FC66FECE" bold="true" box="[151,256,295,321]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815C858132FC66FECE" box="[151,256,295,321]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13392A5DFFFCFF815D038132FC0CFECE" box="[273,362,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1E</figureCitation>
; 
<bibRefCitation id="EF934B29FFFCFF815D6B8132FF1AFECE" author="Ghani, N. &amp; Tirmizi, N. M." box="[377,636,294,321]" pageId="6" pageNumber="436" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi 1995</bibRefCitation>
: 
<figureCitation id="13392A5DFFFCFF815E9D8132FF8CFECE" box="[655,746,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A</figureCitation>
); posterior margin bearing median tooth in males (
<figureCitation id="13392A5DFFFCFF81592E8132FDA9FEEA" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A, B, D</figureCitation>
) (versus median tooth absent in males in 
<taxonomicName id="4C024D5BFFFCFF815EBD815EFE58FEEB" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[687,830,331,356]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815EBD815EFE58FEEB" box="[687,830,331,356]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815F54815EFEC8FEEA" bold="true" box="[838,942,331,357]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815F54815EFEC8FEEA" box="[838,942,331,357]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13392A5DFFFCFF815FAC815EF973FEEA" box="[958,1045,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1E</figureCitation>
; 
<bibRefCitation id="EF934B29FFFCFF815831815EF844FEEA" author="Ghani, N. &amp; Tirmizi, N. M." box="[1059,1314,330,357]" pageId="6" pageNumber="436" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi 1995</bibRefCitation>
: 
<figureCitation id="13392A5DFFFCFF815926815EF8E8FEEA" box="[1332,1422,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A</figureCitation>
); blunt teeth present on the lateral sides of posterior margin (
<figureCitation id="13392A5DFFFCFF815F3D817AFEB2FE06" box="[815,980,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Figs. 1A, B, D</figureCitation>
) (versus pointed teeth on lateral sides of posterior margin in 
<taxonomicName id="4C024D5BFFFCFF815D608186FC99FE23" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[370,511,403,428]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815D608186FC99FE23" box="[370,511,403,428]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815E178186FF0CFE22" bold="true" box="[517,618,403,429]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815E178186FF0CFE22" box="[517,618,403,429]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13392A5DFFFCFF815E658186FFAAFE22" box="[631,716,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1E</figureCitation>
; 
<bibRefCitation id="EF934B29FFFCFF815EC58186FEA8FE22" author="Ghani, N. &amp; Tirmizi, N. M." box="[727,974,402,429]" pageId="6" pageNumber="436" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi 1995</bibRefCitation>
: 
<figureCitation id="13392A5DFFFCFF815FCC8186F953FE22" box="[990,1077,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1A</figureCitation>
); thoracic sternum tuberculated, prominent at margins (
<figureCitation id="13392A5DFFFCFF815D8F81A2FC92FE5E" box="[413,500,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1C</figureCitation>
) (versus thoracic sternum smooth, tuberculation only on margins in 
<taxonomicName id="4C024D5BFFFCFF81591F81A2F8FBFE5F" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[1293,1437,439,464]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF81591F81A2F8FBFE5F" box="[1293,1437,439,464]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815C8581CEFD99FE7A" bold="true" box="[151,255,475,501]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815C8581CEFD99FE7A" box="[151,255,475,501]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13392A5DFFFCFF815D1D81CEFC02FE7A" box="[271,356,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1F</figureCitation>
); male G1 apical process comparatively long (0.3× as long as whole length), slightly sinuous-shape, with tapering ending (
<figureCitation id="13392A5DFFFCFF815DC881EAFF32FD96" box="[474,596,511,537]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="6" pageNumber="436">Fig. 4A–D</figureCitation>
) (versus apical process comparatively short (0.2× as long as whole length), more sinuous-shape, with spatulate ending in 
<taxonomicName id="4C024D5BFFFCFF815EB28236FE49FDB3" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[672,815,547,572]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815EB28236FE49FDB3" box="[672,815,547,572]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815F258236FEF9FDB2" bold="true" box="[823,927,547,573]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815F258236FEF9FDB2" box="[823,927,547,573]" pageId="6" pageNumber="436" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13392A5DFFFCFF815FBC8236F943FDB2" box="[942,1061,547,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1668]" captionTargetBox="[159,1427,641,1616]" captionTargetId="figure-192@7.[151,1436,633,1621]" captionTargetPageId="7" captionText="FIGURE 4. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) G1 shaft; (B) G1 tip; Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (CL 4.2 mm, CW 3.9 mm) (ZMUC CRU885), Bushehr, Iran, Persian Gulf: (C) G1 shaft; (D) G1 tip; Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) G1 shaft; (F) G1 tip." figureDoi="http://doi.org/10.5281/zenodo.8254918" httpUri="https://zenodo.org/record/8254918/files/figure.png" pageId="6" pageNumber="436">Fig. 4E, F</figureCitation>
; 
<bibRefCitation id="EF934B29FFFCFF8158238236F849FDB2" author="Ghani, N. &amp; Tirmizi, N. M." box="[1073,1327,546,573]" pageId="6" pageNumber="436" pagination="143 - 147" refId="ref6537" refString="Ghani, N. &amp; Tirmizi, N. M. (1995) A new leucosid crab from Pakistan waters of the northern Arabian Sea. In: Thomson, M. F. &amp; Tirmizi, N. M. (Eds.), The Arabian Sea: Living Marine Resources and the Environment. Vanguard Books (Pvt) Ltd, Rotterdam, pp. 143 - 147." type="book chapter" year="1995">Ghani &amp; Tirmizi 1995</bibRefCitation>
: 
<figureCitation id="13392A5DFFFCFF8159538236FDCFFDEE" captionStart="FIGURE 1" captionStartId="2.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-17@2.[151,1436,181,1730]" captionTargetPageId="2" captionText="FIGURE 1. Tirmilyra sagittifera (Alcock, 1896) n. comb., lectotype male (CL 5.0 mm, CW 4.5 mm) (ZSI C910/10), Karachi, Pakistan: (A) dorsal view; (B) carapace dorsal view; (C) ventral view. Tirmilyra sagittifera (Alcock, 1896) n. comb., fresh specimen male (ZUMC CRU885) (CL 4.2 mm, CW 3.9 mm) Bushehr, Iran, Persian Gulf: (D) dorsal view. Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh specimen male (CL 8.4 mm, CW 7.6 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (E) dorsal view; (F) ventral view." figureDoi="http://doi.org/10.5281/zenodo.8254910" httpUri="https://zenodo.org/record/8254910/files/figure.png" pageId="6" pageNumber="436">Fig. 1E, F</figureCitation>
), endostome almost reaching inhalent channels (
<figureCitation id="13392A5DFFFCFF815EC58252FE57FDEE" box="[727,817,582,609]" captionStart="FIGURE 5" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[160,1422,822,1692]" captionTargetId="figure-273@8.[151,1436,813,1700]" captionTargetPageId="8" captionText="FIGURE 5. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) frontal view of cephalothorax; (B) female gonopore (right side); Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) frontal view of cephalothorax; (D) female gonopore (right side)." figureDoi="http://doi.org/10.5281/zenodo.8254922" httpUri="https://zenodo.org/record/8254922/files/figure.png" pageId="6" pageNumber="436">Fig. 5A</figureCitation>
) (versus endostome reaching slightly beyond inhalent channels in 
<taxonomicName id="4C024D5BFFFCFF815D0F827EFCCCFD0B" authorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" authorityYear="2023" baseAuthorityName="Patel &amp; Naderloo &amp; Trivedi &amp; Mitra" baseAuthorityYear="1995" box="[285,426,619,644]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="concinnus" status="comb. nov.">
<emphasis id="B976EACAFFFCFF815D0F827EFCCCFD0B" box="[285,426,619,644]" italics="true" pageId="6" pageNumber="436">T. concinnus</emphasis>
</taxonomicName>
<emphasis id="B976EACAFFFCFF815DA3827EFF57FD0A" bold="true" box="[433,561,619,645]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24557B1FFFCFF815DA3827EFF57FD0A" box="[433,561,619,645]" pageId="6" pageNumber="436" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="13392A5DFFFCFF815E2A827EFFEBFD0A" box="[568,653,618,645]" captionStart="FIGURE 5" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[160,1422,822,1692]" captionTargetId="figure-273@8.[151,1436,813,1700]" captionTargetPageId="8" captionText="FIGURE 5. Tirmilyra sagittifera (Alcock, 1896) n. comb., paralectotype female (CL 6.0 mm, CW 5.7 mm) (ZSI C910/10) Karachi, Pakistan: (A) frontal view of cephalothorax; (B) female gonopore (right side); Tirmilyra concinnus (Ghani &amp; Tirmizi, 1995) n. comb., fresh female (CL 9.4 mm, CW 8.4 mm) (ZUTC 6925) Hormoz Island, Persian Gulf: (C) frontal view of cephalothorax; (D) female gonopore (right side)." figureDoi="http://doi.org/10.5281/zenodo.8254922" httpUri="https://zenodo.org/record/8254922/files/figure.png" pageId="6" pageNumber="436">Fig. 5C</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>