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<document id="516B197D85CFFBF66B9495C596957ED5" ID-DOI="10.5281/zenodo.172729" ID-GBIF-Dataset="b184b252-bc32-4fc8-801b-9a8d9e4c64fe" ID-ISSN="1175­5326" ID-Zenodo-Dep="172729" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761247542" checkinUser="plazi" docAuthor="Asakura, Akira" docDate="2006" docId="039D87E1C4091F624B5CFE41DAAFFDAB" docLanguage="en" docName="zt01230p054.pdf" docOrigin="Zootaxa 1230" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Dardanus umbella Asakura, 2006, sp. nov." docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFA4FF99C4131F464A54FFA5D90FFF8D" masterDocTitle="Dardanus sanguinocarpus Degener, 1925 (Crustacea: Decapoda: Anomura: Diogenidae) species complex: a redescription of D. sanguinocarpus and descriptions of three new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="27" updateTime="1698215190567" updateUser="plazi">
<mods:mods id="BAF6593B5FC864537A98978FBE681249" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DE00763079F133EE3F42F78A52384752">Dardanus sanguinocarpus Degener, 1925 (Crustacea: Decapoda: Anomura: Diogenidae) species complex: a redescription of D. sanguinocarpus and descriptions of three new species</mods:title>
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<mods:namePart id="2196612D6426B6421FCF31CB739DCE98">Asakura, Akira</mods:namePart>
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<mods:title id="347FE20CCA37FE75CE825692853B8EA4">Zootaxa</mods:title>
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<mods:part id="02C8C72F62F135382AE4513F193A4315">
<mods:date id="F52E453830B8D90C28930FD433F56D2D">2006</mods:date>
<mods:detail id="0C11E0724B8B4A88F1A42042B5864902" type="volume">
<mods:number id="5F19FE3698F994D9B45C41BFAD7C74DB">1230</mods:number>
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<mods:classification id="FD3EF0DD13EF8FC90109F6C3AB84D0A5">journal article</mods:classification>
<mods:identifier id="B4314DDA81BE0F34B2BE6C2C1FCCB20F" type="DOI">10.5281/zenodo.172729</mods:identifier>
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<mods:identifier id="419473ED080E250FDD9F6AD498C82BF2" type="ISSN">1175­5326</mods:identifier>
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<treatment id="039D87E1C4091F624B5CFE41DAAFFDAB" ID-DOI="http://doi.org/10.5281/zenodo.6259129" ID-GBIF-Taxon="119344937" ID-Zenodo-Dep="6259129" LSID="urn:lsid:plazi:treatment:039D87E1C4091F624B5CFE41DAAFFDAB" httpUri="http://treatment.plazi.org/id/039D87E1C4091F624B5CFE41DAAFFDAB" lastPageId="36" lastPageNumber="37" pageId="26" pageNumber="27">
<subSubSection id="C32E657CC4091F5C4B5CFE41D8DDFDAC" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B8B36F7C4091F5C4B5CFE41DB40FE70" blockId="26.[264,591,483,545]" box="[264,591,483,510]" pageId="26" pageNumber="27">
<heading id="D0C3819BC4091F5C4B5CFE41DB40FE70" bold="true" box="[264,591,483,510]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B940EAE5C4091F5C4B5CFE41DB40FE70" bold="true" box="[264,591,483,510]" pageId="26" pageNumber="27">
<taxonomicName id="4C344D74C4091F5C4B5CFE41D8E5FE73" ID-CoL="36e9230f-2360-492e-a694-ea28a3b04cc9" box="[264,490,484,510]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C4091F5C4B5CFE41D8E5FE73" bold="true" box="[264,490,484,510]" italics="true" pageId="26" pageNumber="27">Dardanus umbella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273579EC4091F5C4BA5FE46DB40FE70" box="[497,591,483,509]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B5CFDA2D8DDFDAC" blockId="26.[264,591,483,545]" box="[264,466,519,545]" pageId="26" pageNumber="27">
(
<figureCitation id="130F2A72C4091F5C4B45FDA2D890FDAC" box="[273,415,519,545]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStart-5="FIGURE 21" captionStart-6="FIGURE 22" captionStartId-0="29.[264,368,1701,1725]" captionStartId-1="30.[264,368,1719,1743]" captionStartId-2="31.[264,368,1680,1704]" captionStartId-3="32.[264,368,1625,1649]" captionStartId-4="33.[264,368,1659,1683]" captionStartId-5="34.[264,368,1792,1816]" captionStartId-6="35.[264,368,1736,1760]" captionTargetBox-0="[344,1249,399,1689]" captionTargetBox-1="[283,1313,326,1689]" captionTargetBox-2="[283,1301,329,1639]" captionTargetBox-3="[278,1288,337,1598]" captionTargetBox-4="[289,1285,430,1609]" captionTargetBox-5="[310,1278,321,1762]" captionTargetBox-6="[302,1312,334,1707]" captionTargetId-0="figure@29.[324,1272,384,1697]" captionTargetId-1="figure@30.[268,1327,326,1716]" captionTargetId-2="figure@31.[263,1322,302,1657]" captionTargetId-3="figure@32.[264,1323,318,1633]" captionTargetId-4="figure@33.[264,1323,416,1629]" captionTargetId-5="figure@34.[280,1315,308,1783]" captionTargetId-6="figure@35.[281,1340,300,1728]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="32" captionTargetPageId-4="33" captionTargetPageId-5="34" captionTargetPageId-6="35" captionText-0="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." captionText-1="FIGURE 17. Dardanus umbella, sp. nov.: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm." captionText-2="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." captionText-3="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." captionText-4="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." captionText-5="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." captionText-6="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri-0="https://zenodo.org/record/172745/files/figure.png" httpUri-1="https://zenodo.org/record/172746/files/figure.png" httpUri-2="https://zenodo.org/record/172747/files/figure.png" httpUri-3="https://zenodo.org/record/172748/files/figure.png" httpUri-4="https://zenodo.org/record/172749/files/figure.png" httpUri-5="https://zenodo.org/record/172750/files/figure.png" httpUri-6="https://zenodo.org/record/172751/files/figure.png" pageId="26" pageNumber="27">Figs. 16–22</figureCitation>
,
<figureCitation id="130F2A72C4091F5C4BFFFDA2D8C5FDAC" box="[427,458,519,545]" captionStart="FIGURE 32" captionStartId="50.[264,368,1014,1038]" captionTargetBox="[274,1293,446,995]" captionTargetId="figure@50.[265,1324,420,1012]" captionTargetPageId="50" captionText="FIGURE 32. Geographical distribution of four species of Dardanus and one undeterminable species close to D. longior. Black circle: D. sanguinocarpus, white star: D. robustus, double circle: D. umbella, black triangle: D. longior, white triangle: Dardanus cf. longior." httpUri="https://zenodo.org/record/172760/files/figure.png" pageId="26" pageNumber="27">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E657CC4091F5C4B5CFDF2DD2FFC9C" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B8B36F7C4091F5C4B5CFDF2D8E9FDFD" blockId="26.[264,1323,599,785]" box="[264,486,599,624]" pageId="26" pageNumber="27">
<emphasis id="B940EAE5C4091F5C4B5CFDF2D8E9FDFD" box="[264,486,599,624]" italics="true" pageId="26" pageNumber="27">Material examined</emphasis>
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CFDDADD2FFC9C" blockId="26.[264,1323,599,785]" pageId="26" pageNumber="27">
<typeStatus id="548F8855C4091F5C4B6CFDDAD8ACFD15" box="[312,419,639,664]" pageId="26" pageNumber="27" type="holotype">
<emphasis id="B940EAE5C4091F5C4B6CFDDAD8ACFD15" box="[312,419,639,664]" italics="true" pageId="26" pageNumber="27">Holotype</emphasis>
</typeStatus>
: Ψ, SL
<quantity id="4CCC9B12C4091F5C4BABFDDADB54FD14" box="[511,603,639,665]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="26" pageNumber="27" unit="mm" value="5.8">5.8 mm</quantity>
, Y­48B, Yap
<collectingCountry id="F3237667C4091F5C48A3FDDADA4DFD14" box="[759,834,639,665]" name="Iceland" pageId="26" pageNumber="27">Island</collectingCountry>
,
<collectingCountry id="F3237667C4091F5C491AFDDADD1FFD14" box="[846,1040,639,665]" name="Micronesia" pageId="26" pageNumber="27">Caroline Islands</collectingCountry>
,
<collectingCountry id="F3237667C4091F5C4E48FDDADDADFD14" box="[1052,1186,639,665]" name="Micronesia" pageId="26" pageNumber="27">Micronesia</collectingCountry>
, 1953, coll. R. Hiatt, JF 1000,
<collectionCode id="ED25AE32C4091F5C4BB3FD02DB33FD4C" LSID="urn:lsid:biocol.org:col:34871" box="[487,572,679,705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
1086590.
<typeStatus id="548F8855C4091F5C48E8FD02DA28FD4D" box="[700,807,679,704]" pageId="26" pageNumber="27" type="paratype">
<emphasis id="B940EAE5C4091F5C48E8FD02DA28FD4D" box="[700,807,679,704]" italics="true" pageId="26" pageNumber="27">Paratype</emphasis>
</typeStatus>
: ♂, SL
<quantity id="4CCC9B12C4091F5C49D1FD02DAEBFD4C" box="[901,996,679,705]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="26" pageNumber="27" unit="mm" value="3.7">3.7 mm</quantity>
,
<geoCoordinate id="EE005030C4091F5C49A5FD02DD9FFD4C" box="[1009,1168,679,705]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="9" value="119.83833">119°50.30’ E</geoCoordinate>
,
<geoCoordinate id="EE005030C4091F5C4ECBFD02DC29FD4C" box="[1183,1318,679,705]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="9" value="4.9700003">4°58.20’ N</geoCoordinate>
,
<quantity id="4CCC9B12C4091F5C4B5CFD6AD840FD64" box="[264,335,719,745]" metricMagnitude="0" metricUnit="m" metricValue="2.7" pageId="26" pageNumber="27" unit="m" value="2.7">2.7 m</quantity>
, st. D5165, Tawi Tawi Group, Sulu Archipelago,
<collectingCountry id="F3237667C4091F5C49F6FD6ADD76FD64" box="[930,1145,719,745]" name="Philippines" pageId="26" pageNumber="27">Philippine Islands</collectingCountry>
,
<date id="FF8A1037C4091F5C4ED2FD6ADC29FD64" box="[1158,1318,719,745]" pageId="26" pageNumber="27" value="1908-02-24">24 Feb. 1908</date>
, Johnston oyster dredge, R/V Albatross, JF 0186c,
<collectionCode id="ED25AE32C4091F5C4901FD52DAA5FC9C" LSID="urn:lsid:biocol.org:col:34871" box="[853,938,759,785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
1086591.
</paragraph>
</subSubSection>
<subSubSection id="C32E657CC4091F624B5CFCE2D8A4FED3" lastPageId="36" lastPageNumber="37" pageId="26" pageNumber="27" type="description">
<paragraph id="8B8B36F7C4091F5C4B5CFCE2D89DFCED" blockId="26.[264,1324,839,1905]" box="[264,402,839,864]" pageId="26" pageNumber="27">
<emphasis id="B940EAE5C4091F5C4B5CFCE2D89DFCED" box="[264,402,839,864]" italics="true" pageId="26" pageNumber="27">Description</emphasis>
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CFCCADA3EFBDC" blockId="26.[264,1324,839,1905]" pageId="26" pageNumber="27">
Shield (
<figureCitation id="130F2A72C4091F5C4BC1FCCAD8E3FC04" box="[405,492,879,905]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
A) as long as broad; anterior margin between rostral lobe and lateral projections nearly straight; lateral margins slightly convex, somewhat irregular, with rows of long setae. Dorsal surface of shield with short rows or tufts of setae; weakly­calcified Yshaped linea present posteriorly; gastric pits distinct. Rostral lobe indistinct. Lateral projections large, bluntly triangular, produced. Posterior carapace lateral elements well calcified, unarmed. Branchiostegites unarmed.
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CFBFAD89BFB7C" blockId="26.[264,1324,839,1905]" pageId="26" pageNumber="27">
Ocular peduncles (
<figureCitation id="130F2A72C4091F5C4842FBFADB64FBF4" box="[534,619,1119,1145]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
A) 0.8–0.9 length of shield, subcylindrical, slightly inflated distally; corneas only very slightly dilated. Ocular acicles (
<figureCitation id="130F2A72C4091F5C49EFFB22DD1FFB2C" box="[955,1040,1159,1185]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
B) broad; distal margins each with 2–5 spines, tips often corneous. Interocular plate (
<figureCitation id="130F2A72C4091F5C498EFB0ADD3FFB44" box="[986,1072,1199,1225]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
B) with pair of small protrusions.
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CFB5ADDFCFAE4" blockId="26.[264,1324,839,1905]" pageId="26" pageNumber="27">
Antennular peduncles (
<figureCitation id="130F2A72C4091F5C4802FB5ADBBFFA94" box="[598,688,1279,1305]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
A) slender, when fully extended, distal margins of ultimate segments reaching bases to distal margins of corneas; ultimate and penultimate segments unarmed; basal segment with ventromesial distal angle bearing small spine.
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CFAD2D858F90C" blockId="26.[264,1324,839,1905]" pageId="26" pageNumber="27">
Antennal peduncles (
<figureCitation id="130F2A72C4091F5C4862FAD2DB80FA1C" box="[566,655,1399,1425]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
A, B), when fully extended, reaching bases of corneas; fifth and fourth segments unarmed; third segment projecting ventrodistally; second segment with dorsomesial distal angle bearing spine, dorsolateral distal angle produced, terminating in simple or bifid spine; first segment with ventrolateral distal angle with spine. Antennal acicle (
<figureCitation id="130F2A72C4091F5C4873F9B2DB70F9BC" box="[551,639,1559,1585]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
B) terminating in strong spine; dorsomesial margin with 2–4 spines; dorsolateral margin with 0–2 subdistal spines. Antennal flagella sparsely setose.
</paragraph>
<paragraph id="8B8B36F7C4091F5C4B6CF92ADBDAF95C" blockId="26.[264,1324,839,1905]" pageId="26" pageNumber="27">
Third maxilliped (
<figureCitation id="130F2A72C4091F5C4847F92ADB64F924" box="[531,619,1679,1705]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
C, D) with well­developed crista dentata (
<figureCitation id="130F2A72C4091F5C4E3CF92ADDCFF924" box="[1128,1216,1679,1705]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
D); basis with 1 or 2 corneous spines (
<figureCitation id="130F2A72C4091F5C480EF912DBBFF95C" box="[602,688,1719,1745]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
D).
</paragraph>
<paragraph id="8B8B36F7C4091F5D4B6CF97ADD2DFC03" blockId="26.[264,1324,839,1905]" lastBlockId="27.[264,1324,284,1870]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Chelipeds vastly unequal, left larger. Left cheliped very stout, more elongate in
<typeStatus id="548F8855C4091F5C4B5CF8A2D862F8AC" box="[264,365,1799,1825]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
male (
<figureCitation id="130F2A72C4091F5C4BE9F8A2DB1DF8AC" box="[445,530,1799,1825]" captionStart="FIGURE 17" captionStartId="30.[264,368,1719,1743]" captionTargetBox="[283,1313,326,1689]" captionTargetId="figure@30.[268,1327,326,1716]" captionTargetPageId="30" captionText="FIGURE 17. Dardanus umbella, sp. nov.: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm." httpUri="https://zenodo.org/record/172746/files/figure.png" pageId="26" pageNumber="27">Fig. 17</figureCitation>
A, B) than in
<typeStatus id="548F8855C4091F5C48E7F8A2DA16F8AC" box="[691,793,1799,1825]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
female (
<figureCitation id="130F2A72C4091F5C49D5F8A2DAD7F8AC" box="[897,984,1799,1825]" captionStart="FIGURE 18" captionStartId="31.[264,368,1680,1704]" captionTargetBox="[283,1301,329,1639]" captionTargetId="figure@31.[263,1322,302,1657]" captionTargetPageId="31" captionText="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="26" pageNumber="27">Fig. 18</figureCitation>
). Dactyl terminating in large corneous claw; cutting edge with 6–10 variously­sized calcareous teeth; upper and outer faces with unarmed to armed with several corneous, or sometimes corneous­tipped, conical spines and short stiff setae. Fixed finger terminating in large corneous claw; cutting edge with 5–8 variously sized calcareous teeth. Palm and fixed finger with outer face bearing numerous prominent corneous­tipped, conical spines, sometimes forming longitudinal rows, and stiff setae of short to medium length; lower half of outer face concave, numerous miniscule setae reminiscent of umbrella­like ribs (
<figureCitation id="130F2A72C4081F5D4E6EFE19DD82FE5B" box="[1082,1165,444,470]" captionStart="FIGURE 18" captionStartId="31.[264,368,1680,1704]" captionTargetBox="[283,1301,329,1639]" captionTargetId="figure@31.[263,1322,302,1657]" captionTargetPageId="31" captionText="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="27" pageNumber="28">Fig. 18</figureCitation>
C) present on lower half of outer face; lower face with 1–3 irregular rows of prominent tricolored conical spines each consisting of distal corneous, middle half corneous, and proximal calcareous portions (
<figureCitation id="130F2A72C4081F5D485CFD91DB6BFDC3" box="[520,612,564,590]" captionStart="FIGURE 18" captionStartId="31.[264,368,1680,1704]" captionTargetBox="[283,1301,329,1639]" captionTargetId="figure@31.[263,1322,302,1657]" captionTargetPageId="31" captionText="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="27" pageNumber="28">Fig. 18</figureCitation>
B), accompanied with 1–3 irregular rows of corneous or corneous­tipped spines; upper face of palm with 1 or 2 irregular rows of prominent calcareous or corneous­tipped, conical spines. Carpus with upper margin bearing 1 or 2 irregular rows of prominent calcareous or corneous­tipped conical spines; outer face with numerous tubercles (
<typeStatus id="548F8855C4081F5D4859FD71DB7BFD63" box="[525,628,724,750]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
male,
<figureCitation id="130F2A72C4081F5D4881FD71DA3CFD63" box="[725,819,724,750]" captionStart="FIGURE 17" captionStartId="30.[264,368,1719,1743]" captionTargetBox="[283,1313,326,1689]" captionTargetId="figure@30.[268,1327,326,1716]" captionTargetPageId="30" captionText="FIGURE 17. Dardanus umbella, sp. nov.: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm." httpUri="https://zenodo.org/record/172746/files/figure.png" pageId="27" pageNumber="28">Fig. 17</figureCitation>
), or numerous prominent calcareous or corneous­tipped spines (
<typeStatus id="548F8855C4081F5D4862FD59DB91FC9B" box="[566,670,764,790]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
female,
<figureCitation id="130F2A72C4081F5D4942FD59DA7AFC9B" box="[790,885,764,790]" captionStart="FIGURE 18" captionStartId="31.[264,368,1680,1704]" captionTargetBox="[283,1301,329,1639]" captionTargetId="figure@31.[263,1322,302,1657]" captionTargetPageId="31" captionText="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="27" pageNumber="28">Fig. 18</figureCitation>
). Merus with ventrolateral margin bearing few large spines (
<typeStatus id="548F8855C4081F5D4817FC81DBA5FCB3" box="[579,682,804,830]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
male,
<figureCitation id="130F2A72C4081F5D4954FC81DA55FCB3" box="[768,858,804,830]" captionStart="FIGURE 17" captionStartId="30.[264,368,1719,1743]" captionTargetBox="[283,1313,326,1689]" captionTargetId="figure@30.[268,1327,326,1716]" captionTargetPageId="30" captionText="FIGURE 17. Dardanus umbella, sp. nov.: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm." httpUri="https://zenodo.org/record/172746/files/figure.png" pageId="27" pageNumber="28">Fig. 17</figureCitation>
), or ventral margin and ventral 0.3 of lateral face bearing numerous corneous or corneous­tipped spines (
<typeStatus id="548F8855C4081F5D4E7DFCE9DD9DFCEB" box="[1065,1170,844,870]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
female:
<figureCitation id="130F2A72C4081F5D4EA8FCE9D827FC03" captionStart="FIGURE 18" captionStartId="31.[264,368,1680,1704]" captionTargetBox="[283,1301,329,1639]" captionTargetId="figure@31.[263,1322,302,1657]" captionTargetPageId="31" captionText="FIGURE 18. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm." httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="27" pageNumber="28">Fig. 18</figureCitation>
); distal margin of lateral face with 0–3 spines. Ischium unarmed.
</paragraph>
<paragraph id="8B8B36F7C4081F5D4B6CFC39DB9AFA33" blockId="27.[264,1324,284,1870]" pageId="27" pageNumber="28">
Right cheliped moderately slender (
<figureCitation id="130F2A72C4081F5D488AFC39DA3DFC3B" box="[734,818,924,950]" captionStart="FIGURE 17" captionStartId="30.[264,368,1719,1743]" captionTargetBox="[283,1313,326,1689]" captionTargetId="figure@30.[268,1327,326,1716]" captionTargetPageId="30" captionText="FIGURE 17. Dardanus umbella, sp. nov.: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm." httpUri="https://zenodo.org/record/172746/files/figure.png" pageId="27" pageNumber="28">Fig. 17</figureCitation>
C–E), generally setose. Dactyl terminating in large corneous claw; cutting edge with 5–8 variously­sized calcareous teeth; upper face with 1–3 irregular rows of prominent corneous, or sometimes corneous­tipped, spines; outer face with few corneous spines. Fixed finger terminating in large corneous claw; cutting edge with 5–8 variously­sized calcareous teeth. Palm and fixed finger with outer faces bearing numerous corneous, or sometimes corneous­tipped, spines, often forming longitudinal rows; lower face with 1 or 2 irregular rows of prominent conical spines accompanied with 1–3 irregular rows of corneous, or corneous­tipped spines; upper margin of palm with 1 or 2 irregular rows of very strong corneous­tipped conical spines. Carpus with upper margin bearing 1 or 2 irregular rows of very strong corneous­tipped conical spines; outer face with several, prominent corneous­tipped spines on upper half and many small corneous­tipped spines near distal margin. Merus with distal half of lateral face bearing many corneous or corneous­tipped spines, sometimes forming short transverse row. Ischium unarmed.
</paragraph>
<paragraph id="8B8B36F7C4081F5A4B6CFA69DBC5FDFB" blockId="27.[264,1324,284,1870]" lastBlockId="28.[264,1325,284,1870]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
Second (
<figureCitation id="130F2A72C4081F5D4BF2FA69DB0CFA6B" box="[422,515,1484,1510]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
) and right third (
<figureCitation id="130F2A72C4081F5D48B9FA69DA45FA6B" box="[749,842,1484,1510]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
A–C) pereopods generally similar, but armament and proportion somewhat different between second pair and right third; second pair more slender than right third; of second pair, left slightly shorter than right. Dactyls of second pair and right third all 1.2–1.3 length of propodi, each terminating in strong corneous claw; dorsal surfaces with irregular rows of tufts of stiff setae, and, on second, setae sometimes forming short obliquely transverse rows (
<figureCitation id="130F2A72C4081F5D49E6F931DD07F923" box="[946,1032,1684,1710]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
C); few strong spine­like setae present dorsodistally (
<figureCitation id="130F2A72C4081F5D480FF919DBCFF95B" box="[603,704,1724,1750]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="27" pageNumber="28">Figs. 19</figureCitation>
D, 22B) or laterally (
<figureCitation id="130F2A72C4081F5D499CF919DD2AF95B" box="[968,1061,1724,1750]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
B); lateral and mesial faces generally convex (second,
<figureCitation id="130F2A72C4081F5D48DCF941DBD2F973" box="[648,733,1764,1790]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
B, C) or each with longitudinal faint sulcus (third,
<figureCitation id="130F2A72C4081F5D4B5CF8A9D850F8AB" box="[264,351,1804,1830]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
B, C), with few longitudinal rows of tufts of stiff setae; ventral margins each with row of 3–6 corneous spines. Propodi 1.7–1.8 (second,
<figureCitation id="130F2A72C4081F5D49CAF891DAFBF8C3" box="[926,1012,1844,1870]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
E) or 2.2–2.3 (right third,
<figureCitation id="130F2A72C40F1F5A4B5CFEB9D852FEBB" box="[264,349,284,310]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
A) length of carpi; dorsal faces flat and broad (second) or comparatively narrower (right third), armed with numerous (second) or comparatively fewer numbers of (right third) corneous spines and with long stiff setae, dorsodistal margins armed with corneous spines; lateral faces with few longitudinal rows of tufts of stiff setae. Carpi 0.5–0.6 (second,
<figureCitation id="130F2A72C40F1F5A4B25FE19D8CDFE5B" box="[369,450,444,470]" captionStart="FIGURE 19" captionStartId="32.[264,368,1625,1649]" captionTargetBox="[278,1288,337,1598]" captionTargetId="figure@32.[264,1323,318,1633]" captionTargetPageId="32" captionText="FIGURE 19. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm." httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="28" pageNumber="29">Fig. 19</figureCitation>
A) or 0.6–0.7 (right third,
<figureCitation id="130F2A72C40F1F5A48A0FE19DA47FE5B" box="[756,840,444,470]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
A) length of meri; dorsolateral and dorsal margins with several corneous­tipped spines distally and large dorsodistal spine (second), or with only large dorsodistal spine (right third). Meri with ventrolateral distal angles each with sharp spine and ventral margins each with row of several spines (second), or meri unarmed (right third). Ischia unarmed.
</paragraph>
<paragraph id="8B8B36F7C40F1F5A4B6CFD21DB06F983" blockId="28.[264,1325,284,1870]" pageId="28" pageNumber="29">
Left third pereopod (
<figureCitation id="130F2A72C40F1F5A4879FD21DB9FFD13" box="[557,656,644,670]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Figs. 20</figureCitation>
,
<figureCitation id="130F2A72C40F1F5A48C9FD21DBB4FD13" box="[669,699,644,670]" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">21</figureCitation>
) generally very stout. Dactyl (
<figureCitation id="130F2A72C40F1F5A4E70FD21DD89FD13" box="[1060,1158,644,670]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Figs. 20</figureCitation>
A, B, 21A–C) 1.4–1.5 length of propodus, very broad, terminating in strong corneous claw; lateral surface (
<figureCitation id="130F2A72C40F1F5A4B26FD71D8C1FD63" box="[370,462,724,750]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
B) with dorsal row of corneous spines (
<figureCitation id="130F2A72C40F1F5A4990FD71DD2FFD63" box="[964,1056,724,750]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
B­a), dorsal half with longitudinal concavity (
<figureCitation id="130F2A72C40F1F5A487EFD59DB89FC9B" box="[554,646,764,790]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
B­b), ventral half (
<figureCitation id="130F2A72C40F1F5A4924FD59DAC5FC9B" box="[880,970,764,790]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
B­c) with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen in
<figureCitation id="130F2A72C40F1F5A4B5CFCE9D854FCEB" box="[264,347,844,870]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
B due to their tiny size); ventral margin somewhat irregular proximally, with row of small corneous spines (
<figureCitation id="130F2A72C40F1F5A4843FCD1DB64FC03" box="[535,619,884,910]" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
C) and rows of stiff setae (
<figureCitation id="130F2A72C40F1F5A49CAFCD1DAF0FC03" box="[926,1023,884,910]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Figs. 20</figureCitation>
B, 21B); mesial face (
<figureCitation id="130F2A72C40F1F5A4EAFFCD1D829FC3B" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
B) strongly convex, with few irregular rows of tufts of short stiff setae; dorsal margin (
<figureCitation id="130F2A72C40F1F5A4B45FC61D869FC53" box="[273,358,964,990]" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
B) with rows of dense stiff setae. Propodus (
<figureCitation id="130F2A72C40F1F5A4921FC61DAD7FC53" box="[885,984,964,990]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Figs. 20</figureCitation>
C, D, 21D, E) 1.5–1.6 length of carpus, very broad; dorsal margin with series of furrows producing row of lobes, lateral face of each lobe sometimes armed dorsodistally with slender corneous spines or spinelike setae (
<figureCitation id="130F2A72C40F1F5A4BD2FB99D8D3FBDB" box="[390,476,1084,1110]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
C­a), dorsomesial surface of each lobe with 1–3 strong spines and dense setae (
<figureCitation id="130F2A72C40F1F5A4B01FBC1D8A1FBF3" box="[341,430,1124,1150]" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
D); lateral face (
<figureCitation id="130F2A72C40F1F5A4821FBC1DBC2FBF3" box="[629,717,1124,1150]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
C) with dorsal half bearing very deep longitudinal concavity (
<figureCitation id="130F2A72C40F1F5A4BD9FB29D8EBFB2B" box="[397,484,1164,1190]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
C­b) with scattered stiff setae, ventral half with 2–4 irregular rows of large protuberances (
<figureCitation id="130F2A72C40F1F5A4850FB11DB54FB43" box="[516,603,1204,1230]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
C­c) each bearing 1 to several stiff setae or spiniform setae and with miniscule setae reminiscent of umbrella­like ribs (
<figureCitation id="130F2A72C40F1F5A4995FB79DD17FB7B" box="[961,1048,1244,1270]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
D); ventral margin with series of deep furrows producing row of large lobes, lateral face of each lobe with marginal setae or spine­like setae (
<figureCitation id="130F2A72C40F1F5A48E4FA89DA05FACB" box="[688,778,1324,1350]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
C­d), ventromesial margin of each lobe (
<figureCitation id="130F2A72C40F1F5A4EAFFA89D82AFAE3" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
E) with 0–2 very large corneous spines and row of setae; mesial face (
<figureCitation id="130F2A72C40F1F5A4EDBFAF1DDE7FAE3" box="[1167,1256,1364,1390]" captionStart="FIGURE 21" captionStartId="34.[264,368,1792,1816]" captionTargetBox="[310,1278,321,1762]" captionTargetId="figure@34.[280,1315,308,1783]" captionTargetPageId="34" captionText="FIGURE 21. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm." httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
A, E) strongly convex, with few rows of tufts of short setae, distal margin with 1–6 corneous spines. Carpus 0.7–0.8 length of merus; lateral face (
<figureCitation id="130F2A72C40F1F5A49E8FA01DD15FA33" box="[956,1050,1444,1470]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
A) with several large corneous­tipped spines. Merus and ischium (
<figureCitation id="130F2A72C40F1F5A4940FA69DA67FA6B" box="[788,872,1484,1510]" captionStart="FIGURE 20" captionStartId="33.[264,368,1659,1683]" captionTargetBox="[289,1285,430,1609]" captionTargetId="figure@33.[264,1323,416,1629]" captionTargetPageId="33" captionText="FIGURE 20. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot; c &quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm." httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
A) with armament and setation similar to those of right third.
</paragraph>
<paragraph id="8B8B36F7C40F1F5A4B6CF9B9DA31F9D3" blockId="28.[264,1325,284,1870]" pageId="28" pageNumber="29">
Sternite of third pereopods with anterior lobe (
<figureCitation id="130F2A72C40F1F5A49DFF9B9DAE9F9BB" box="[907,998,1564,1590]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
E) rectangular, with small protrusion bearing short transverse row of setae.
</paragraph>
<paragraph id="8B8B36F7C40F1F5A4B6CF9C9D8D9F95B" blockId="28.[264,1325,284,1870]" pageId="28" pageNumber="29">
Fourth pereopod (
<figureCitation id="130F2A72C40F1F5A485AF9C9DB6BF90B" box="[526,612,1644,1670]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
F) semichelate; dactyl with 5 or 6 short corneous spines on lateral face ventrally; propodal rasp well developed; carpus with sharp corneous­tipped dorsodistal spine.
</paragraph>
<paragraph id="8B8B36F7C40F1F5A4B6CF941DD6DF973" blockId="28.[264,1325,284,1870]" box="[312,1122,1764,1790]" pageId="28" pageNumber="29">Fifth pereopod chelate; rasps of dactyl and propodus well developed.</paragraph>
<paragraph id="8B8B36F7C40F1F5B4B6CF8A9DB8EFED3" blockId="28.[264,1325,284,1870]" lastBlockId="29.[264,1323,284,350]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Female pleon (
<figureCitation id="130F2A72C40F1F5A4BB3F8A9DB33F8AB" box="[487,572,1804,1830]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
D) with second to fifth left pleopods, fringed with setae; second to fourth large, triramous; fifth small, with slender exopod; extremely large fleshy membranous protuberance with scattered setae present between fourth and fifth pleopods. Pleon of
<typeStatus id="548F8855C40E1F5B4B26FEE1D8D8FED3" box="[370,471,324,350]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
male missing.
</paragraph>
<caption id="DF4B667FC40E1F5B4B5CF900DB9FF8E7" httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="29" pageNumber="30" targetBox="[344,1249,399,1689]" targetPageId="29">
<paragraph id="8B8B36F7C40E1F5B4B5CF900DB9FF8E7" blockId="29.[264,1323,1701,1898]" pageId="29" pageNumber="30">
<emphasis id="B940EAE5C40E1F5B4B5CF900D893F930" bold="true" box="[264,412,1701,1725]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
<taxonomicName id="4C344D74C40E1F5B4BFCF903DB7CF930" box="[424,627,1702,1725]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C40E1F5B4BFCF903DB7CF930" box="[424,627,1702,1725]" italics="true" pageId="29" pageNumber="30">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC40E1F5B48D1F900DBD0F930" box="[645,735,1701,1725]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B940EAE5C40E1F5B48D1F900DBD0F930" bold="true" box="[645,735,1701,1725]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF4B667FC40D1F584B5CF912DDBAF8B3" httpUri="https://zenodo.org/record/172746/files/figure.png" pageId="30" pageNumber="31" targetBox="[283,1313,326,1689]" targetPageId="30">
<paragraph id="8B8B36F7C40D1F584B5CF912DDBAF8B3" blockId="30.[264,1324,1719,1854]" pageId="30" pageNumber="31">
<emphasis id="B940EAE5C40D1F584B5CF912D896F942" bold="true" box="[264,409,1719,1743]" pageId="30" pageNumber="31">FIGURE 17.</emphasis>
<taxonomicName id="4C344D74C40D1F584BF6F912DB65F943" box="[418,618,1719,1742]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C40D1F584BF6F912DB65F943" box="[418,618,1719,1742]" italics="true" pageId="30" pageNumber="31">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC40D1F58482DF912DBDFF942" box="[633,720,1719,1743]" pageId="30" pageNumber="31" rank="species">
<emphasis id="B940EAE5C40D1F58482DF912DBDFF942" bold="true" box="[633,720,1719,1743]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
: paratype, ɗ, SL 2.5 mm, Philippine, USNM 1086591. Left cheliped: A, chela and carpus, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, merus, outer. Right cheliped: C, chela and carpus, outer; D, merus and ischium, outer; E, chela and carpus, lower. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF4B667FC40C1F594B5CF935DBF6F8BE" httpUri="https://zenodo.org/record/172747/files/figure.png" pageId="31" pageNumber="32" targetBox="[283,1301,329,1639]" targetPageId="31">
<paragraph id="8B8B36F7C40C1F594B5CF935DBF6F8BE" blockId="31.[264,1323,1680,1843]" pageId="31" pageNumber="32">
<emphasis id="B940EAE5C40C1F594B5CF935D893F925" bold="true" box="[264,412,1680,1704]" pageId="31" pageNumber="32">FIGURE 18.</emphasis>
<taxonomicName id="4C344D74C40C1F594BFCF934DB7CF925" box="[424,627,1681,1704]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C40C1F594BFCF934DB7CF925" box="[424,627,1681,1704]" italics="true" pageId="31" pageNumber="32">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC40C1F5948D1F935DBD0F925" box="[645,735,1680,1704]" pageId="31" pageNumber="32" rank="species">
<emphasis id="B940EAE5C40C1F5948D1F935DBD0F925" bold="true" box="[645,735,1680,1704]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left cheliped: A, whole, outer, area encircled by dashed line indicates area where miniscule setae reminiscent of umbrella­like ribs occur; B, chela and carpus, inner lower; C, lower portion of outer face of palm, enlarged, indicating setae reminiscent of umbrella­like ribs, normal setae and corneous­tipped spines. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF4B667FC4331F664B5CF9FCDBB6F93B" httpUri="https://zenodo.org/record/172748/files/figure.png" pageId="32" pageNumber="33" targetBox="[278,1288,337,1598]" targetPageId="32">
<paragraph id="8B8B36F7C4331F664B5CF9FCDBB6F93B" blockId="32.[264,1323,1625,1718]" pageId="32" pageNumber="33">
<emphasis id="B940EAE5C4331F664B5CF9FCD893F9FC" bold="true" box="[264,412,1625,1649]" pageId="32" pageNumber="33">FIGURE 19.</emphasis>
<taxonomicName id="4C344D74C4331F664BFCF9FFDB7CF9FC" box="[424,627,1626,1649]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C4331F664BFCF9FFDB7CF9FC" box="[424,627,1626,1649]" italics="true" pageId="32" pageNumber="33">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC4331F6648D1F9FCDBD0F9FC" box="[645,735,1625,1649]" pageId="32" pageNumber="33" rank="species">
<emphasis id="B940EAE5C4331F6648D1F9FCDBD0F9FC" bold="true" box="[645,735,1625,1649]" pageId="32" pageNumber="33">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Left second pereopod: A, whole, lateral; B, dactyl, lateral; C, dactyl, mesial; D, dactyl, dorsal; E, propodus, dorsal. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8B8B36F7C4331F664B6CF941DBA3F8AB" blockId="32.[264,1322,1764,1870]" pageId="32" pageNumber="33">
Uropods (
<figureCitation id="130F2A72C4331F664BE7F941DB00F973" box="[435,527,1764,1790]" captionStart="FIGURE 22" captionStartId="35.[264,368,1736,1760]" captionTargetBox="[302,1312,334,1707]" captionTargetId="figure@35.[281,1340,300,1728]" captionTargetPageId="35" captionText="FIGURE 22. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm." httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="32" pageNumber="33">Fig. 22</figureCitation>
D) markedly asymmetrical, left larger than right; endopods and exopods with well­developed rasps.
</paragraph>
<paragraph id="8B8B36F7C4331F674B6CF891DDC5FE0B" blockId="32.[264,1322,1764,1870]" lastBlockId="33.[264,1324,284,390]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Telson (
<figureCitation id="130F2A72C4331F664BC9F891D8F8F8C3" box="[413,503,1844,1870]" captionStart="FIGURE 16" captionStartId="29.[264,368,1701,1725]" captionTargetBox="[344,1249,399,1689]" captionTargetId="figure@29.[324,1272,384,1697]" captionTargetPageId="29" captionText="FIGURE 16. Dardanus umbella, sp. nov.: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. A, shield and cephalic appendages, dorsal. B, interocular plate, right ocular acicle, and second and first (partial) segments and acicle of right antennal peduncle, dorsal, setae omitted. Right third maxilliped: C, inner; D, ischium and basis, outer. E, anterior lobe of sternite of third pereopods. F, chela and carpus of left fourth pereopod, lateral, setae omitted. Telson: G, dorsal; H, posterior lobes, ventral. Scale 1 mm." httpUri="https://zenodo.org/record/172745/files/figure.png" pageId="32" pageNumber="33">Fig. 16</figureCitation>
G, H) with lateral constrictions; marginal area partially calcified; posterior lobes separated by median cleft, left larger than right, with 5–7 (right) or 9–13 (left) ventral corneous spines near terminal margin, spines on left sometimes very long, terminal margins fringed with many short bristle­like setae and several long setae.
</paragraph>
<caption id="DF4B667FC4321F674B5CF9DEDDEFF8EE" httpUri="https://zenodo.org/record/172749/files/figure.png" pageId="33" pageNumber="34" targetBox="[289,1285,430,1609]" targetPageId="33">
<paragraph id="8B8B36F7C4321F674B5CF9DEDDEFF8EE" blockId="33.[264,1325,1659,1891]" pageId="33" pageNumber="34">
<emphasis id="B940EAE5C4321F674B5CF9DED893F91E" bold="true" box="[264,412,1659,1683]" pageId="33" pageNumber="34">FIGURE 20.</emphasis>
<taxonomicName id="4C344D74C4321F674BFCF9D9DB7CF91E" box="[424,627,1660,1683]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C4321F674BFCF9D9DB7CF91E" box="[424,627,1660,1683]" italics="true" pageId="33" pageNumber="34">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC4321F6748D1F9DEDBD0F91E" box="[645,735,1659,1683]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B940EAE5C4321F6748D1F9DEDBD0F91E" bold="true" box="[645,735,1659,1683]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Lateral face of left third pereopod: A, whole; B, dactyl, a, dorsal row of corneous spines, b, dorsal longitudinal concavity, c, ventral area with scattered corneous spines or spiniform setae and miniscule setae reminiscent of umbrella­like ribs (not seen due to their tiny size, see D for enlarged figure); C, propodus, a, dorsal row of large lobes, b, dorsal deep longitudinal concavity, c, ventral area with irregular rows of large protuberances, d, ventral row of lobes; D, enlarged figure of area &quot;c&quot;, indicating setae reminiscent of umbrella­like ribs and normal setae. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF4B667FC4311F644B5CF8A5DD5FF8D1" httpUri="https://zenodo.org/record/172750/files/figure.png" pageId="34" pageNumber="35" targetBox="[310,1278,321,1762]" targetPageId="34">
<paragraph id="8B8B36F7C4311F644B5CF8A5DD5FF8D1" blockId="34.[264,1322,1792,1884]" pageId="34" pageNumber="35">
<emphasis id="B940EAE5C4311F644B5CF8A5D893F895" bold="true" box="[264,412,1792,1816]" pageId="34" pageNumber="35">FIGURE 21.</emphasis>
<taxonomicName id="4C344D74C4311F644BFCF8A4DB7CF895" box="[424,627,1793,1816]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C4311F644BFCF8A4DB7CF895" box="[424,627,1793,1816]" italics="true" pageId="34" pageNumber="35">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC4311F6448D1F8A5DBD0F895" box="[645,735,1792,1816]" pageId="34" pageNumber="35" rank="species">
<emphasis id="B940EAE5C4311F6448D1F8A5DBD0F895" bold="true" box="[645,735,1792,1816]" pageId="34" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Mesial face of left third pereopod: A, dactyl and propodus; B, dactyl; C, ventral portion of dactyl, setae omitted; D, dorsal portion of propodus; E, propodus. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF4B667FC4301F654B5CF96DDB4DF8A8" httpUri="https://zenodo.org/record/172751/files/figure.png" pageId="35" pageNumber="36" targetBox="[302,1312,334,1707]" targetPageId="35">
<paragraph id="8B8B36F7C4301F654B5CF96DDB4DF8A8" blockId="35.[264,1322,1736,1829]" pageId="35" pageNumber="36">
<emphasis id="B940EAE5C4301F654B5CF96DD893F96D" bold="true" box="[264,412,1736,1760]" pageId="35" pageNumber="36">FIGURE 22.</emphasis>
<taxonomicName id="4C344D74C4301F654BFCF96CDB7CF96D" box="[424,627,1737,1760]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="B940EAE5C4301F654BFCF96CDB7CF96D" box="[424,627,1737,1760]" italics="true" pageId="35" pageNumber="36">Dardanus umbella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A273579EC4301F6548D1F96DDBD0F96D" box="[645,735,1736,1760]" pageId="35" pageNumber="36" rank="species">
<emphasis id="B940EAE5C4301F6548D1F96DDBD0F96D" bold="true" box="[645,735,1736,1760]" pageId="35" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, Ψ, SL 3.9 mm, Yap, Micronesia, USNM 1086590. Right third pereopod: A, whole, lateral; B, dactyl, mesial; C, dactyl, lateral. D, left side of pleon of female. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8B8B36F7C4371F624B5CFEB9D897FEB8" blockId="36.[264,427,284,350]" box="[264,408,284,309]" pageId="36" pageNumber="37">
<emphasis id="B940EAE5C4371F624B5CFEB9D897FEB8" box="[264,408,284,309]" italics="true" pageId="36" pageNumber="37">Color in life</emphasis>
</paragraph>
<paragraph id="8B8B36F7C4371F624B6CFEE1D8A4FED3" blockId="36.[264,427,284,350]" box="[312,427,324,350]" pageId="36" pageNumber="37">Unknown</paragraph>
</subSubSection>
<subSubSection id="C32E657CC4371F624B5CFE31DAAFFDAB" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8B8B36F7C4371F624B5CFE31D861FE20" blockId="36.[264,1323,404,550]" box="[264,366,404,429]" pageId="36" pageNumber="37">
<emphasis id="B940EAE5C4371F624B5CFE31D861FE20" box="[264,366,404,429]" italics="true" pageId="36" pageNumber="37">Remarks</emphasis>
</paragraph>
<paragraph id="8B8B36F7C4371F624B6CFE19DAAFFDAB" blockId="36.[264,1323,404,550]" pageId="36" pageNumber="37">
The specimens of this species in USNM were initially identified as
<taxonomicName id="4C344D74C4371F624ED8FE19D8CCFE70" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sanguinocarpus">
<emphasis id="B940EAE5C4371F624ED8FE19DC0FFE58" box="[1164,1280,444,469]" italics="true" pageId="36" pageNumber="37">Dardanus</emphasis>
cf.
<emphasis id="B940EAE5C4371F624B5CFE41D8CCFE70" box="[264,451,484,509]" italics="true" pageId="36" pageNumber="37">sanguinocarpus</emphasis>
</taxonomicName>
by J. Forest, suggesting that he had already recognized morphological differences between
<taxonomicName id="4C344D74C4371F624BAEFDA9DBD5FDA8" box="[506,730,524,549]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sanguinocarpus">
<emphasis id="B940EAE5C4371F624BAEFDA9DBD5FDA8" box="[506,730,524,549]" italics="true" pageId="36" pageNumber="37">D. sanguinocarpus</emphasis>
</taxonomicName>
and this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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