plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/AB/87/03AB8793FF92AB67FF3BFED0FE88F8B0.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

603 lines
115 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="944AE291993131620A242BCD65A2B3C6" ID-CLB-Dataset="201894" ID-DOI="10.11646/zootaxa.5214.3.6" ID-GBIF-Dataset="1008c02f-024f-43b8-8f4f-8e057949d5d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389472" ID-ZooBank="3FE60779-E207-433F-B827-4C4D68C17432" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970704146" checkinUser="plazi" docAuthor="Osawa, Masayuki &amp; Fujita, Yoshihisa" docDate="2022" docId="03AB8793FF92AB67FF3BFED0FE88F8B0" docLanguage="en" docName="zootaxa.5214.3.6.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eutrichopagurus crassoculus Osawa &amp; Fujita 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="447" masterDocId="FF92FFEBFF93AB60FFACFFBFFFD8FFAC" masterDocTitle="A new species of the genus Eutrichopagurus Komai, 2015 (Crustacea: Decapoda Anomura: Paguridae) from a submarine cave in the Ryukyu Islands, southwestern Japan" masterLastPageNumber="450" masterPageNumber="440" pageNumber="441" updateTime="1699413976561" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7EA9F8AC556FDFEB8E4902A43EBD8EE1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4BB32E7305394E35310950705C97667A">
<mods:title id="E4288A63F303E0268F485C5E8097B9DB">A new species of the genus Eutrichopagurus Komai, 2015 (Crustacea: Decapoda Anomura: Paguridae) from a submarine cave in the Ryukyu Islands, southwestern Japan</mods:title>
</mods:titleInfo>
<mods:name id="C4B91AFA7DDD514987171D46E8B9E2BA" type="personal">
<mods:role id="D64EB492C4E4D68EE6984D78EFB4A695">
<mods:roleTerm id="27F2F56E76847A6F6478BFB32FFE2982">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="40EF113DB2C145713EB9F2427C2EC909">Osawa, Masayuki</mods:namePart>
<mods:nameIdentifier id="4FD8F6B9C83258280E99C2741B5BECCA" type="ORCID">0000-0001-8600-7550</mods:nameIdentifier>
<mods:affiliation id="91795B3C3370D7249AC36FF01C8009FF">Estuary Research Center, Shimane University, 1060 Nishikawatsu-cho, Matsue, Shimane, 690 - 8504 Japan osawam @ soc. shimane-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8600 - 7550</mods:affiliation>
<mods:nameIdentifier id="E8FF195EB10E6C75B32AD4974001EE1C" type="email">osawam@soc.shimane-u.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="C8D318FF62752DB26C31D2F9EADDE420" type="personal">
<mods:role id="39E2A84EFE28E8A160FE8289D31F6F5D">
<mods:roleTerm id="0565C6ACC44E33FDFF1D8E15999C25BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C35B6389788924F33FD68E395125AACE">Fujita, Yoshihisa</mods:namePart>
<mods:affiliation id="D63FFC8AF27D354C9DD5E6C40798EC3A">Okinawa Prefectual University of Arts, 1 - 4 Shuri-Tounokura, Naha, Okinawa 903 - 8602, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2ABD579BEE80B9628808D34DC191873B">text</mods:typeOfResource>
<mods:relatedItem id="E8634EC7A86040570CD7DC70FDA0AE51" type="host">
<mods:titleInfo id="1E92C4E6723B5C31FB10E46DC03CFD64">
<mods:title id="DF36E38DE9C69AC1403757235DE669DE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="26FCB3C4A699EB89DE06CEA0B529E395">
<mods:date id="6916FC6F3C464A056B5FAAC74E714A7D">2022</mods:date>
<mods:detail id="92BE1279334F46FEB6EEAFBBE3DE1478" type="pubDate">
<mods:number id="E10EF0232C61F6DCCC3BBC0C27965DCE">2022-12-02</mods:number>
</mods:detail>
<mods:detail id="F1E07A02B941DE114DF90CEC3B392611" type="volume">
<mods:number id="0EB9519C9765B016D2C553D12F732A77">5214</mods:number>
</mods:detail>
<mods:detail id="CF07F6ABE02B189DB3A416510ED425FD" type="issue">
<mods:number id="D538416A731251E085F66E296B6F3ECF">3</mods:number>
</mods:detail>
<mods:extent id="985B71845686FE3E4CB544A622E00FD3" unit="page">
<mods:start id="E26026790C059687471D91F510BD0C44">440</mods:start>
<mods:end id="E92897A3DCBBC1AB252BFE06B390F2C9">450</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AEFB4B1EF3E615AB54937E0D5CE4A473">journal article</mods:classification>
<mods:identifier id="A5105C42B48B15020B022EC360FEFB84" type="CLB-Dataset">201894</mods:identifier>
<mods:identifier id="61391B7B185AB6E5ACDFB902DF492CF7" type="DOI">10.11646/zootaxa.5214.3.6</mods:identifier>
<mods:identifier id="95EF72A40DCA6C0A31E4034E289BD71F" type="GBIF-Dataset">1008c02f-024f-43b8-8f4f-8e057949d5d8</mods:identifier>
<mods:identifier id="623D0234BBBF5CE33331D0707CC31EE4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7D585CDD4955916A60BE3035D5212464" type="Zenodo-Dep">7389472</mods:identifier>
<mods:identifier id="B7D1207B489DCDFA364B54E9B628A702" type="ZooBank">3FE60779-E207-433F-B827-4C4D68C17432</mods:identifier>
</mods:mods>
<treatment id="03AB8793FF92AB67FF3BFED0FE88F8B0" ID-DOI="http://doi.org/10.5281/zenodo.7400374" ID-GBIF-Taxon="205794363" ID-Zenodo-Dep="7400374" LSID="urn:lsid:plazi:treatment:03AB8793FF92AB67FF3BFED0FE88F8B0" httpUri="http://treatment.plazi.org/id/03AB8793FF92AB67FF3BFED0FE88F8B0" lastPageId="7" lastPageNumber="447" pageId="1" pageNumber="441">
<subSubSection id="C318650EFF92AB61FF3BFED0FDAFFE01" pageId="1" pageNumber="441" type="nomenclature">
<paragraph id="8BBD3685FF92AB61FF3BFED0FDE9FE25" blockId="1.[151,631,367,465]" box="[151,561,367,394]" pageId="1" pageNumber="441">
<heading id="D0F581E9FF92AB61FF3BFED0FDE9FE25" bold="true" box="[151,561,367,394]" fontSize="11" level="1" pageId="1" pageNumber="441" reason="1">
<emphasis id="B976EA97FF92AB61FF3BFED0FDE9FE25" bold="true" box="[151,561,367,394]" pageId="1" pageNumber="441">
<taxonomicName id="4C024D06FF92AB61FF3BFED0FE33FE26" ID-CoL="d01c7c8e-95f7-4afb-ae5f-9793fa6e6926" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[151,491,367,394]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="441" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF92AB61FF3BFED0FE33FE26" bold="true" box="[151,491,367,394]" italics="true" pageId="1" pageNumber="441">Eutrichopagurus crassoculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557ECFF92AB61FE5EFED0FDE9FE25" box="[498,561,367,393]" pageId="1" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD3685FF92AB61FF3BFE2BFDAFFE01" blockId="1.[151,631,367,465]" box="[151,631,403,430]" pageId="1" pageNumber="441">[New Japanese name: Kurayami-yadokari]</paragraph>
</subSubSection>
<subSubSection id="C318650EFF92AB61FF3BFE08FECAFE7D" box="[151,274,439,465]" pageId="1" pageNumber="441" type="description">
<paragraph id="8BBD3685FF92AB61FF3BFE08FECAFE7D" blockId="1.[151,631,367,465]" box="[151,274,439,465]" pageId="1" pageNumber="441">
(
<figureCitation id="13392A00FF92AB61FF33FE08FED2FE7D" box="[159,266,439,465]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1759,1783]" captionStartId-1="3.[151,250,1811,1835]" captionStartId-2="5.[151,250,1951,1975]" captionStartId-3="7.[151,250,1292,1316]" captionTargetBox-0="[173,1418,188,1730]" captionTargetBox-1="[163,1428,181,1783]" captionTargetBox-2="[157,1429,783,1924]" captionTargetBox-3="[155,1432,777,1269]" captionTargetId-0="figure-17@2.[164,1423,181,1736]" captionTargetId-1="figure-26@3.[151,1436,181,1787]" captionTargetId-2="figure-267@5.[151,1436,777,1928]" captionTargetId-3="figure-362@7.[155,1432,777,1269]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." captionText-1="FIGURE 2. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted); E, left cheliped, mesial view; F, same, lateral view; G, same, chela, dorsal view (setae omitted); H, same, carpus, dorsal view (setae omitted). Scale: 1.0 mm." captionText-2="FIGURE 3. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right pereopod 2, lateral view; B, same, dactylus, mesial view (setae drawn only on mesial surface); C, left pereopod 3, lateral view; H, same, dactylus, mesial view (setae drawn only on mesial surface). Scale: 1.0 mm." captionText-3="FIGURE 4. Eutrichopagurus crassoculus n. sp., paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.7389474" figureDoi-1="http://doi.org/10.5281/zenodo.7389478" figureDoi-2="http://doi.org/10.5281/zenodo.7389480" figureDoi-3="http://doi.org/10.5281/zenodo.7389486" httpUri-0="https://zenodo.org/record/7389474/files/figure.png" httpUri-1="https://zenodo.org/record/7389478/files/figure.png" httpUri-2="https://zenodo.org/record/7389480/files/figure.png" httpUri-3="https://zenodo.org/record/7389486/files/figure.png" pageId="1" pageNumber="441">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C318650EFF92AB61FF3BFE40FAE5FDCD" pageId="1" pageNumber="441" type="materials_examined">
<paragraph id="8BBD3685FF92AB61FF3BFE40FAE5FDCD" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
<emphasis id="B976EA97FF92AB61FF3BFE40FE92FDB5" bold="true" box="[151,330,511,537]" pageId="1" pageNumber="441">Type material.</emphasis>
<materialsCitation id="3B6A3CD8FF92AB61FEFAFE40FE8AFDCD" ID-GBIF-Occurrence="3986693302" collectingDate="2020-09-24" collectionCode="SCUBA, T" collectorName="Y. Fujita &amp; Fujii" county="Hedo Dome" elevation="4" location="Island" municipality="Kunigami" pageId="1" pageNumber="441" specimenCode="RUMF-ZC 6115" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B976EA97FF92AB61FEFAFE40FE65FDB5" box="[342,445,511,537]" italics="true" pageId="1" pageNumber="441">
<typeStatus id="54B98827FF92AB61FEFAFE40FE65FDB5" box="[342,445,511,537]" pageId="1" pageNumber="441" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DBA49EFEFF92AB61FE7DFE40FD48FDB6" box="[465,656,511,538]" collectionCode="RUMF-ZC" pageId="1" pageNumber="441">RUMF-ZC 6115</specimenCode>
(HD 2020-56), male (sl
<quantity id="4CFA9B60FF92AB61FC12FE40FBC5FDB5" box="[958,1053,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="441" unit="mm" value="1.2">1.2 mm</quantity>
),
<collectingCounty id="62DC4E09FF92AB61FB99FE40FB1AFDB6" box="[1077,1218,511,538]" pageId="1" pageNumber="441">Hedo Dome</collectingCounty>
(= Ginama) cave,
<collectingMunicipality id="6BD9ACFFFF92AB61FF3BFD9CFED2FD91" box="[151,266,547,573]" pageId="1" pageNumber="441">Kunigami</collectingMunicipality>
,
<collectingRegion id="49C6F867FF92AB61FEBBFD9CFEA4FD91" box="[279,380,547,573]" country="Japan" name="Okinawa" pageId="1" pageNumber="441">Okinawa</collectingRegion>
<location id="8EDD605EFF92AB61FE2AFD9CFE15FD91" LSID="urn:lsid:plazi:treatment:03AB8793FF92AB67FF3BFED0FE88F8B0:8EDD605EFF92AB61FE2AFD9CFE15FD91" box="[390,461,547,573]" county="Hedo Dome" municipality="Kunigami" name="Island" pageId="1" pageNumber="441" stateProvince="Okinawa">Island</location>
,
<location id="8EDD605EFF92AB61FE76FD9CFD55FD91" LSID="urn:lsid:plazi:treatment:03AB8793FF92AB67FF3BFED0FE88F8B0:8EDD605EFF92AB61FE76FD9CFD55FD91" box="[474,653,547,573]" county="Hedo Dome" municipality="Kunigami" name="Ryukyu Islands" pageId="1" pageNumber="441" stateProvince="Okinawa">Ryukyu Islands</location>
, zone 2,
<elevation id="002FD1B6FF92AB61FD5AFD9CFC51FD91" box="[758,905,547,574]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="441" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<quantity id="4CFA9B60FF92AB61FD5AFD9CFCE7FD92" box="[758,831,547,574]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="441" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</quantity>
depth
</elevation>
,
<date id="FFBC1045FF92AB61FC3BFD9CFBA0FD91" box="[919,1144,547,573]" pageId="1" pageNumber="441" value="2020-09-24">
<collectingDate id="EFF8E9ADFF92AB61FC3BFD9CFBA0FD91" box="[919,1144,547,573]" pageId="1" pageNumber="441" value="2020-09-24">24 September 2020</collectingDate>
</date>
,
<collectionCode id="ED13AE40FF92AB61FB29FD9CFB3DFD91" box="[1157,1253,547,573]" pageId="1" pageNumber="441">SCUBA</collectionCode>
, collected by
<collectorName id="26F75353FF92AB61FA29FD9BFF00FDCD" pageId="1" pageNumber="441">Y. Fujita</collectorName>
&amp;
<collectionCode id="ED13AE40FF92AB61FF50FDF7FED6FDCE" box="[252,270,584,610]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="441">T</collectionCode>
.
<collectorName id="26F75353FF92AB61FEB5FDF8FE8AFDCD" box="[281,338,583,609]" pageId="1" pageNumber="441">Fujii</collectorName>
</materialsCitation>
.
<materialsCitation id="3B6A3CD8FF92AB61FEF0FDF7FAE2FDCD" ID-GBIF-Occurrence="3986693301" box="[348,1338,583,610]" collectingDate="2020-09-24" collectionCode="RUMF-ZC" collectorName="Y. Fujita &amp; Fujii" county="Hedo Dome" elevation="4" location="Island" municipality="Kunigami" pageId="1" pageNumber="441" specimenCode="RUMF-ZC 6116" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="B976EA97FF92AB61FEF0FDF7FE1BFDCD" box="[348,451,584,609]" italics="true" pageId="1" pageNumber="441">
<typeStatus id="54B98827FF92AB61FEF0FDF7FE1BFDCD" box="[348,451,584,609]" pageId="1" pageNumber="441" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCode id="DBA49EFEFF92AB61FE7DFDF8FD54FDCD" box="[465,652,583,609]" collectionCode="RUMF-ZC" pageId="1" pageNumber="441">RUMF-ZC 6116</specimenCode>
(HD 2020-54),
<specimenCount id="9D04FD0CFF92AB61FCECFDF8FC53FDCD" box="[832,907,583,609]" pageId="1" pageNumber="441" type="male">1 male</specimenCount>
(sl
<quantity id="4CFA9B60FF92AB61FC19FDF8FBD7FDCD" box="[949,1039,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="441" unit="mm" value="1.2">1.2 mm</quantity>
), same data as in
<typeStatus id="54B98827FF92AB61FB78FDF8FAE2FDCD" box="[1236,1338,583,609]" pageId="1" pageNumber="441" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C318650EFF92AB66FF6BFDD4FF6DFEB1" lastPageId="6" lastPageNumber="446" pageId="1" pageNumber="441" type="description">
<paragraph id="8BBD3685FF92AB61FF6BFDD4FCFDFD29" blockId="1.[151,1437,511,1906]" box="[199,805,619,645]" pageId="1" pageNumber="441">
<emphasis id="B976EA97FF92AB61FF6BFDD4FE81FD29" bold="true" box="[199,345,619,645]" pageId="1" pageNumber="441">Description.</emphasis>
Eleven pairs of deeply quadriserial gills.
</paragraph>
<paragraph id="8BBD3685FF92AB61FF6BFD30FE28FC09" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
Shield (
<figureCitation id="13392A00FF92AB61FE8CFD30FE44FD05" box="[288,412,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1A, B</figureCitation>
) subtriangular in general outline, 1.1–1.2 times as long as wide; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin roundly truncate, slightly concave on median part; dorsal surface slightly convex transversely, with 1 median, 1 submedian, and 1 lateral pairs of few short setae on anterior half. Rostrum triangular, terminating in acute tip, distinctly overreaching lateral projections. Lateral projections obtusely triangular, each with short marginal spine. Posterior carapace (
<figureCitation id="13392A00FF92AB61FAC3FCA0FF61FCF1" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1A</figureCitation>
) with lateral lobes very narrow. Cardiac sulci not discernible. Sulci cardiobranchiales extending to posterior carapace margin, converging posteriorly. Posteromedian and posterolateral plates poorly calcified; branchiostegites membranous, almost glabrous.
</paragraph>
<paragraph id="8BBD3685FF92AB61FF6BFC10FA5DFB0E" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
Ocular peduncle (including cornea) (
<figureCitation id="13392A00FF92AB61FDF5FC10FD13FC65" box="[601,715,943,969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1B, C</figureCitation>
) stout, about half-length of shield, widest at about midlength, with sparse tufts of setae distally on each dorsomesial surface; cornea greatly reduced, rounded on distal margin, much narrower than peduncle; lateral surface strongly convex on proximal half and concave and somewhat decalcified on distal part including cornea (in preserved condition, showing strongly concave appearance, but in fresh condition, only weakly concave;
<figureCitation id="13392A00FF92AB61FE34FB80FE2BFBF5" box="[408,499,1087,1113]" captionStart="FIGURE 4" captionStartId="7.[151,250,1292,1316]" captionTargetBox="[155,1432,777,1269]" captionTargetId="figure-362@7.[155,1432,777,1269]" captionTargetPageId="7" captionText="FIGURE 4. Eutrichopagurus crassoculus n. sp., paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7389486" httpUri="https://zenodo.org/record/7389486/files/figure.png" pageId="1" pageNumber="441">Fig. 4A</figureCitation>
); mesial surface flattish, with bluntly delimited dorsomesial margin. Ocular acicle ovate, with small submarginal spine reaching proximal 0.2 of ocular peduncle; dorsal surface flattish. Interocular lobe separated from ocular acicles, overhung by rostrum, visible in dorsal view; anterior surface slightly convex.
</paragraph>
<paragraph id="8BBD3685FF92AB61FF6BFB14FA45FA31" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
Antennular peduncle (
<figureCitation id="13392A00FF92AB61FE6CFB14FDE0FB69" box="[448,568,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1B, D</figureCitation>
) long, when fully extended, overreaching distal corneal margin by full length of article 3 and distal one-third of article 2. Article 3 3.2 times as long as high, approximately 2.1 times as long as article 2, somewhat widened distally, with prominent tuft of long setae (length subequal to that of dorsal flagellum) on dorsodistal margin. Article 1 with distolateral margin not markedly produced; statocyst lobe slightly inflated, with small spine on lateral face. Dorsal flagellum 1.4 times longer than peduncular article 3, consisting of short aesthetasc-bearing proximal portion (about 10 articles) and long distal portion (about 10 articles, about 2.8 length of aesthetasc-bearing portion); ventral flagellum about half length of dorsal flagellum, consisting of about 10 articles.
</paragraph>
<paragraph id="8BBD3685FF92AB61FF6BFA18FAC8F94D" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
Antennal peduncle (
<figureCitation id="13392A00FF92AB61FE02FA18FDFFFA6D" box="[430,551,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1B, E</figureCitation>
) overreaching distal corneal margin by nearly full length of article 5. Article 5 slightly flattened, with few setae marginally. Article 4 with few short setae ventromesially. Article 3 unarmed but with oblong capsule-like structure and few short setae on ventromesial distal margin. Article 2 dorsolateral distal angle produced, barely reaching midlength of article 4, terminating in simple spine, bearing few distal setae; dorsomesial distal angle with small spine; ventromesial margin with few short setae. Article 1 with minute distal spine on lateral surface; ventromesial distal margin somewhat produced anteriorly, but without armature. Antennal acicle overreaching distal corneal margin by distal one third length and reaching midlength of peduncular article 5, terminating in small spine; dorsomesial margin with row of sparse setae. Antennal flagellum (
<figureCitation id="13392A00FF92AB61FB07F91CFB2EF911" box="[1195,1270,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig.1F</figureCitation>
) 3.1–3.3 times shield length, overreaching extended right cheliped; each article with 0–3 short setae on distal margin.
</paragraph>
<paragraph id="8BBD3685FF92AB61FF6BF954FCC1F8DE" blockId="1.[151,1437,511,1906]" pageId="1" pageNumber="441">
Mouthparts not dissected. Maxilliped 3 (
<figureCitation id="13392A00FF92AB61FD30F954FCC3F8A9" box="[668,795,1771,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="1" pageNumber="441">Fig. 1G, H</figureCitation>
) moderately stout; carpus and merus unarmed; ischium with crista dentata composed of about 10 small closely set corneous teeth and 1 accessory tooth, dorsal margin with minute distal spine; basis-ischium fusion incomplete; basis with few minute denticles on ventromesial margin; exopod reaching slightly short of distal margin of carpus.
</paragraph>
<caption id="DF7D660DFF91AB62FF3BF960FA98F85F" ID-DOI="http://doi.org/10.5281/zenodo.7389474" ID-Zenodo-Dep="7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="2" pageNumber="442" startId="2.[151,250,1759,1783]" targetBox="[173,1418,188,1730]" targetPageId="2">
<paragraph id="8BBD3685FF91AB62FF3BF960FA98F85F" blockId="2.[151,1436,1759,2036]" pageId="2" pageNumber="442">
<emphasis id="B976EA97FF91AB62FF3BF960FECBF954" bold="true" box="[151,275,1759,1784]" pageId="2" pageNumber="442">FIGURE 1.</emphasis>
<taxonomicName id="4C024D06FF91AB62FEB5F95FFD9BF954" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[281,579,1760,1784]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="442" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF91AB62FEB5F95FFD9BF954" box="[281,579,1760,1784]" italics="true" pageId="2" pageNumber="442">Eutrichopagurus crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF91AB62FDE5F95FFD5AF954" bold="true" box="[585,642,1760,1784]" pageId="2" pageNumber="442">
<taxonomicNameLabel id="A24557ECFF91AB62FDE5F95FFD5AF954" box="[585,642,1760,1784]" pageId="2" pageNumber="442" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm.
</paragraph>
</caption>
<caption id="DF7D660DFF90AB63FF3BF8ACFEE8F83A" ID-DOI="http://doi.org/10.5281/zenodo.7389478" ID-Zenodo-Dep="7389478" httpUri="https://zenodo.org/record/7389478/files/figure.png" pageId="3" pageNumber="443" startId="3.[151,250,1811,1835]" targetBox="[163,1428,181,1783]" targetPageId="3">
<paragraph id="8BBD3685FF90AB63FF3BF8ACFEE8F83A" blockId="3.[151,1437,1811,1943]" pageId="3" pageNumber="443">
<emphasis id="B976EA97FF90AB63FF3BF8ACFECBF887" bold="true" box="[151,275,1811,1835]" pageId="3" pageNumber="443">FIGURE 2.</emphasis>
<taxonomicName id="4C024D06FF90AB63FEB6F8ACFD9CF887" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[282,580,1811,1835]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF90AB63FEB6F8ACFD9CF887" box="[282,580,1811,1835]" italics="true" pageId="3" pageNumber="443">Eutrichopagurus crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF90AB63FDE6F8ACFD5BF887" bold="true" box="[586,643,1811,1835]" pageId="3" pageNumber="443">
<taxonomicNameLabel id="A24557ECFF90AB63FDE6F8ACFD5BF887" box="[586,643,1811,1835]" pageId="3" pageNumber="443" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted); E, left cheliped, mesial view; F, same, lateral view; G, same, chela, dorsal view (setae omitted); H, same, carpus, dorsal view (setae omitted). Scale: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BBD3685FF97AB64FF6BFF28FE73FBBD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="444">
Chelipeds (
<figureCitation id="13392A00FF97AB64FEE9FF28FE2EFF1D" box="[325,502,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1811,1835]" captionTargetBox="[163,1428,181,1783]" captionTargetId="figure-26@3.[151,1436,181,1787]" captionTargetPageId="3" captionText="FIGURE 2. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted); E, left cheliped, mesial view; F, same, lateral view; G, same, chela, dorsal view (setae omitted); H, same, carpus, dorsal view (setae omitted). Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389478" httpUri="https://zenodo.org/record/7389478/files/figure.png" pageId="4" pageNumber="444">Fig. 2A, B, E, F</figureCitation>
) subequal in length, dissimilar with right much more robust than left. Right cheliped (
<figureCitation id="13392A00FF97AB64FF33FF04FEC4FF79" box="[159,284,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1811,1835]" captionTargetBox="[163,1428,181,1783]" captionTargetId="figure-26@3.[151,1436,181,1787]" captionTargetPageId="3" captionText="FIGURE 2. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted); E, left cheliped, mesial view; F, same, lateral view; G, same, chela, dorsal view (setae omitted); H, same, carpus, dorsal view (setae omitted). Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389478" httpUri="https://zenodo.org/record/7389478/files/figure.png" pageId="4" pageNumber="444">Fig. 2A–D</figureCitation>
) moderately stout, moderately large compared to body. Chela elongate subovate in dorsal view, 1.8 times as long as wide (greatest width at midlength of palm), 1.7 times longer than carpus; dorsomesial margin somewhat narrowing distally; dorsolateral margin gently arched. Dactylus 0.7 length of palm, weakly curved in dorsal and mesial views, with sparse tufts of setae on surfaces, terminating in small corneous claw; dorsal and mesial surfaces proximally with sparse granules, dorsomesial margin not delimited; occlusal margin with 3 subacute calcareous teeth on submedian part and row of small corneous teeth on distal concaved part. Palm with sparse short to moderately long setae and covering of coarse granules on surfaces, granules on fixed finger sparse; dorsomesial margin delimited by row of short granular ridges; dorsolateral margin weakly delimited; dorsal and ventral surfaces gently convex. Fixed finger terminating in small corneous claw; occlusal margin with low rounded calcareous tooth proximally and prominent roundly triangular calcareous tooth subdistally, concaved distal part with row of small corneous teeth. Carpus slightly widened distally in dorsal view, 1.4–1.5 times longer than distal width, 0.9 length of merus, bearing sparse bristle-like setae on dorsomesial and dorsomesial margins and short transverse ridges and granules dorsally on mesial and lateral surfaces; few short setae on ventral surface; all surfaces with coarse granules, fewer on dorsal surface; dorsomesial margin with 3 or 4 distal spines followed by some short granular ridges; dorsolateral margin with blunt distal projection followed by some short granular ridges; mesial surface dorsally with 2 distal spines and few short granular ridges, distomesial margin granular; lateral surface with few short granular ridges dorsally and distal small spine ventrally, distolateral margin with row of coarse granules; ventral surface convex. Merus with very low, short transverse ridges and some bristle-like and simple setae on dorsal surface, dorsodistal margin unarmed; mesial surface with coarse granules and few distal bristle-like setae ventrally, ventromesial margin with 2–4 distinct distal spines followed by some short ridges or minute spines; lateral surface with few short setae and short granular ridges and single granules ventrally; ventrolateral margin 3 spines on distal half followed by some short ridges; ventral surface slightly convex on proximal half, with coarse granules. Ischium with sparse setae on dorsodistal margin and ventral surface; ventromesial margin unarmed. Coxa laterally with minute distal spine.
</paragraph>
<paragraph id="8BBD3685FF97AB64FF6BFBA4FCA8F885" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="444">
Left cheliped (
<figureCitation id="13392A00FF97AB64FEC6FBA4FE3AFB99" box="[362,482,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1811,1835]" captionTargetBox="[163,1428,181,1783]" captionTargetId="figure-26@3.[151,1436,181,1787]" captionTargetPageId="3" captionText="FIGURE 2. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted); E, left cheliped, mesial view; F, same, lateral view; G, same, chela, dorsal view (setae omitted); H, same, carpus, dorsal view (setae omitted). Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389478" httpUri="https://zenodo.org/record/7389478/files/figure.png" pageId="4" pageNumber="444">Fig. 2E–H</figureCitation>
) slender. Chela 2.7–2.9 times longer than wide (greatest width at midlength of palm), 1.3–1.5 times longer than carpus; dorsolateral margin weakly undulate on distal two-thirds Dactylus 1.5 times longer than palm, gently curved distally in dorsal view, terminating in small corneous claw; surfaces with sparse setae, dorsal surface proximally with coarse granules, otherwise nearly smooth; mesial surface proximally with arc of granules, dorsomesial margin not delimited; occlusal margin with row of minute corneous teeth (teeth on proximal half widely to narrowly spaced, those on distal half contiguous). Palm with sparse setae on dorsal and ventral surfaces; dorsal surface convex, elevated with some proximal subacute or spinulose tubercles on midline, covered with minute granules, dorsomesial and dorsolateral margins each delimited by row of small tubercles; lateral surface covered with granules; ventral surface gently convex, nearly smooth. Fixed finger distinctly curved distally in dorsal view, with scattered setae on dorsal and ventral surfaces, terminating in small corneous claw; occlusal margin bordered with thin corneous plate. Carpus slightly widened distally in dorsal view, 2.8–3.2 times longer than distal width, 0.9 length of merus; dorsolateral and dorsomesial margins with 4 (1 distal and 3 median) and 2 distal spines followed by few short ridges, respectively, spines and short ridges bearing bristle-like and simple setae; mesial surface smooth, with few bristle-like setae dorsally and distally; lateral surface with some scaley ridges medially, distolateral margin with row of small tubercles and few short bristle-like setae; ventrodistal margin with 2 small distal spines; ventral surface convex, with few very short setae. Merus with row of short ridges bearing few bristle-like and simple setae on distal half of dorsal surface, dorsodistal margin unarmed; mesial surface smooth, with few very short setae ventrally, ventromesial margin with row of 4 or 5 small spines and terminal bristle-like setae on distal half and with some short somewhat elevated ridges and bristle-like setae on proximal half; lateral surface with sparse short ridges bearing short setae, ventrolateral margin with 2 or 3 small spines on distal one third followed by some short somewhat elevated ridges; ventral surface flattish, bearing few short setae. Ischium unarmed, with few setae. Coxa laterally with minute distal spine.
</paragraph>
<paragraph id="8BBD3685FF97AB65FF6BF88CFE33FEC9" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,753]" lastPageId="5" lastPageNumber="445" pageId="4" pageNumber="444">
Ambulatory legs (
<figureCitation id="13392A00FF97AB64FE38F88CFDD7F8E1" box="[404,527,1843,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1951,1975]" captionTargetBox="[157,1429,783,1924]" captionTargetId="figure-267@5.[151,1436,777,1928]" captionTargetPageId="5" captionText="FIGURE 3. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right pereopod 2, lateral view; B, same, dactylus, mesial view (setae drawn only on mesial surface); C, left pereopod 3, lateral view; H, same, dactylus, mesial view (setae drawn only on mesial surface). Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389480" httpUri="https://zenodo.org/record/7389480/files/figure.png" pageId="4" pageNumber="444">Fig. 3A–D</figureCitation>
) moderately long and slender; surfaces mostly smooth. Dactyli 1.0–1.1 times as long as propodi, in dorsal view slightly curved mesially, in lateral view slightly curved ventrally; dorsal margins each with irregularly arranged row of setae, setae on distal one-third more closely set than on those on remaining part; lateral and mesial faces almost glabrous; ventral margins each with row of 3 or 4 (second) and 5 or 6 (third) slender corneous spines. Propodi distinctly longer than carpi, slightly narrowing distally, 5.0–5.1 times as long as maximum high; dorsal surfaces each with sparse setae; lateral and mesial faces glabrous; ventral margins each with 1 or 2 slender corneous spines at distal end and 1 or 2 corneous spines on median one third. Carpi unarmed, with sparse setae on dorsal surfaces; ventrodistal margin unarmed. Meri 3.6–3.8 (second) and 3.0–3.9 (third) times as long as maximum high, with sparse setae on dorsal surfaces; lateral surfaces almost glabrous; ventral margins with 1–3 small spines on posterior half (second) and unarmed (third). Ischia unarmed, with sparse tufts of setae distally on dorsal and ventral margins.
</paragraph>
<paragraph id="8BBD3685FF96AB65FF6BFED0FD56FE7D" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="445">
Pereopod 4 (
<figureCitation id="13392A00FF96AB65FEF0FED0FE72FE25" box="[348,426,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="5" pageNumber="445">Fig. 1I</figureCitation>
) semichelate. Dactylus slightly curved. Propodus with rasp consisting of single row of corneous scales. Carpus unarmed on dosrsal surface. Dactylus to merus with sparse tufts of setae on dorsal surfaces, dorsodistal tuft of setae on merus prominent.
</paragraph>
<paragraph id="8BBD3685FF96AB65FF6BFE64FC5EFDCD" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="445">
Pereopods 5 chelate. Coxae (
<figureCitation id="13392A00FF96AB65FDA7FE64FDAFFE59" box="[523,631,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="5" pageNumber="445">Fig. 1J–L</figureCitation>
) asymmetrical; right coxa with long, filamentous sexual tube (about 4.7 times of coxal length) directed posterolaterally and recurved toward anterior between dorsal surface of ischium and base of pleon, with hooked part on about distal one third, prominent tuft of setae at base of sexual tube; left coxa with very short sexual tube, directed mesially and tapering distally.
</paragraph>
<paragraph id="8BBD3685FF96AB65FF6BFDD4FD04FD05" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="445">
Thoracic sternite 6 anterior lobe (
<figureCitation id="13392A00FF96AB65FD92FDD4FD42FD29" box="[574,666,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="5" pageNumber="445">Fig. 1M</figureCitation>
) transversely ovate, with pair of short setae anterolaterally. Thoracic sternite 8 (
<figureCitation id="13392A00FF96AB65FEA2FD30FE83FD05" box="[270,347,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="5" pageNumber="445">Fig. 1J</figureCitation>
) consisting of 2 contiguous lobes.
</paragraph>
<paragraph id="8BBD3685FF96AB65FF6BFD0CFBB7FD61" blockId="5.[151,1437,151,753]" box="[199,1135,690,717]" pageId="5" pageNumber="445">Pleon dextrally twisted. Male with unpaired, unequally biramous left pleopods 3–5.</paragraph>
<paragraph id="8BBD3685FF96AB65FF6BFD68FCFDFD5D" blockId="5.[151,1437,151,753]" box="[199,805,727,753]" pageId="5" pageNumber="445">Uropods markedly asymmetrical, protopods unarmed.</paragraph>
<caption id="DF7D660DFF96AB65FF3BF820FCE9F852" ID-DOI="http://doi.org/10.5281/zenodo.7389480" ID-Zenodo-Dep="7389480" httpUri="https://zenodo.org/record/7389480/files/figure.png" pageId="5" pageNumber="445" startId="5.[151,250,1951,1975]" targetBox="[157,1429,783,1924]" targetPageId="5">
<paragraph id="8BBD3685FF96AB65FF3BF820FCE9F852" blockId="5.[151,1436,1951,2047]" pageId="5" pageNumber="445">
<emphasis id="B976EA97FF96AB65FF3BF820FECDF81B" bold="true" box="[151,277,1951,1975]" pageId="5" pageNumber="445">FIGURE 3.</emphasis>
<taxonomicName id="4C024D06FF96AB65FEB2F820FD92F81B" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[286,586,1951,1975]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="445" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF96AB65FEB2F820FD92F81B" box="[286,586,1951,1975]" italics="true" pageId="5" pageNumber="445">Eutrichopagurus crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF96AB65FDFFF820FD56F81B" bold="true" box="[595,654,1951,1975]" pageId="5" pageNumber="445">
<taxonomicNameLabel id="A24557ECFF96AB65FDFFF820FD56F81B" box="[595,654,1951,1975]" pageId="5" pageNumber="445" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, right pereopod 2, lateral view; B, same, dactylus, mesial view (setae drawn only on mesial surface); C, left pereopod 3, lateral view; H, same, dactylus, mesial view (setae drawn only on mesial surface). Scale: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BBD3685FF95AB66FF6BFF28FAE3FF79" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
Telson (
<figureCitation id="13392A00FF95AB66FE8CFF28FEAFFF1D" box="[288,375,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="6" pageNumber="446">Fig. 1N</figureCitation>
) with lateral indentations at posterior one-third; posterior lobes subtriangular, terminal margins oblique, each with 3–5 minute acute or blunt spines, posterolateral margins each with fringe of long setae.
</paragraph>
<paragraph id="8BBD3685FF95AB66FF6BFF61FF6DFEB1" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<emphasis id="B976EA97FF95AB66FF6BFF61FE97FF54" bold="true" box="[199,335,222,248]" pageId="6" pageNumber="446">Coloration.</emphasis>
In fresh condition, body and appendage are entirely translucent white with pale brown tint (
<figureCitation id="13392A00FF95AB66FAC3FF60FF70FEB1" captionStart="FIGURE 4" captionStartId="7.[151,250,1292,1316]" captionTargetBox="[155,1432,777,1269]" captionTargetId="figure-362@7.[155,1432,777,1269]" captionTargetPageId="7" captionText="FIGURE 4. Eutrichopagurus crassoculus n. sp., paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7389486" httpUri="https://zenodo.org/record/7389486/files/figure.png" pageId="6" pageNumber="446">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C318650EFF95AB66FF6BFE99FE4EFC2D" pageId="6" pageNumber="446" type="distribution">
<paragraph id="8BBD3685FF95AB66FF6BFE99FF30FEC9" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<emphasis id="B976EA97FF95AB66FF6BFE99FEB9FEEC" bold="true" box="[199,353,294,320]" pageId="6" pageNumber="446">Distribution.</emphasis>
So far only known from the
<typeStatus id="54B98827FF95AB66FD7CFE98FCD8FEED" box="[720,768,295,321]" pageId="6" pageNumber="446">type</typeStatus>
locality,
<collectingRegion id="49C6F867FF95AB66FCDFFE98FC00FEED" box="[883,984,295,321]" country="Japan" name="Okinawa" pageId="6" pageNumber="446">Okinawa</collectingRegion>
Island (Ryukyu Islands, southwestern
<collectingCountry id="F3157615FF95AB66FF3BFEF4FF04FEC9" box="[151,220,331,357]" name="Japan" pageId="6" pageNumber="446">Japan</collectingCountry>
).
</paragraph>
<paragraph id="8BBD3685FF95AB66FF6BFED1FE76FDB5" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<emphasis id="B976EA97FF95AB66FF6BFED1FEF2FE24" bold="true" box="[199,298,366,392]" pageId="6" pageNumber="446">Habitat.</emphasis>
The
<specimenCount id="9D04FD0CFF95AB66FECEFED0FE1BFE25" box="[354,451,367,393]" pageId="6" pageNumber="446" type="generic" typeStatus="type">two type</specimenCount>
specimens were obtained from the area between “main tube” zone and “innermost part” zones in Hedo Dome cave (cf.
<bibRefCitation id="EF934B74FF95AB66FE5BFE2CFD36FE01" author="Osawa, M. &amp; Fujita, Y." box="[503,750,403,429]" pageId="6" pageNumber="446" pagination="463 - 482" refId="ref6924" refString="Osawa, M. &amp; Fujita, Y. (2019) Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan. Zootaxa, 4560 (3), 463 - 482. https: // doi. org / 10.11646 / zootaxa. 4560.3.3" type="journal article" year="2019">Osawa &amp; Fujita 2019</bibRefCitation>
), having an anchialine environment with salinity cline. They each carried a shell of gastropod referred to an unidentified species of the genus
<taxonomicName id="4C024D06FF95AB66FBEBFE08FA85FE7D" authority="Iredale, 1918" authorityName="Iredale" authorityYear="1918" box="[1095,1373,439,465]" class="Gastropoda" family="Colloniidae" genus="Collonista" kingdom="Animalia" order="Trochida" pageId="6" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="B976EA97FF95AB66FBEBFE08FB64FE7D" box="[1095,1212,439,465]" italics="true" pageId="6" pageNumber="446">Collonista</emphasis>
<bibRefCitation id="EF934B74FF95AB66FB6AFE08FA85FE7D" author="Iredale, T." box="[1222,1373,439,465]" pageId="6" pageNumber="446" pagination="28 - 40" refId="ref6302" refString="Iredale, T. (1918) Molluscan nomenclatural problems and solutions. No. 1. Proceedings of the Malacological Society of London, 13 (1 - 2), 28 - 40. https: // doi. org / 10.1093 / oxfordjournals. mollus. a 063674" type="journal article" year="1918">Iredale, 1918</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13392A00FF95AB66FAC3FE08FF06FE59" captionStart="FIGURE 4" captionStartId="7.[151,250,1292,1316]" captionTargetBox="[155,1432,777,1269]" captionTargetId="figure-362@7.[155,1432,777,1269]" captionTargetPageId="7" captionText="FIGURE 4. Eutrichopagurus crassoculus n. sp., paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7389486" httpUri="https://zenodo.org/record/7389486/files/figure.png" pageId="6" pageNumber="446">Fig. 4B; H</figureCitation>
. Kubo, personal communication). Numerous empty and living shells of that species were found in the same environment of the cave.
</paragraph>
<paragraph id="8BBD3685FF95AB66FF6BFD9CFE4EFC2D" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
Prior to this study, hermit crabs found in Hedo Dome cave included 12 species (
<bibRefCitation id="EF934B74FF95AB66FB99FD9CFAF0FD91" author="Osawa, M. &amp; Fujita, Y." box="[1077,1320,547,573]" pageId="6" pageNumber="446" pagination="463 - 482" refId="ref6924" refString="Osawa, M. &amp; Fujita, Y. (2019) Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan. Zootaxa, 4560 (3), 463 - 482. https: // doi. org / 10.11646 / zootaxa. 4560.3.3" type="journal article" year="2019">Osawa &amp; Fujita 2019</bibRefCitation>
):
<taxonomicName id="4C024D06FF95AB66FA96FD9CFE69FDCD" authority="Lewinsohn, 1982" authorityName="Lewinsohn" authorityYear="1982" class="Malacostraca" family="Diogenidae" genus="Aniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="retipes">
<emphasis id="B976EA97FF95AB66FA96FD9CFF3AFDCC" italics="true" pageId="6" pageNumber="446">Aniculus retipes</emphasis>
<bibRefCitation id="EF934B74FF95AB66FF46FDF8FE69FDCD" author="Lewinsohn, C." box="[234,433,583,609]" pageId="6" pageNumber="446" pagination="76 - 82" refId="ref6387" refString="Lewinsohn, C. (1982) Aniculus retipes n. sp. ein neuer Einsiedlerkrebs aus dem Roten Meer (Decapoda, Anomura). Crustaceana, 42 (1 - 2), 76 - 82. https: // doi. org / 10.1163 / 156854082 X 00713" type="journal article" year="1982">Lewinsohn, 1982</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FE12FDF8FCC6FDCD" authority="Morgan, 1992" authorityName="Morgan" authorityYear="1992" box="[446,798,583,609]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="jalur">
<emphasis id="B976EA97FF95AB66FE12FDF8FDACFDCD" box="[446,628,583,609]" italics="true" pageId="6" pageNumber="446">Paguristes jalur</emphasis>
<bibRefCitation id="EF934B74FF95AB66FDD0FDF8FCC6FDCD" author="Morgan, G. J." box="[636,798,583,609]" pageId="6" pageNumber="446" pagination="163 - 174" refId="ref6666" refString="Morgan, G. J. (1992) The hermit crabs (Crustacea: Decapoda: Coenobitidae, Diogenidae, Paguridae) of Christmas and Cocos (Keeling) Islands, Indian Ocean, with description of a new species of Paguristes. Raffles Bulletin of Zoology, 49 (2), 163 - 174." type="journal article" year="1992">Morgan, 1992</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FC87FDF8FADBFDCD" authority="Asakura, 2004" authorityName="Asakura" authorityYear="2004" box="[811,1283,583,609]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="B976EA97FF95AB66FC87FDF8FB8FFDCD" box="[811,1111,583,609]" italics="true" pageId="6" pageNumber="446">Pseudopaguristes shidarai</emphasis>
<bibRefCitation id="EF934B74FF95AB66FBF2FDF8FADBFDCD" author="Asakura, A." box="[1118,1283,583,609]" pageId="6" pageNumber="446" pagination="153 - 168" refId="ref5169" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura, 2004</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C024D06FF95AB66FABEFDF8FA40FDCD" authorityName="Ortmann" authorityYear="1892" box="[1298,1432,583,609]" class="Malacostraca" family="Diogenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
, three species);
<taxonomicName id="4C024D06FF95AB66FE97FDD4FCCAFD29" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[315,786,619,645]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B976EA97FF95AB66FE97FDD4FDE3FD29" box="[315,571,619,645]" italics="true" pageId="6" pageNumber="446">Catapaguroides foresti</emphasis>
<bibRefCitation id="EF934B74FF95AB66FD93FDD4FCCAFD29" author="McLaughlin, P. A." box="[575,786,619,645]" pageId="6" pageNumber="446" pagination="495 - 504" refId="ref6438" refString="McLaughlin, P. A. (2002) A new species of the genus Catapaguroides (Decapoda: Anomura: Paguroidea: Paguridae) from Guam, Micronesia. Crustaceana, 75 (3 - 4), 495 - 504. https: // doi. org / 10.1163 / 156854002760095543" type="journal article" year="2002">McLaughlin, 2002</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FCB6FDD4FAF8FD29" authority="Komai, Yamada &amp; Shirakawa, 2010" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[794,1312,619,645]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="B976EA97FF95AB66FCB6FDD4FC53FD29" box="[794,907,619,645]" italics="true" pageId="6" pageNumber="446">C. longior</emphasis>
<bibRefCitation id="EF934B74FF95AB66FC3DFDD4FAF8FD29" author="Komai, T. &amp; Yamada, Y. &amp; Shirakawa, N." box="[913,1312,619,645]" pageId="6" pageNumber="446" pagination="32 - 42" refId="ref6223" refString="Komai, T., Yamada, Y. &amp; Shirakawa, N. (2010) A new hermit crab species of the genus Catapaguroides A. Milne-Edwards &amp; Bouvier (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, Japan, and additional record of C. foresti McLaughlin. Zootaxa, 2690 (1), 32 - 42. https: // doi. org / 10.11646 / zootaxa. 2690.1.3" type="journal article" year="2010">Komai, Yamada &amp; Shirakawa, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FA85FDD4FDB8FD05" authority="Osawa, Fujita &amp; Okuno, 2006" authorityName="Osawa, Fujita &amp; Okuno" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B976EA97FF95AB66FA85FDD4FF2EFD05" italics="true" pageId="6" pageNumber="446">Pagurixus longipes</emphasis>
<bibRefCitation id="EF934B74FF95AB66FEACFD30FDB8FD05" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[256,608,655,681]" pageId="6" pageNumber="446" pagination="27 - 45" refId="ref6983" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148 (1), 27 - 45. https: // doi. org / 10.11646 / zootaxa. 1148.1.2" type="journal article" year="2006">Osawa, Fujita &amp; Okuno, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FDC2FD30FBCAFD05" authority="Komai &amp; Asakura, 1995" authorityName="Komai &amp; Asakura" authorityYear="1995" box="[622,1042,654,681]" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B976EA97FF95AB66FDC2FD30FD33FD04" box="[622,747,655,680]" italics="true" pageId="6" pageNumber="446">P. nomurai</emphasis>
<bibRefCitation id="EF934B74FF95AB66FD59FD30FBCAFD05" author="Komai, T. &amp; Asakura, A." box="[757,1042,654,681]" pageId="6" pageNumber="446" pagination="341 - 354" refId="ref5861" refString="Komai, T. &amp; Asakura, A. (1995) Pagurixus nomurai, new species, and addition al record of Pagurixus maorus (Nobili, 1906), hermit crabs from Kume-jima Island, the Ryukyus, Japan (Decapoda: Anomura: Paguridae). Journal of Crustacean Biology, 15 (2), 341 - 354. https: // doi. org / 10.2307 / 1548961" type="journal article" year="1995">Komai &amp; Asakura, 1995</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FB8CFD30FEF3FD61" authority="Osawa &amp; Okuno, 2007" authorityName="Osawa &amp; Okuno" authorityYear="2007" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B976EA97FF95AB66FB8CFD30FAFFFD05" box="[1056,1319,655,681]" italics="true" pageId="6" pageNumber="446">Pylopaguropsis bellula</emphasis>
<bibRefCitation id="EF934B74FF95AB66FA9CFD30FEF3FD61" author="Osawa, M. &amp; Okuno, J." pageId="6" pageNumber="446" pagination="29 - 46" refId="ref7048" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FE90FD0CFD23FD61" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[316,763,691,717]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="B976EA97FF95AB66FE90FD0CFE19FD61" box="[316,449,691,717]" italics="true" pageId="6" pageNumber="446">P. fimbriata</emphasis>
<bibRefCitation id="EF934B74FF95AB66FE66FD0CFD23FD61" author="McLaughlin, P. A. &amp; Haig, J." box="[458,763,691,717]" pageId="6" pageNumber="446" pagination="123 - 171" refId="ref6552" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FCA4FD0CFB9DFD61" authority="Asakura, 2000" authorityName="Asakura" authorityYear="2000" box="[776,1093,691,717]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="B976EA97FF95AB66FCA4FD0CFC4BFD61" box="[776,915,691,717]" italics="true" pageId="6" pageNumber="446">P. granulata</emphasis>
<bibRefCitation id="EF934B74FF95AB66FC30FD0CFB9DFD61" author="Asakura, A." box="[924,1093,691,717]" pageId="6" pageNumber="446" pagination="70 - 108" refId="ref5121" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura, 2000</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C024D06FF95AB66FBFDFD0CFF0AFD5D" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="B976EA97FF95AB66FBFDFD0CFB7EFD61" box="[1105,1190,691,717]" italics="true" pageId="6" pageNumber="446">P. keijii</emphasis>
<bibRefCitation id="EF934B74FF95AB66FB02FD0CFF0AFD5D" author="McLaughlin, P. A. &amp; Haig, J." pageId="6" pageNumber="446" pagination="123 - 171" refId="ref6552" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C024D06FF95AB66FEBFFD68FD17FD5D" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[275,719,727,753]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="B976EA97FF95AB66FEBFFD68FE49FD5C" box="[275,401,727,752]" italics="true" pageId="6" pageNumber="446">P. speciosa</emphasis>
<bibRefCitation id="EF934B74FF95AB66FE30FD68FD17FD5D" author="McLaughlin, P. A. &amp; Haig, J." box="[412,719,727,753]" pageId="6" pageNumber="446" pagination="123 - 171" refId="ref6552" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C024D06FF95AB66FD4CFD68FC8DFD5D" authorityName="Latreille" authorityYear="1802" box="[736,853,727,753]" class="Malacostraca" family="Paguridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
, nine species). The present new species represents the 13th hermit crab species found in the cave. Among these species, only
<taxonomicName id="4C024D06FF95AB66FC4DFD44FB3BFCB9" authorityName="McLaughlin" authorityYear="2002" box="[993,1251,763,789]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B976EA97FF95AB66FC4DFD44FB3BFCB9" box="[993,1251,763,789]" italics="true" pageId="6" pageNumber="446">Catapaguroides foresti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D06FF95AB66FB5DFD44FABDFCB9" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[1265,1381,763,789]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="B976EA97FF95AB66FB5DFD44FABDFCB9" box="[1265,1381,763,789]" italics="true" pageId="6" pageNumber="446">C. longior</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C024D06FF95AB66FF3BFCA0FE62FC95" authorityName="Asakura" authorityYear="2000" box="[151,442,799,825]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="B976EA97FF95AB66FF3BFCA0FE62FC95" box="[151,442,799,825]" italics="true" pageId="6" pageNumber="446">Pylopaguropsis granulata</emphasis>
</taxonomicName>
occurred in “silty room” zone leading to “innermost part” zone of Hedo Dome cave. The other species were found in “main tube” zone and the present new species is obtained from the area adjacent to “innermost part” zone.
</paragraph>
</subSubSection>
<subSubSection id="C318650EFF95AB66FF6BFC34FECCFC41" pageId="6" pageNumber="446" type="etymology">
<paragraph id="8BBD3685FF95AB66FF6BFC34FECCFC41" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<emphasis id="B976EA97FF95AB66FF6BFC34FE96FC09" bold="true" box="[199,334,907,933]" pageId="6" pageNumber="446">Etymology.</emphasis>
The specific name is derived from the combination of the Latin,
<emphasis id="B976EA97FF95AB66FB96FC34FB56FC08" box="[1082,1166,907,932]" italics="true" pageId="6" pageNumber="446">crassus</emphasis>
(= stout) and
<emphasis id="B976EA97FF95AB66FA9DFC34FAA2FC09" box="[1329,1402,907,933]" italics="true" pageId="6" pageNumber="446">oculus</emphasis>
(= eye), in reference to the inflated proximal portion of the ocular peduncle of the new species. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C318650EFF95AB67FF6BFC48FE88F8B0" lastPageId="7" lastPageNumber="447" pageId="6" pageNumber="446" type="discussion">
<paragraph id="8BBD3685FF95AB66FF6BFC48FBE1FAD5" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<emphasis id="B976EA97FF95AB66FF6BFC48FEE3FBBD" bold="true" box="[199,315,1015,1041]" pageId="6" pageNumber="446">Remarks.</emphasis>
In the preserved condition, the ocular peduncles of both the
<typeStatus id="54B98827FF95AB66FC6BFC48FBF1FBBD" box="[967,1065,1015,1041]" pageId="6" pageNumber="446" type="holotype">holotype</typeStatus>
and a
<typeStatus id="54B98827FF95AB66FBC0FC48FB14FBBD" box="[1132,1228,1015,1041]" pageId="6" pageNumber="446" type="paratype">paratype</typeStatus>
available for study have a strongly concave appearance on each distolateral margin including the cornea. Nevertheless, this condition is likely due to shrinkage by ethanol preservation because the ocular peduncles of the fresh specimens are seemingly only weakly concave on that margin (
<figureCitation id="13392A00FF95AB66FDE4FBDCFD7DFBD1" box="[584,677,1123,1149]" captionStart="FIGURE 4" captionStartId="7.[151,250,1292,1316]" captionTargetBox="[155,1432,777,1269]" captionTargetId="figure-362@7.[155,1432,777,1269]" captionTargetPageId="7" captionText="FIGURE 4. Eutrichopagurus crassoculus n. sp., paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7389486" httpUri="https://zenodo.org/record/7389486/files/figure.png" pageId="6" pageNumber="446">Fig. 4A</figureCitation>
). Meanwhile, the
<specimenCount id="9D04FD0CFF95AB66FCDFFBDCFC0EFBD1" box="[883,982,1123,1149]" pageId="6" pageNumber="446" type="generic">two type</specimenCount>
specimens have an oblong capsule-like structure on the ventromesial distal margin of each of the left and right antennal article 3 (
<figureCitation id="13392A00FF95AB66FBD0FB38FB09FB0D" box="[1148,1233,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1783]" captionTargetBox="[173,1418,188,1730]" captionTargetId="figure-17@2.[164,1423,181,1736]" captionTargetPageId="2" captionText="FIGURE 1. Eutrichopagurus crassoculus n. sp., holotype, male (sl 1.2 mm), RUMF-ZC 6115. A, carapace, dorsal view (setae omitted); B, shield and cephalic appendages, dorsal view (setae drawn only on left side; distolateral margin of ocular peduncle strongly concave due to shrinkage by ethanol preservation); C, left ocular peduncle and ocular acicle, mesial view; D, right antennular article 3 and flagella, lateral view (aesthetasc on proximal portion of dorsal flagellum drawn simplified); E, left antennal peduncle, ventral view (setae omitted; arrow showing capsule-like structure); F, left antennal flagellum, lateral view; G, left maxilliped 3, lateral view (flagellum of exopod concealed); H, same, ischium and basis, ventral view; I, left pereopod 4, lateral view; J, coxae of pereopods 5 and thoracic sternite 8, ventral view; K, basis and coxa of right pereopod 5, lateral view; L, same (sexual tube in extended condition); M, thoracic sternite 6, ventral view; N, telson, dorsal view. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7389474" httpUri="https://zenodo.org/record/7389474/files/figure.png" pageId="6" pageNumber="446">Fig. 1E</figureCitation>
). Similar capsulelike structures have been reported in
<taxonomicName id="4C024D06FF95AB66FDE4FB14FB96FB69" authority="Komai &amp; Poupin, 2012" authorityName="Komai &amp; Poupin" authorityYear="2012" box="[584,1102,1195,1221]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B976EA97FF95AB66FDE4FB14FCEDFB68" box="[584,821,1195,1221]" italics="true" pageId="6" pageNumber="446">Trichopagurus asper</emphasis>
<bibRefCitation id="EF934B74FF95AB66FC93FB14FB96FB69" author="Komai, T. &amp; Poupin, J." box="[831,1102,1195,1221]" pageId="6" pageNumber="446" pagination="56 - 68" refId="ref6077" refString="Komai, T. &amp; Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai &amp; Poupin, 2012</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D06FF95AB66FB27FB14FF0AFB45" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="tenuidactylus">
<emphasis id="B976EA97FF95AB66FB27FB14FAE7FB69" box="[1163,1343,1195,1221]" italics="true" pageId="6" pageNumber="446">T. tenuidactylus</emphasis>
<bibRefCitation id="EF934B74FF95AB66FAE5FB14FF0AFB45" author="Komai, T." pageId="6" pageNumber="446" pagination="274 - 282" refId="ref5736" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai, 2013</bibRefCitation>
</taxonomicName>
; such several structures are present on the ventral surfaces of both cheliped meri and only a single structure is present on the ventromesial margin of the right cheliped merus in the two species, respectively.
<bibRefCitation id="EF934B74FF95AB66FB76FB4CFF3AFA9D" author="Komai, T. &amp; Poupin, J." pageId="6" pageNumber="446" pagination="56 - 68" refId="ref6077" refString="Komai, T. &amp; Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai &amp; Poupin (2012)</bibRefCitation>
considered they possibly represent egg sacs of other invertebrates or parasites, but it is unknown if the capsule-like structures are identical among
<bibRefCitation id="EF934B74FF95AB66FDD2FA84FC7DFAF9" author="Komai, T. &amp; Poupin, J." box="[638,933,1339,1365]" pageId="6" pageNumber="446" pagination="56 - 68" refId="ref6077" refString="Komai, T. &amp; Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai &amp; Poupin’s (2012)</bibRefCitation>
,
<bibRefCitation id="EF934B74FF95AB66FC1DFA84FBBCFAF9" author="Komai, T." box="[945,1124,1339,1365]" pageId="6" pageNumber="446" pagination="274 - 282" refId="ref5736" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai’s (2013)</bibRefCitation>
, and the present specimens, and if they might be actually external ornamentation belonging to the hermit crabs.
</paragraph>
<paragraph id="8BBD3685FF95AB66FF6BFA3CFD11F885" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
In their description of
<taxonomicName id="4C024D06FF95AB66FE11FA3CFBAFFA31" authority="Komai, Rahayu &amp; Fujita, 2021" authorityName="Komai, Rahayu &amp; Fujita" authorityYear="2021" box="[445,1143,1411,1437]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B976EA97FF95AB66FE11FA3CFCC4FA31" box="[445,796,1411,1437]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus granulimanus</emphasis>
<bibRefCitation id="EF934B74FF95AB66FC8DFA3CFBAFFA31" author="Komai, T. &amp; Rahayu, D. L. &amp; Fujita, Y." box="[801,1143,1411,1437]" pageId="6" pageNumber="446" pagination="378 - 402" refId="ref6134" refString="Komai, T., Rahayu, D. L. &amp; Fujita, Y. (2021) Hermit crabs of the family Paguridae (Crustacea: Decapoda: Anomura) from Christmas and Cocos (Keeling) Islands, Indian Ocean Territory of Australia, with descriptions of three new species and reappraisal of the genus Kumepagurus Komai &amp; Osawa, 2012. Raffles Bulletin of Zoology, 69, 378 - 402. https: // doi. org / 10.26107 / RBZ- 2021 - 0059" type="journal article" year="2021">Komai, Rahayu &amp; Fujita, 2021</bibRefCitation>
</taxonomicName>
from Thunderdome Cave, a submarine cave of
<collectingCountry id="F3157615FF95AB66FED0FA18FDE0FA6D" box="[380,568,1447,1473]" name="Christmas Island" pageId="6" pageNumber="446">Christmas Island</collectingCountry>
in the southeastern Indian Ocean,
<bibRefCitation id="EF934B74FF95AB66FC15FA18FB4BFA6D" author="Komai, T. &amp; Rahayu, D. L. &amp; Fujita, Y." box="[953,1171,1447,1473]" pageId="6" pageNumber="446" pagination="378 - 402" refId="ref6134" refString="Komai, T., Rahayu, D. L. &amp; Fujita, Y. (2021) Hermit crabs of the family Paguridae (Crustacea: Decapoda: Anomura) from Christmas and Cocos (Keeling) Islands, Indian Ocean Territory of Australia, with descriptions of three new species and reappraisal of the genus Kumepagurus Komai &amp; Osawa, 2012. Raffles Bulletin of Zoology, 69, 378 - 402. https: // doi. org / 10.26107 / RBZ- 2021 - 0059" type="journal article" year="2021">
Komai
<emphasis id="B976EA97FF95AB66FBA7FA18FBE4FA6D" box="[1035,1084,1447,1473]" italics="true" pageId="6" pageNumber="446">et al</emphasis>
. (2021)
</bibRefCitation>
discussed the supposed morphological distinctions between the genera
<taxonomicName id="4C024D06FF95AB66FD1BFA74FCADFA49" authorityName="Komai" authorityYear="2015" box="[695,885,1483,1509]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FD1BFA74FCADFA49" box="[695,885,1483,1509]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D06FF95AB66FC1DFA74FB23FA49" authority="Bouvier, 1897" authorityName="Bouvier" authorityYear="1897" box="[945,1275,1483,1509]" class="Malacostraca" family="Paguridae" genus="Cestopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FC1DFA74FB95FA49" box="[945,1101,1483,1509]" italics="true" pageId="6" pageNumber="446">Cestopagurus</emphasis>
<bibRefCitation id="EF934B74FF95AB66FBFAFA74FB23FA49" author="Bouvier, E. L." box="[1110,1275,1483,1509]" pageId="6" pageNumber="446" pagination="229 - 233" refId="ref5219" refString="Bouvier, E. L. (1897) Sur deux paguriens nouveaux trouves par M. Coutiere dans le recif madreporique, Djibouti. Bulletin du Museum national d'Histoire naturelle, Paris, 6, 229 - 233." type="journal article" year="1897">Bouvier, 1897</bibRefCitation>
</taxonomicName>
. As in Komai
<emphasis id="B976EA97FF95AB66FF3BFA50FF14F9A5" box="[151,204,1519,1545]" italics="true" pageId="6" pageNumber="446">et al</emphasis>
.’s (2021) species, the assignment of the present new species to the genus
<taxonomicName id="4C024D06FF95AB66FB90FA50FB22F9A5" authorityName="Komai" authorityYear="2015" box="[1084,1274,1519,1545]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FB90FA50FB22F9A5" box="[1084,1274,1519,1545]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus</emphasis>
</taxonomicName>
is considered provisional, because only the male specimens were available for study, and the original diagnosis of the genus was based only on female characters. The present new species has diagnostic characters of
<taxonomicName id="4C024D06FF95AB66FBC1F988FAF3F9FD" authorityName="Komai" authorityYear="2015" box="[1133,1323,1591,1617]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FBC1F988FAF3F9FD" box="[1133,1323,1591,1617]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus</emphasis>
</taxonomicName>
including eleven pairs of deeply quadriserial phyllobranchiate gills, a prominent tuft of setae on the dorsodistal margin of the article 3 of the antennular peduncle, and the posterolateral margins of the telson each bearing a tuft of long setae (cf.
<bibRefCitation id="EF934B74FF95AB66FEAFF91CFE49F911" author="Komai, T." box="[259,401,1698,1725]" pageId="6" pageNumber="446" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">Komai 2015</bibRefCitation>
;
<bibRefCitation id="EF934B74FF95AB66FE30F91CFDB2F911" author="Komai, T. &amp; Rahayu, D. L. &amp; Fujita, Y." box="[412,618,1699,1725]" pageId="6" pageNumber="446" pagination="378 - 402" refId="ref6134" refString="Komai, T., Rahayu, D. L. &amp; Fujita, Y. (2021) Hermit crabs of the family Paguridae (Crustacea: Decapoda: Anomura) from Christmas and Cocos (Keeling) Islands, Indian Ocean Territory of Australia, with descriptions of three new species and reappraisal of the genus Kumepagurus Komai &amp; Osawa, 2012. Raffles Bulletin of Zoology, 69, 378 - 402. https: // doi. org / 10.26107 / RBZ- 2021 - 0059" type="journal article" year="2021">
Komai
<emphasis id="B976EA97FF95AB66FE43F91CFDF8F911" box="[495,544,1699,1725]" italics="true" pageId="6" pageNumber="446">et al</emphasis>
. 2021
</bibRefCitation>
). The right sexual tube of the new species is directed posterolaterally and filamentous in shape as described in
<taxonomicName id="4C024D06FF95AB66FD9DF978FD34F94D" authorityName="Komai, Rahayu &amp; Fujita" authorityYear="2021" box="[561,748,1735,1761]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B976EA97FF95AB66FD9DF978FD34F94D" box="[561,748,1735,1761]" italics="true" pageId="6" pageNumber="446">E. granulimanus</emphasis>
</taxonomicName>
, whereas that of
<taxonomicName id="4C024D06FF95AB66FC06F978FB39F94D" authority="Komai, 2015" authorityName="Komai" authorityYear="2015" box="[938,1249,1734,1761]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF95AB66FC06F978FB9DF94D" box="[938,1093,1735,1761]" italics="true" pageId="6" pageNumber="446">E. shirakawai</emphasis>
<bibRefCitation id="EF934B74FF95AB66FBE0F978FB39F94D" author="Komai, T." box="[1100,1249,1734,1761]" pageId="6" pageNumber="446" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">Komai, 2015</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="54B98827FF95AB66FAB8F978FA9CF94D" box="[1300,1348,1735,1761]" pageId="6" pageNumber="446">type</typeStatus>
species of the genus and described from shallow subtidal waters around Okinawa Island, Ryukyu Islands, is unknown because no male of the species has been recorded.
</paragraph>
<paragraph id="8BBD3685FF95AB66FF6BF88CFB60F815" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="446">
<taxonomicName id="4C024D06FF95AB66FF6BF88CFE5DF8E1" authorityName="Komai" authorityYear="2015" box="[199,389,1843,1869]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FF6BF88CFE5DF8E1" box="[199,389,1843,1869]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus</emphasis>
</taxonomicName>
is also substantially similar to
<taxonomicName id="4C024D06FF95AB66FD45F88CFB42F8E1" authority="de Saint Laurent, 1968" authorityName="de Saint Laurent" authorityYear="1968" box="[745,1178,1843,1869]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FD45F88CFC56F8E1" box="[745,910,1843,1869]" italics="true" pageId="6" pageNumber="446">Trichopagurus</emphasis>
de
<bibRefCitation id="EF934B74FF95AB66FC15F88CFB42F8E1" author="Saint Laurent, M. de" box="[953,1178,1843,1869]" pageId="6" pageNumber="446" pagination="210 - 222" refId="ref7115" refString="Saint Laurent, M. de. (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. II. Cestopagurus Bouvier (Crustaces Decapodes Paguridae). Bulletin du Museum national d'Histoire naturelle, Paris, 2 e Serie, 42, 210 - 222." type="journal article" year="1968">
Saint Laurent,
<quantity id="4CFA9B60FF95AB66FBCEF88CFB42F8E1" box="[1122,1178,1843,1869]" metricMagnitude="1" metricUnit="m" metricValue="4.99872" pageId="6" pageNumber="446" unit="in" value="1968.0">1968</quantity>
</bibRefCitation>
</taxonomicName>
in having the strongly produced triangular rostrum, a distinct tuft of setae on each posterolateral margin of the telson and the male right sexual tube being directed posterolaterally, but the deeply quadriserial, instead of biserial, gills distinguish the former from the latter (cf.
<bibRefCitation id="EF934B74FF95AB66FE13F820FD65F815" author="Komai, T. &amp; Osawa, M." box="[447,701,1950,1977]" pageId="6" pageNumber="446" pagination="1 - 20" refId="ref6010" refString="Komai, T. &amp; Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai &amp; Osawa 2005</bibRefCitation>
;
<bibRefCitation id="EF934B74FF95AB66FD65F820FC13F815" author="Komai, T. &amp; Poupin, J." box="[713,971,1951,1977]" pageId="6" pageNumber="446" pagination="56 - 68" refId="ref6077" refString="Komai, T. &amp; Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai &amp; Poupin 2012</bibRefCitation>
;
<bibRefCitation id="EF934B74FF95AB66FC7AF820FBBCF815" author="Komai, T." box="[982,1124,1951,1977]" pageId="6" pageNumber="446" pagination="274 - 282" refId="ref5736" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
,
<bibRefCitation id="EF934B74FF95AB66FBC3F821FB74F815" author="Komai, T." box="[1135,1196,1950,1977]" pageId="6" pageNumber="446" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BBD3685FF95AB67FF6BF87CFC54FD5D" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,150,753]" lastPageId="7" lastPageNumber="447" pageId="6" pageNumber="446">
Among two known species of the genus
<taxonomicName id="4C024D06FF95AB66FD26F87CFC90F871" authorityName="Komai" authorityYear="2015" box="[650,840,1987,2013]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA97FF95AB66FD26F87CFC90F871" box="[650,840,1987,2013]" italics="true" pageId="6" pageNumber="446">Eutrichopagurus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D06FF95AB66FCFAF87CFC20F871" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[854,1016,1987,2013]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF95AB66FCFAF87CFC20F871" box="[854,1016,1987,2013]" italics="true" pageId="6" pageNumber="446">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF95AB66FC53F87CFBE5F871" bold="true" box="[1023,1085,1987,2013]" pageId="6" pageNumber="446">
<taxonomicNameLabel id="A24557ECFF95AB66FC53F87CFBE5F871" box="[1023,1085,1987,2013]" pageId="6" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
may be closer to
<taxonomicName id="4C024D06FF95AB67FAADF87CFEF3FF1D" authority="Komai, 2015" authorityName="Komai" authorityYear="2015" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" lastPageNumber="447" order="Decapoda" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF95AB66FAADF87CFA44F871" box="[1281,1436,1987,2013]" italics="true" pageId="6" pageNumber="446">E. shirakawai</emphasis>
<bibRefCitation id="EF934B74FF94AB67FF3BFF28FEF3FF1D" author="Komai, T." box="[151,299,150,177]" pageId="7" pageNumber="447" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">
Komai,
<quantity id="4CFA9B60FF94AB67FF5FFF29FEF3FF1D" box="[243,299,150,177]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="7" pageNumber="447" unit="in" value="2015.0">2015</quantity>
</bibRefCitation>
</taxonomicName>
in having reduced corneas basally bearing sparse setae and no small corneous spines or bristle-like setae on the dorsal surface of the meri of the ambulatory legs, but is distinguishable by the following numerous characters. The proximolateral margin of each ocular peduncle is strongly convex in
<taxonomicName id="4C024D06FF94AB67FBE6FF60FB35FF55" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[1098,1261,223,249]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FBE6FF60FB35FF55" box="[1098,1261,223,249]" italics="true" pageId="7" pageNumber="447">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FB58FF60FAEAFF55" bold="true" box="[1268,1330,223,249]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FB58FF60FAEAFF55" box="[1268,1330,223,249]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, whereas it is gently convex in
<taxonomicName id="4C024D06FF94AB67FE3FFEBCFDE8FEB1" authorityName="Komai" authorityYear="2015" box="[403,560,259,285]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FE3FFEBCFDE8FEB1" box="[403,560,259,285]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
. The ocular acicle is short and ovate in the new species, instead of elongate and narrowly triangular in
<taxonomicName id="4C024D06FF94AB67FE61FE98FDB3FEED" authorityName="Komai" authorityYear="2015" box="[461,619,295,321]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FE61FE98FDB3FEED" box="[461,619,295,321]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
. The antennular peduncle overreaches the distal corneal margin by the full length of the article 3 and the distal one third of the article 2, rather only approximately the full length of the article
<quantity id="4CFA9B60FF94AB67FF48FED0FEC8FE25" box="[228,272,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="447" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C024D06FF94AB67FEBBFED0FE6AFE25" authorityName="Komai" authorityYear="2015" box="[279,434,367,393]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FEBBFED0FE6AFE25" box="[279,434,367,393]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
. The antennal flagellum has at most three short setae on the distal margin of each article, whereas three to five, short and long setae are present on the distal margin, which make the flagellum somewhat setose, in
<taxonomicName id="4C024D06FF94AB67FEA5FE08FE7DFE7D" authorityName="Komai" authorityYear="2015" box="[265,421,439,465]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FEA5FE08FE7DFE7D" box="[265,421,439,465]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
. The merus of maxilliped 3 is unarmed on both the dorsal and ventral margins, but it has distinct dorsodistal and ventromedian spines in
<taxonomicName id="4C024D06FF94AB67FD00FE64FC9FFE59" authorityName="Komai" authorityYear="2015" box="[684,839,475,501]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FD00FE64FC9FFE59" box="[684,839,475,501]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EF934B74FF94AB67FCD7FE64FBD2FE59" author="Komai, T." box="[891,1034,474,501]" pageId="7" pageNumber="447" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">Komai 2015</bibRefCitation>
: fig. 1B, E, F). The right cheliped is proportionally much shorter in
<taxonomicName id="4C024D06FF94AB67FE5BFE40FD43FDB5" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[503,667,511,537]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FE5BFE40FD43FDB5" box="[503,667,511,537]" italics="true" pageId="7" pageNumber="447">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FD0EFE40FD39FDB5" bold="true" box="[674,737,511,537]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FD0EFE40FD39FDB5" box="[674,737,511,537]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C024D06FF94AB67FC92FE40FC02FDB5" authorityName="Komai" authorityYear="2015" box="[830,986,511,537]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FC92FE40FC02FDB5" box="[830,986,511,537]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
; the chela and carpus are 1.8 versus 2.0 and 1.4–1.5 versus 2.1 times longer than the distal widths, respectively. The dactylus of the left cheliped is clearly longer than the palm in
<taxonomicName id="4C024D06FF94AB67FE37FDF8FDE5FDCD" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[411,573,583,609]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FE37FDF8FDE5FDCD" box="[411,573,583,609]" italics="true" pageId="7" pageNumber="447">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FDEFFDF8FD58FDCD" bold="true" box="[579,640,583,609]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FDEFFDF8FD58FDCD" box="[579,640,583,609]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, than in
<taxonomicName id="4C024D06FF94AB67FD72FDF8FCA0FDCD" authorityName="Komai" authorityYear="2015" box="[734,888,583,609]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FD72FDF8FCA0FDCD" box="[734,888,583,609]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
; it is 1.5 versus 0.9 length of the palm. The merus of both chelipeds of
<taxonomicName id="4C024D06FF94AB67FED1FDD4FDF8FD29" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[381,544,619,645]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FED1FDD4FDF8FD29" box="[381,544,619,645]" italics="true" pageId="7" pageNumber="447">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FD8BFDD4FDBEFD29" bold="true" box="[551,614,619,645]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FD8BFDD4FDBEFD29" box="[551,614,619,645]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has sparse bristle-like setae on the dorsodistal margin and adjacent to the ventromesial distal margin, but such setae are absent in
<taxonomicName id="4C024D06FF94AB67FCA2FD30FC72FD05" authorityName="Komai" authorityYear="2015" box="[782,938,655,681]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="shirakawai">
<emphasis id="B976EA97FF94AB67FCA2FD30FC72FD05" box="[782,938,655,681]" italics="true" pageId="7" pageNumber="447">E. shirakawai</emphasis>
</taxonomicName>
. The ambulatory legs are mostly smooth on all the surfaces, instead of the entire surfaces of the dactylus to carpus and the ventral parts of the lateral surfaces of the merus with numerous fine granules (cf.
<bibRefCitation id="EF934B74FF94AB67FD2CFD68FCD7FD5D" author="Komai, T." box="[640,783,726,753]" pageId="7" pageNumber="447" pagination="250 - 260" refId="ref5800" refString="Komai, T. (2015) A new genus and new species of Paguridae (Crustacea: Decapoda: Anomura) from shallow subtidal waters in Okinawa Island, the Ryukyu Islands, Japan. Zootaxa, 3918 (2), 250 - 260. http: // dx. doi. org / 10.11646 / zootaxa. 3918.2.6" type="journal article" year="2015">Komai 2015</bibRefCitation>
: figs. 2–4).
</paragraph>
<caption id="DF7D660DFF94AB67FF3BFAB3FC75FAE4" ID-DOI="http://doi.org/10.5281/zenodo.7389486" ID-Zenodo-Dep="7389486" httpUri="https://zenodo.org/record/7389486/files/figure.png" pageId="7" pageNumber="447" startId="7.[151,250,1292,1316]" targetBox="[155,1432,777,1269]" targetPageId="7">
<paragraph id="8BBD3685FF94AB67FF3BFAB3FC75FAE4" blockId="7.[151,1437,1292,1352]" pageId="7" pageNumber="447">
<emphasis id="B976EA97FF94AB67FF3BFAB3FECEFA88" bold="true" box="[151,278,1292,1316]" pageId="7" pageNumber="447">FIGURE 4.</emphasis>
<taxonomicName id="4C024D06FF94AB67FE8CFAB3FD95FA88" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[288,589,1292,1316]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FE8CFAB3FD95FA88" box="[288,589,1292,1316]" italics="true" pageId="7" pageNumber="447">Eutrichopagurus crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FDFBFAB3FD4BFA88" bold="true" box="[599,659,1292,1316]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FDFBFAB3FD4BFA88" box="[599,659,1292,1316]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, male (sl 1.2 mm), RUMF-ZC 61156. A, entire animal in fresh condition, dorsal view; B, living animal carrying gastropod shell, dorsal view.
</paragraph>
</caption>
<paragraph id="8BBD3685FF94AB67FF6BFAC9FE88F8B0" blockId="7.[151,1437,1398,1820]" pageId="7" pageNumber="447">
Besides the strongly inflated proximolateral margin of the ocular peduncle, the reduced corneas, and the absence of small corneous spines or bristle-like setae on the dorsal surface of the ambulatory meri,
<taxonomicName id="4C024D06FF94AB67FB01FA24FA8AFA18" authority="Osawa &amp; Fujita, 2022" authorityName="Osawa &amp; Fujita" authorityYear="2022" box="[1197,1362,1434,1460]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="crassoculus" status="sp. nov.">
<emphasis id="B976EA97FF94AB67FB01FA24FA8AFA18" box="[1197,1362,1434,1460]" italics="true" pageId="7" pageNumber="447">E. crassoculus</emphasis>
</taxonomicName>
<emphasis id="B976EA97FF94AB67FAF7FA25FA44FA18" bold="true" box="[1371,1436,1434,1460]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A24557ECFF94AB67FAF7FA25FA44FA18" box="[1371,1436,1434,1460]" pageId="7" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C024D06FF94AB67FE85FA00FE3EFA74" authorityName="Komai, Rahayu &amp; Fujita" authorityYear="2021" box="[297,486,1470,1496]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B976EA97FF94AB67FE85FA00FE3EFA74" box="[297,486,1470,1496]" italics="true" pageId="7" pageNumber="447">E. granulimanus</emphasis>
</taxonomicName>
in the following particulars: the comparatively longer shield (1.1–1.2 times versus approximately as long as wide); the proportionally longer article 3 of the antennular peduncle (3.2 versus 2.8 times as long as high); the merus of maxilliped 3 being unarmed on the dorsal and ventral margins (a dorsodistal and a ventromedian spines are present in
<taxonomicName id="4C024D06FF94AB67FD84F994FD3CF9E8" authorityName="Komai, Rahayu &amp; Fujita" authorityYear="2021" box="[552,740,1578,1604]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B976EA97FF94AB67FD84F994FD3CF9E8" box="[552,740,1578,1604]" italics="true" pageId="7" pageNumber="447">E. granulimanus</emphasis>
</taxonomicName>
); the proportionally longer dactylus of the right cheliped (0.7 versus 0.4 length of the palm); the proportionally shorter carpus of the right cheliped (1.4–1.5 versus 1.7 times longer than distal widths); the palm of the left cheliped with numerous minute granules, a distinct median elevation on the dorsal surface, and clearly delimited dorsomesial and dorsolateral margins (the dorsal surface being smooth and gently convex, without the delineation of dorsolateral and dorsomesial margins in
<taxonomicName id="4C024D06FF94AB67FBDFF904FAE8F978" authorityName="Komai, Rahayu &amp; Fujita" authorityYear="2021" box="[1139,1328,1722,1748]" class="Malacostraca" family="Paguridae" genus="Eutrichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="granulimanus">
<emphasis id="B976EA97FF94AB67FBDFF904FAE8F978" box="[1139,1328,1722,1748]" italics="true" pageId="7" pageNumber="447">E. granulimanus</emphasis>
</taxonomicName>
); and the dactyli of the ambulatory legs with less number of ventral spines (three to six versus eight or nine) (cf.
<bibRefCitation id="EF934B74FF94AB67FABEF961FF0BF8B0" author="Komai, T. &amp; Rahayu, D. L. &amp; Fujita, Y." pageId="7" pageNumber="447" pagination="378 - 402" refId="ref6134" refString="Komai, T., Rahayu, D. L. &amp; Fujita, Y. (2021) Hermit crabs of the family Paguridae (Crustacea: Decapoda: Anomura) from Christmas and Cocos (Keeling) Islands, Indian Ocean Territory of Australia, with descriptions of three new species and reappraisal of the genus Kumepagurus Komai &amp; Osawa, 2012. Raffles Bulletin of Zoology, 69, 378 - 402. https: // doi. org / 10.26107 / RBZ- 2021 - 0059" type="journal article" year="2021">
Komai
<emphasis id="B976EA97FF94AB67FAC9F960FA4DF954" box="[1381,1429,1758,1784]" italics="true" pageId="7" pageNumber="447">et al</emphasis>
. 2021
</bibRefCitation>
: figs. 2, 3).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink