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<document id="70BCAC7546BD2C9DEF45FB9242C8DD4D" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55522FFDDFF4AFC4BFCE6BED2" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus secretus Bernhauer 1935, comb. nov." docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="26" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="5F4DA0A2EA98086D3A84D77F70FBC3D5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6D965D4A58BFD8A1AC58C7D15D4F1887">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="DC53917C235CB101E0D215FE6A3DAE04" type="personal">
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<mods:namePart id="02ABA62D541C62BA9FDF33C4FE9BFA4D">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="87780A2C099DF9CD8329DDA9C9B563D3" type="personal">
<mods:role id="061176BEBE07530E7BCEBC65514477E3">
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<mods:namePart id="B92BD09E316EDA47FEAE31328A21A60D">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="63DB2C76E6DA4056AFEB25F8C4F39BE1">text</mods:typeOfResource>
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<mods:title id="3A65940925BBA66A7A937D66814DCFC4">Zootaxa</mods:title>
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<mods:part id="DC9C81CD597A8A16E4786D0CE3C9D680">
<mods:date id="BEE9BB81D470AA08D49AB44715DD1880">2015</mods:date>
<mods:detail id="5EE0DC1BB77FBBD5237CA98A834D12EA" type="volume">
<mods:number id="11858C6CF48D0997C07C81141AC7F323">4038</mods:number>
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<mods:start id="15768EF584E72C0857010D15ED09531F">1</mods:start>
<mods:end id="CCE3592012165D080EEA6AC7EDEF6E8A">94</mods:end>
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<mods:classification id="98E164BB410518523FF3D2FF2BC8D9E3">journal article</mods:classification>
<mods:identifier id="C33C0A87FEC496CAA07610CFF09490E5" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="0FCFF7A97B40472A89EAA02DEDFC2A82" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
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<treatment id="039D87F55522FFDDFF4AFC4BFCE6BED2" ID-DOI="http://doi.org/10.5281/zenodo.6101623" ID-GBIF-Taxon="127693953" ID-Zenodo-Dep="6101623" LSID="urn:lsid:plazi:treatment:039D87F55522FFDDFF4AFC4BFCE6BED2" httpUri="http://treatment.plazi.org/id/039D87F55522FFDDFF4AFC4BFCE6BED2" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="C32E65685522FFD8FF4AFC4BFEA7BA0D" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8B8B36E35522FFD8FF4AFC4BFCC8BA22" blockId="25.[151,823,1005,1064]" box="[151,823,1005,1031]" pageId="25" pageNumber="26">
<heading id="D0C3818F5522FFD8FF4AFC4BFCC8BA22" bold="true" box="[151,823,1005,1031]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B940EAF15522FFD8FF4AFC4BFCC8BA22" bold="true" box="[151,823,1005,1031]" pageId="25" pageNumber="26">
<taxonomicName id="4C344D605522FFD8FF4AFC4BFD56BA22" ID-CoL="56RYL" authority="Bernhauer, 1935" authorityName="Bernhauer" authorityYear="1935" box="[151,681,1005,1031]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus" status="comb. nov.">
<emphasis id="B940EAF15522FFD8FF4AFC4BFE3DBA22" bold="true" box="[151,450,1005,1031]" italics="true" pageId="25" pageNumber="26">Thyreocephalus secretus</emphasis>
(
<bibRefCitation id="EFA54B125522FFD8FE0DFC4BFD5EBA22" author="Bernhauer" box="[464,673,1005,1031]" pageId="25" pageNumber="26" refString="Bernhauer, M. (1935) Neue Staphyliniden vom belgischen Kongo. Revue de Zoologie et de Botanique Africaines, 27 (1), 97 - 106." type="journal article" year="1935">Bernhauer, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A273578A5522FFD8FD6CFC4BFCC8BA22" box="[689,823,1005,1031]" pageId="25" pageNumber="26">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36E35522FFD8FF4AFBA9FEA7BA0D" blockId="25.[151,823,1005,1064]" box="[151,344,1039,1064]" pageId="25" pageNumber="26">
(
<figureCitation id="130F2A665522FFD8FF42FBA9FF03BA0D" box="[159,252,1039,1064]" captionStart="FIGURE 29" captionStartId="16.[151,250,1593,1615]" captionTargetBox="[162,1433,201,1578]" captionTargetId="figure@16.[151,1436,193,1581]" captionTargetPageId="16" captionText="FIGURE 29. Distribution of species in Thyreocephalus pilosus group (● = T. africanus (Bernhauer), Δ = T. brunneiventris (Tottenham), ▲ = T. pilosus (Roth), ☆ T. pseudoafricanus sp. nov., ○ = T. secretus (Bernhauer), ★ = T. tsingidianus sp. nov.)." httpUri="https://zenodo.org/record/289884/files/figure.png" pageId="25" pageNumber="26">Figs. 29</figureCitation>
,
<figureCitation id="130F2A665522FFD8FED5FBA9FEAFBA0D" box="[264,336,1039,1064]" captionStart="FIGURES 51 56" captionStartId="26.[151,264,1911,1933]" captionTargetBox="[198,1405,209,1872]" captionTargetId="figure@26.[181,1405,193,1890]" captionTargetPageId="26" captionText="FIGURES 51 56. Thyreocephalus secretus (Bernhauer); 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere." httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="25" pageNumber="26">5156</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF4AFBFEFCF5BA4B" box="[151,778,1112,1134]" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8B8B36E35522FFD8FF4AFBFEFCF5BA4B" blockId="25.[151,778,1112,1134]" box="[151,778,1112,1134]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB2411CD5522FFD8FF4AFBFEFCF5BA4B" box="[151,778,1112,1134]" pageId="25" pageNumber="26">
<taxonomicName id="4C344D605522FFD8FF4AFBFEFDD7BA4B" ID-CoL="3CBNS" authority="Bernhauer, 1935: 101" authorityName="Bernhauer" authorityPageNumber="101" authorityYear="1935" box="[151,552,1112,1134]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FF4AFBFEFEBBBA4B" box="[151,324,1112,1134]" italics="true" pageId="25" pageNumber="26">Eulissus secretus</emphasis>
<treatmentCitation id="0A9510F25522FFD8FE97FBFEFDD7BA4B" author="Bernhauer" box="[330,552,1112,1134]" page="101" pageId="25" pageNumber="26" year="1935">
<bibRefCitation id="EFA54B125522FFD8FE97FBFEFE0ABA4B" author="Bernhauer" box="[330,501,1112,1134]" pageId="25" pageNumber="26" refString="Bernhauer, M. (1935) Neue Staphyliniden vom belgischen Kongo. Revue de Zoologie et de Botanique Africaines, 27 (1), 97 - 106." type="journal article" year="1935">Bernhauer, 1935</bibRefCitation>
: 101
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25522FFD8FDEFFBFFFCF9BA4B" author="Herman" box="[562,774,1112,1134]" page="3629" pageId="25" pageNumber="26" year="2001">
<bibRefCitation id="EFA54B125522FFD8FDEFFBFFFD39BA4B" author="Herman" box="[562,710,1112,1134]" pageId="25" pageNumber="26" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3629
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF4AFB3CFDA3BBF6" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8B8B36E35522FFD8FF4AFB3CFC9CBA96" blockId="25.[151,1437,1178,2031]" box="[151,867,1178,1203]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF4AFB3CFECABA96" bold="true" box="[151,309,1178,1203]" pageId="25" pageNumber="26">
<typeStatus id="548F88415522FFD8FF4AFB3CFF2DBA96" box="[151,210,1178,1203]" pageId="25" pageNumber="26">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32376735522FFD8FEE1FB3CFD62BA96" box="[316,669,1178,1203]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">Democratic Republic of Congo</collectingCountry>
, Lulua, Kapanga.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AFB18FD44BB42" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFB18FD74BAF2" bold="true" box="[199,651,1214,1239]" pageId="25" pageNumber="26">
<typeStatus id="548F88415522FFD8FF1AFB18FEFDBAF2" box="[199,258,1214,1239]" pageId="25" pageNumber="26">Type</typeStatus>
material examined.
<typeStatus id="548F88415522FFD8FDC8FB18FD74BAF2" box="[533,651,1214,1239]" pageId="25" pageNumber="26" type="lectotype">Lectotype</typeStatus>
</emphasis>
(by present designation) ♂:
<collectingCountry id="F32376735522FFD8FC23FB18FEFEBADE" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FC23FB18FEFEBADE" bold="true" pageId="25" pageNumber="26">DEMOCRATIC REPUBLIC OF CONGO</emphasis>
</collectingCountry>
: “Musée du
<collectingCountry id="F32376735522FFD8FE73FB45FE04BADE" box="[430,507,1251,1275]" name="Republic of the Congo" pageId="25" pageNumber="26">Congo</collectingCountry>
, Lulua: Kapanga”, “
<date id="FF8A10235522FFD8FD20FB45FC98BADE" box="[765,871,1251,1275]" pageId="25" pageNumber="26" value="1932-12">XII.1932</date>
, Overlaet”, “
<taxonomicName id="4C344D605522FFD8FBD0FB42FB40BADE" authority="Brnh." authorityName="Brnh." box="[1037,1215,1250,1275]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FBD0FB42FB97BADE" box="[1037,1128,1252,1275]" italics="true" pageId="25" pageNumber="26">secretus</emphasis>
, Brnh.
</taxonomicName>
<typeStatus id="548F88415522FFD8FB12FB45FAE9BADE" box="[1231,1302,1251,1275]" pageId="25" pageNumber="26">Typus</typeStatus>
,
<taxonomicName id="4C344D605522FFD8FAF6FB44FA77BADE" box="[1323,1416,1250,1275]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15522FFD8FAF6FB44FA77BADE" box="[1323,1416,1250,1275]" italics="true" pageId="25" pageNumber="26">Eulissus</emphasis>
</taxonomicName>
”, “
<taxonomicName id="4C344D605522FFD8FF7EFAA0FDD0BB3A" authority="Bernh." authorityName="Bernh." box="[163,559,1286,1311]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FF7EFAA0FE3FBB3B" box="[163,448,1286,1311]" italics="true" pageId="25" pageNumber="26">Thyreocephalus secretus</emphasis>
(Bernh.)
</taxonomicName>
, J. Janák det. 2006”. The specimen is chosen as
<typeStatus id="548F88415522FFD8FB7FFAA0FAF2BB3A" box="[1186,1293,1286,1311]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
. The labels “
<typeStatus id="548F88415522FFD8FF7CFA8DFEDFBB66" box="[161,288,1323,1347]" pageId="25" pageNumber="26" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="4C344D605522FFD8FEF6FA8CFD05BB66" authority="Bh., Janak &amp; Bordoni" authorityName="Bh., Janak &amp; Bordoni" box="[299,762,1322,1347]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FEF6FA8CFE11BB66" box="[299,494,1322,1347]" italics="true" pageId="25" pageNumber="26">Eulissus secretus</emphasis>
Bh., Janák &amp; Bordoni
</taxonomicName>
des. 2014” and “
<taxonomicName id="4C344D605522FFD8FC11FA8CFAD4BB66" authority="Bh." authorityName="Bh." box="[972,1323,1322,1347]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FC11FA8CFB1BBB66" box="[972,1252,1322,1347]" italics="true" pageId="25" pageNumber="26">Thyreocephalus secretus</emphasis>
(Bh.)
</taxonomicName>
, Janák &amp; Bordoni det. 2014” were added to this specimen.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AFAD4FDA3BBF6" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFAD4FBB4BBAE" bold="true" box="[199,1099,1394,1419]" pageId="25" pageNumber="26">
Additional material examined.
<collectingCountry id="F32376735522FFD8FD9BFAD4FBB4BBAE" box="[582,1099,1394,1419]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">DEMOCRATIC REPUBLIC OF CONGO</collectingCountry>
</emphasis>
:
<specimenCount id="9D32FD6A5522FFD8FB80FAD5FB74BBAF" box="[1117,1163,1395,1419]" pageId="25" pageNumber="26" type="generic">1 ♂</specimenCount>
, 1 ♀: Lulua: Kapanga,
<date id="FF8A10235522FFD8FF4AFA30FF09BB8A" box="[151,246,1430,1455]" pageId="25" pageNumber="26" value="1932-12">xii.1932</date>
, F. G. Overlaet (
<collectionCode id="ED25AE265522FFD8FE6CFA31FDF7BB8A" box="[433,520,1431,1455]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="25" pageNumber="26">MRAC</collectionCode>
,
<collectionCode id="ED25AE265522FFD8FDC9FA31FDABBB8A" box="[532,596,1431,1455]" pageId="25" pageNumber="26">JJRC</collectionCode>
);
<specimenCount id="9D32FD6A5522FFD8FDB4FA31FD68BB8B" box="[617,663,1431,1455]" pageId="25" pageNumber="26" type="generic">3 ♂</specimenCount>
, 1 ♀: Lulua: Kapanga,
<date id="FF8A10235522FFD8FC78FA30FC11BB8A" box="[933,1006,1430,1455]" pageId="25" pageNumber="26" value="1933-01">i.1933</date>
, F. G. Overlaet (
<collectionCode id="ED25AE265522FFD8FB77FA31FAFCBB8A" box="[1194,1283,1431,1455]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="25" pageNumber="26">MRAC</collectionCode>
);
<specimenCount id="9D32FD6A5522FFD8FACAFA31FABDBB8B" box="[1303,1346,1431,1455]" pageId="25" pageNumber="26" type="generic">1 ♂</specimenCount>
: Lulua: Kapanga,
<date id="FF8A10235522FFD8FED5FA1CFEA6BBF6" box="[264,345,1466,1491]" pageId="25" pageNumber="26" value="1933-02">ii.1933</date>
, F. G. Overlaet (
<collectionCode id="ED25AE265522FFD8FDD2FA1DFDAFBBF6" box="[527,592,1467,1491]" pageId="25" pageNumber="26">JJRC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF1AFA78FE7BB982" pageId="25" pageNumber="26" type="description">
<paragraph id="8B8B36E35522FFD8FF1AFA78FB01B88E" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFA78FEA6BBD2" bold="true" box="[199,345,1502,1527]" pageId="25" pageNumber="26">Description.</emphasis>
Body length
<quantity id="4CCC9B065522FFD8FE27FA78FD71BBD3" box="[506,654,1502,1527]" metricMagnitude="-2" metricUnit="m" metricValue="2.025" metricValueMax="2.15" metricValueMin="1.9" pageId="25" pageNumber="26" unit="mm" value="20.25" valueMax="21.5" valueMin="19.0">1921.5 mm</quantity>
; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B065522FFD8FF4AF9A4FED1B83F" box="[151,302,1538,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.2" metricValueMin="1.05" pageId="25" pageNumber="26" unit="mm" value="11.25" valueMax="12.0" valueMin="10.5">10.512 mm</quantity>
. Head, pronotum and scutellum black, elytra and abdomen reddish yellow, metasternum black, abdominal tergites 3 and 4 with median black spots; all sternites yellowish, antennae and legs brown dark. Shiny. Head and pronotum with micro-punctation. Head and pronotum and related punctation as in
<figureCitation id="130F2A665522FFD8FB6DF9ECFAFCB846" box="[1200,1283,1610,1635]" captionStart="FIGURES 51 56" captionStartId="26.[151,264,1911,1933]" captionTargetBox="[198,1405,209,1872]" captionTargetId="figure@26.[181,1405,193,1890]" captionTargetPageId="26" captionText="FIGURES 51 56. Thyreocephalus secretus (Bernhauer); 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere." httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="25" pageNumber="26">Fig. 51</figureCitation>
; labrum as in
<figureCitation id="130F2A665522FFD8FF4AF9C8FF16B8A2" box="[151,233,1646,1671]" captionStart="FIGURES 51 56" captionStartId="26.[151,264,1911,1933]" captionTargetBox="[198,1405,209,1872]" captionTargetId="figure@26.[181,1405,193,1890]" captionTargetPageId="26" captionText="FIGURES 51 56. Thyreocephalus secretus (Bernhauer); 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere." httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="25" pageNumber="26">Fig. 52</figureCitation>
. Elytra as long as pronotum, dilated posteriad and there wider than pronotum, with rounded humeral angles. Surface with sparse punctation, arranged in three series, one near suture, one median and one lateral.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AF910FA74B932" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<collectingCountry id="F32376735522FFD8FF1AF910FEFBB8EA" box="[199,260,1718,1743]" name="Malta" pageId="25" pageNumber="26">Male</collectingCountry>
. Lateral part of temple with series of moderately coarse punctures. Tergite 10 and sternite 9 of male genital segment as in
<figureCitation id="130F2A665522FFD8FE5AF97CFDF2B8D6" box="[391,525,1754,1779]" captionStart="FIGURES 51 56" captionStartId="26.[151,264,1911,1933]" captionTargetBox="[198,1405,209,1872]" captionTargetId="figure@26.[181,1405,193,1890]" captionTargetPageId="26" captionText="FIGURES 51 56. Thyreocephalus secretus (Bernhauer); 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere." httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="25" pageNumber="26">Figs. 53, 54</figureCitation>
. Aedeagus (
<figureCitation id="130F2A665522FFD8FD49F97CFCE6B8D6" box="[660,793,1754,1779]" captionStart="FIGURES 51 56" captionStartId="26.[151,264,1911,1933]" captionTargetBox="[198,1405,209,1872]" captionTargetId="figure@26.[181,1405,193,1890]" captionTargetPageId="26" captionText="FIGURES 51 56. Thyreocephalus secretus (Bernhauer); 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere." httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="25" pageNumber="26">Figs. 55, 56</figureCitation>
) ovoid, elongate,
<quantity id="4CCC9B065522FFD8FC3CF97CFBBBB8D7" box="[993,1092,1754,1779]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="25" pageNumber="26" unit="mm" value="1.65">1.65 mm</quantity>
long, with sub-triangular, long median lobe, with rounded apex; parameres symmetrical; inner sac narrow and long, apparently devoid of scales.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AF884FBC7B91F" blockId="25.[151,1437,1178,2031]" box="[199,1080,1826,1851]" pageId="25" pageNumber="26">Female. Lateral parts of temple impunctate or at most with few fine punctures.</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AF8E0FE7BB982" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AF8E0FE2EB97A" bold="true" box="[199,465,1862,1887]" pageId="25" pageNumber="26">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D605522FFD8FE01F8E0FD0BB97B" box="[476,756,1862,1887]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FE01F8E0FD0BB97B" box="[476,756,1862,1887]" italics="true" pageId="25" pageNumber="26">Thyreocephalus secretus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C344D605522FFD8FC0EF8E1FB68B97A" box="[979,1175,1862,1887]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneiventris">
<emphasis id="B940EAF15522FFD8FC0EF8E1FB68B97A" box="[979,1175,1862,1887]" italics="true" pageId="25" pageNumber="26">T. brunneiventris</emphasis>
</taxonomicName>
and differs in smaller black spots on the base of the abdomen, yellow lateral margins of tergite 3 and in rounded apical margin of median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF1AF814FEADB9CA" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8B8B36E35522FFD8FF1AF814FEADB9CA" blockId="25.[151,1437,1178,2031]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AF814FE9FB9EE" bold="true" box="[199,352,1970,1995]" pageId="25" pageNumber="26">Distribution.</emphasis>
The species is hitherto known only from the
<typeStatus id="548F88415522FFD8FCAEF815FC5CB9EE" box="[883,931,1971,1995]" pageId="25" pageNumber="26">type</typeStatus>
locality situated in
<collectingCountry id="F32376735522FFD8FB57F814FF1DB9CA" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">Democratic Republic of Congo</collectingCountry>
(
<figureCitation id="130F2A665522FFD8FF2CF870FEBAB9CA" box="[241,325,2006,2031]" captionStart="FIGURE 29" captionStartId="16.[151,250,1593,1615]" captionTargetBox="[162,1433,201,1578]" captionTargetId="figure@16.[151,1436,193,1581]" captionTargetPageId="16" captionText="FIGURE 29. Distribution of species in Thyreocephalus pilosus group (● = T. africanus (Bernhauer), Δ = T. brunneiventris (Tottenham), ▲ = T. pilosus (Roth), ☆ T. pseudoafricanus sp. nov., ○ = T. secretus (Bernhauer), ★ = T. tsingidianus sp. nov.)." httpUri="https://zenodo.org/record/289884/files/figure.png" pageId="25" pageNumber="26">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4B666B5521FFDBFF4AF8D1FDE2B989" httpUri="https://zenodo.org/record/289889/files/figure.png" pageId="26" pageNumber="27" targetBox="[198,1405,209,1872]" targetPageId="26">
<paragraph id="8B8B36E35521FFDBFF4AF8D1FDE2B989" blockId="26.[151,1435,1911,1964]" pageId="26" pageNumber="27">
<emphasis id="B940EAF15521FFDBFF4AF8D1FEB0B9A8" bold="true" box="[151,335,1911,1933]" pageId="26" pageNumber="27">FIGURES 5156</emphasis>
.
<taxonomicName id="4C344D605521FFDBFE83F8D1FD1FB9A8" authority="Bernhauer" authorityName="Bernhauer" box="[350,736,1911,1934]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15521FFDBFE83F8D1FDA5B9AB" box="[350,602,1911,1934]" italics="true" pageId="26" pageNumber="27">Thyreocephalus secretus</emphasis>
(Bernhauer)
</taxonomicName>
; 51, head and pronotum; 52, labrum; 53, male tergite 10; 54, male sternite 9; 55, aedeagus; 56, paramere.
</paragraph>
</caption>
<caption id="DF4B666B5520FFDAFF4AF814FB7CB9C2" httpUri="https://zenodo.org/record/289890/files/figure.png" pageId="27" pageNumber="28" targetBox="[225,1367,193,1936]" targetPageId="27">
<paragraph id="8B8B36E35520FFDAFF4AF814FB7CB9C2" blockId="27.[151,1436,1970,2023]" pageId="27" pageNumber="28">
<emphasis id="B940EAF15520FFDAFF4AF814FEB0B9ED" bold="true" box="[151,335,1970,1992]" pageId="27" pageNumber="28">FIGURES 5763</emphasis>
.
<taxonomicName id="4C344D605520FFDAFE80F814FCDDB9ED" authority="Tottenham" authorityName="Tottenham" box="[349,802,1970,1992]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brunneiventris">
<emphasis id="B940EAF15520FFDAFE80F814FD65B9ED" box="[349,666,1970,1992]" italics="true" pageId="27" pageNumber="28">Thyreocephalus brunneiventris</emphasis>
(Tottenham)
</taxonomicName>
; 57, head and pronotum; 58, labrum; 59, male tergite 10; 60, male sternite 9; 61, aedeagus of paratype; 62, apical part of median lobe of holotype; 63, paramere.
</paragraph>
</caption>
<subSubSection id="C32E65685527FFDDFF1AFF31FCE6BED2" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B8B36E35527FFDDFF1AFF31FCE6BED2" blockId="28.[151,1436,151,248]" pageId="28" pageNumber="29">
<emphasis id="B940EAF15527FFDDFF1AFF31FEC4BE95" bold="true" box="[199,315,151,176]" pageId="28" pageNumber="29">Remarks.</emphasis>
<collectingCountry id="F32376735527FFDDFE99FF3EFE9CBE95" box="[324,355,152,176]" name="American Samoa" pageId="28" pageNumber="29">As</collectingCountry>
the superior line of pronotal hypomeron is turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto anterior margin of pronotum, the species was transferred from
<taxonomicName id="4C344D605527FFDDFE02FF79FDC3BEDD" box="[479,572,223,248]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15527FFDDFE02FF79FDC3BEDD" box="[479,572,223,248]" italics="true" pageId="28" pageNumber="29">Eulissus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C344D605527FFDDFDBDFF79FCEDBEDD" box="[608,786,223,248]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15527FFDDFDBDFF79FCEDBEDD" box="[608,786,223,248]" italics="true" pageId="28" pageNumber="29">Thyreocephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>