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<mods:title id="4986A44EBAB712B0408718148918FFF7">A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
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<taxonomicName id="4C344D55186FFFD9FC7CFEB9FB6149F6" ID-CoL="63LD4" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[1002,1261,381,405]" class="Arachnida" family="Lycosidae" genus="Passiena" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAC4186FFFD9FC7CFEB9FB6149F6" bold="true" box="[1002,1261,381,405]" pageId="1" pageNumber="480">
<emphasis id="B940EAC4186FFFD9FC7CFEB9FBC049F6" bold="true" box="[1002,1100,381,405]" italics="true" pageId="1" pageNumber="480">Passiena</emphasis>
<bibRefCitation id="EFA54B27186FFFD9FBC3FEB9FB6149F6" author="Thorell T" box="[1109,1261,381,405]" pageId="1" pageNumber="480" pagination="132 - 172" refId="ref5071" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
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<typeStatus id="548F8874186FFFD9FCBBFE79FCE549B6" box="[813,873,445,469]" pageId="1" pageNumber="480">Type</typeStatus>
species.
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<taxonomicName id="4C344D55186FFFD9FC4AFE7AFAE949B6" ID-CoL="4DY64" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[988,1381,445,469]" class="Arachnida" family="Lycosidae" genus="Passiena" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B940EAC4186FFFD9FC4AFE7AFB3B49B6" box="[988,1207,446,469]" italics="true" pageId="1" pageNumber="480">Passiena spinicrus</emphasis>
<bibRefCitation id="EFA54B27186FFFD9FB55FE79FAE949B6" author="Thorell T" box="[1219,1381,445,469]" pageId="1" pageNumber="480" pagination="132 - 172" refId="ref5071" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
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, from
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.
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<paragraph id="8B8B36D6186FFFD9FCBBFDD9FAD14BF6" blockId="1.[811,1452,541,917]" pageId="1" pageNumber="480">
<emphasis id="B940EAC4186FFFD9FCBBFDD9FC224A56" bold="true" box="[813,942,541,565]" pageId="1" pageNumber="480">Comments.</emphasis>
The genus is currently known from four species. Two of them, including the type species
<taxonomicName id="4C344D55186FFFD9FB44FDFAFCEF4A16" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" class="Arachnida" family="Lycosidae" genus="Passiena" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B940EAC4186FFFD9FB44FDFAFAC34A36" box="[1234,1359,574,597]" italics="true" pageId="1" pageNumber="480">P. spinicrus</emphasis>
<bibRefCitation id="EFA54B27186FFFD9FAC1FDF9FCEF4A16" author="Thorell T" pageId="1" pageNumber="480" pagination="132 - 172" refId="ref5071" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
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, occur in Southeast Asia, and two others (seemingly misplaced in the genus) are restricted to Africa (WSC, 2020). The type species is known by the
<typeStatus id="548F8874186FFFD9FB31FD59FA884AD6" box="[1191,1284,669,693]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
female. It was redescribed and illustrated for the first time by
<bibRefCitation id="EFA54B27186FFFD9FADCFD79FCFB4A96" author="Lehtinen PT" pageId="1" pageNumber="480" pagination="398 - 407" refId="ref4766" refString="Lehtinen PT (2005) Review of the Oriental wolf spider genus Passiena (Lycosidae, Pardosinae). Journal of Arachnology, 33: 398 - 407." type="journal article" year="2005">Lehtinen (2005)</bibRefCitation>
. In the same work,
<bibRefCitation id="EFA54B27186FFFD9FBDCFD19FB744A96" author="Lehtinen PT" box="[1098,1272,733,757]" pageId="1" pageNumber="480" pagination="398 - 407" refId="ref4766" refString="Lehtinen PT (2005) Review of the Oriental wolf spider genus Passiena (Lycosidae, Pardosinae). Journal of Arachnology, 33: 398 - 407." type="journal article" year="2005">Lehtinen (2005)</bibRefCitation>
described a new species,
<taxonomicName id="4C344D55186FFFD9FC1CFD3AFB4B4B76" authority="Lehtinen, 2005" authorityName="Lehtinen" authorityYear="2005" box="[906,1223,765,789]" class="Arachnida" family="Lycosidae" genus="Passiena" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="torbjoerni">
<emphasis id="B940EAC4186FFFD9FC1CFD3AFB9A4B76" box="[906,1046,765,789]" italics="true" pageId="1" pageNumber="480">P. torbjoerni</emphasis>
<bibRefCitation id="EFA54B27186FFFD9FBB6FD39FB4B4B76" author="Lehtinen PT" box="[1056,1223,765,789]" pageId="1" pageNumber="480" pagination="398 - 407" refId="ref4766" refString="Lehtinen PT (2005) Review of the Oriental wolf spider genus Passiena (Lycosidae, Pardosinae). Journal of Arachnology, 33: 398 - 407." type="journal article" year="2005">Lehtinen, 2005</bibRefCitation>
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, based on both male and female. The palp of the new species was illustrated but lacked essential details, such as the shape of the intact palp and the shape of the embolic division. Nevertheless, it was the first description of the palp of a
<taxonomicName id="4C344D55186FFFD9FB2FFCBAFA954BF6" authorityName="Thorell" authorityYear="1890" box="[1209,1305,894,917]" class="Arachnida" family="Lycosidae" genus="Passiena" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAC4186FFFD9FB2FFCBAFA954BF6" box="[1209,1305,894,917]" italics="true" pageId="1" pageNumber="480">Passiena</emphasis>
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male.
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We tried to compare the new species with two other congeners occurring in Asia, but failed to found
<typeStatus id="548F8874186FFFD9FB39FC1AFB6B4B95" box="[1199,1255,990,1014]" pageId="1" pageNumber="480">types</typeStatus>
listed as deposited in the Natural History Museum in Stockholm and Zoological Museum University of Turku (
<bibRefCitation id="EFA54B27186FFFD9FBE6FBD9FA9B4C56" author="Lehtinen PT" box="[1136,1303,1053,1077]" pageId="1" pageNumber="480" pagination="398 - 407" refId="ref4766" refString="Lehtinen PT (2005) Review of the Oriental wolf spider genus Passiena (Lycosidae, Pardosinae). Journal of Arachnology, 33: 398 - 407." type="journal article" year="2005">Lehtinen, 2005</bibRefCitation>
). Fortunately, the junior author has the original illustrations of these species made for Pekka Lehtinens publication.
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