302 lines
56 KiB
XML
302 lines
56 KiB
XML
<document id="D0E2B7DD5BB3217401F75CE08B7A8EC3" ID-DOI="10.11646/zootaxa.5004.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5140802" ID-ZooBank="86B5901B-A1B8-490A-9926-81B54D2C1D73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627453265329" checkinUser="plazi" docAuthor="Trýzna, Miloš & Baňař, Petr" docDate="2021" docId="039D1929D860CC1CFF5EFB78F0FDB814" docLanguage="en" docName="zootaxa.5004.4.4.pdf" docOrigin="Zootaxa 5004 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Portentus Tryzna & Banar 2021, gen. nov." docType="treatment" docUuid="A236CAE1-3103-4938-B004-C6955568646C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="553" masterDocId="FFA46151D861CC1EFFC9FF8BF561BF4B" masterDocTitle="Description of a new genus and species, Portentus marschneri (Coleoptera: Anthribidae), from Penang National Park, Malaysia" masterLastPageNumber="563" masterPageNumber="551" pageNumber="552" updateTime="1699024961175" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="6DC4A4A133C38E6039EA1AA989E7457F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3AB8D8A871AFC197D049190418EC7523">
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<mods:title id="612683CF97B7DD072BFB5E31942F2890">Description of a new genus and species, Portentus marschneri (Coleoptera: Anthribidae), from Penang National Park, Malaysia</mods:title>
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<mods:name id="1504F301F051ABA76EF4383F9DE8F580" type="personal">
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<mods:role id="D9BD1E74D83CA5882EECFED80AEED163">
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<mods:roleTerm id="0A990A138219D0EDFC6E5556DEDC2589">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6D1F7CD8061988161D6DBBFF7AA2D155">Trýzna, Miloš</mods:namePart>
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<mods:affiliation id="1B50AA0D89A2CB7F9BBF79876CE22D72">Municipal Museum of Ústí nad Labem, Natural History Department, Masarykova 1000 / 3, CZ- 400 01 Ústí nad Labem, Czech Republic. Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic.</mods:affiliation>
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</mods:name>
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<mods:name id="42B705C42BFC30ACAB34283708E5CF27" type="personal">
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<mods:role id="0C99A4002E6AADCD762A7C64061FBF55">
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<mods:roleTerm id="82D8304F7F447AD992C82C18EF7414BF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6C48C1E368EA00E4BE513A0016C16592">Baňař, Petr</mods:namePart>
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<mods:affiliation id="EA858E58A6AB58CAFF364812165B506C">Mendel University in Brno, Faculty of AgriSciences, Department of Zoology, Fisheries, Hydrobiology and Apiculture, Zemědělská 1, Brno, CZ- 613 00, Czech Republic.</mods:affiliation>
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<mods:typeOfResource id="1A95C3BF8B052A82F9C6008AA7877139">text</mods:typeOfResource>
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<mods:title id="6B682580AC22D2F7CB82CC134B16D1A3">Zootaxa</mods:title>
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<mods:part id="5632F8B74D992A660E2F790EB9A65CFA">
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<mods:date id="7DBED35DBFC545D25700C092F2B5FA8F">2021</mods:date>
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<mods:number id="A254AF9B88C42EAA05B331E36C0DDB7E">2021-07-22</mods:number>
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<mods:detail id="5264F0243AC2A878FD1A080B7E033522" type="volume">
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<mods:number id="E8A67352F23A82D03789D16E1C4F2CF9">5004</mods:number>
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<mods:detail id="B194750F7288613308C977184DAD7A07" type="issue">
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<mods:number id="863DAE97A2DCB515BA52D3CD98624567">4</mods:number>
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<mods:start id="B7599B73086AAFB2E32886CE0493BEF1">551</mods:start>
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<mods:end id="AB532B2EA285B807699D5724D59850AD">563</mods:end>
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<mods:classification id="6C5DD17460F359730BC7AE5EFB5A98E9">journal article</mods:classification>
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<mods:identifier id="6318360AD66DFFB03B39BA27ACC7C1E6" type="DOI">10.11646/zootaxa.5004.4.4</mods:identifier>
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<mods:identifier id="58E24AEADEEFFF2858D92950E828BDB2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039D1929D860CC1CFF5EFB78F0FDB814" LSID="urn:lsid:zoobank.org:act:A236CAE1-3103-4938-B004-C6955568646C" httpUri="http://treatment.plazi.org/id/039D1929D860CC1CFF5EFB78F0FDB814" lastPageId="2" lastPageNumber="553" pageId="1" pageNumber="552">
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<subSubSection id="C32EFBB4D860CC1FFF5EFB78F736BA46" box="[151,599,1266,1293]" pageId="1" pageNumber="552" type="nomenclature">
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<paragraph id="8B8BA83FD860CC1FFF5EFB78F736BA46" blockId="1.[151,599,1266,1329]" box="[151,599,1266,1293]" pageId="1" pageNumber="552">
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<heading id="D0C31F53D860CC1FFF5EFB78F736BA46" bold="true" box="[151,599,1266,1293]" fontSize="11" level="1" pageId="1" pageNumber="552" reason="1">
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<emphasis id="B940742DD860CC1FFF5EFB78F736BA46" bold="true" box="[151,599,1266,1293]" pageId="1" pageNumber="552">
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<taxonomicName id="4C34D3BCD860CC1FFF5EFB78F483BA46" authority="Tryzna & Banar" authorityName="Tryzna & Banar" authorityYear="2021" box="[151,482,1266,1293]" class="Insecta" family="Anthribidae" genus="Portentus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B940742DD860CC1FFF5EFB78F46DBA46" bold="true" box="[151,268,1267,1293]" italics="true" pageId="1" pageNumber="552">Portentus</emphasis>
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Trýzna & Baňař
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</taxonomicName>
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<taxonomicNameLabel id="A273C956D860CC1FFE23FB78F736BA46" box="[490,599,1267,1293]" pageId="1" pageNumber="552" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EFBB4D860CC1FFF5EFA9CF478BA7A" box="[151,281,1303,1329]" pageId="1" pageNumber="552" type="description">
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<paragraph id="8B8BA83FD860CC1FFF5EFA9CF478BA7A" blockId="1.[151,599,1266,1329]" box="[151,281,1303,1329]" pageId="1" pageNumber="552">
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(
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<figureCitation id="130FB4BAD860CC1FFF56FA9CF470BA7A" box="[159,273,1303,1329]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–11" captionStart-3="FIGURES 12–18" captionStart-4="FIGURES 19–22" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1565,1589]" captionStartId-2="5.[151,264,1864,1888]" captionStartId-3="6.[151,264,1943,1967]" captionStartId-4="7.[151,264,1492,1516]" captionTargetBox-0="[184,1403,191,1909]" captionTargetBox-1="[170,1432,183,1537]" captionTargetBox-2="[151,1435,849,1841]" captionTargetBox-3="[159,1428,181,1915]" captionTargetBox-4="[154,1435,181,1456]" captionTargetId-0="figure-17@3.[178,1409,181,1919]" captionTargetId-1="figure-18@4.[151,1435,181,1542]" captionTargetId-2="figure-293@5.[151,1435,849,1841]" captionTargetId-3="figure-18@6.[159,1428,181,1918]" captionTargetId-4="figure-17@7.[151,1435,181,1469]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." captionText-1="FIGURES 5–8. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 5, ventral habitus (all left tibiae and tarsi omitted); 6, head in ventral view; 7, right antenna in dorsal view; 8, abdomen in ventral view. Scale bars in mm." captionText-2="FIGURES 9–11. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 9, right hindwing; 10, metanotum in dorsal view; 11, abdominal tergites in dorsal view. Scale bars in mm. Abbreviations: A1, A2, A3 = anal vein 1, 2, 3 respectively; Ac = anal cell; alc = alacrista; C = costa; Cu = cubitus; J = jugal vein; M = media; R = radius; Rc = radial cell; Rs = radial sector; Sc = subcosta; sc3 = metascutum; scl3 = metascutellum." captionText-3="FIGURES 12–18. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 12, aedeagus in ventral view; 13, apex of aedeagus in ventral view; 14, segment VIII in ventral view; 15, sternite of segment IX (= spiculum gastrale) in ventral view; 16, pygidium in dorsal view; 17, apex of tegmen in ventral view; 18, tegmen in ventral view. Scale bars in mm." captionText-4="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi-0="http://doi.org/10.5281/zenodo.5140804" figureDoi-1="http://doi.org/10.5281/zenodo.5140806" figureDoi-2="http://doi.org/10.5281/zenodo.5140808" figureDoi-3="http://doi.org/10.5281/zenodo.5140810" figureDoi-4="http://doi.org/10.5281/zenodo.5140812" httpUri-0="https://zenodo.org/record/5140804/files/figure.png" httpUri-1="https://zenodo.org/record/5140806/files/figure.png" httpUri-2="https://zenodo.org/record/5140808/files/figure.png" httpUri-3="https://zenodo.org/record/5140810/files/figure.png" httpUri-4="https://zenodo.org/record/5140812/files/figure.png" pageId="1" pageNumber="552">Figs 1–22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EFBB4D860CC1FFF5EFAD7F1D4BAAA" pageId="1" pageNumber="552" type="type_taxon">
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<paragraph id="8B8BA83FD860CC1FFF5EFAD7F164BA3F" blockId="1.[151,1029,1371,1396]" box="[151,1029,1371,1396]" pageId="1" pageNumber="552">
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<emphasis id="B940742DD860CC1FFF5EFAD7F442BA38" bold="true" box="[151,291,1371,1396]" pageId="1" pageNumber="552">
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<typeStatus id="548F169DD860CC1FFF5EFAD7F5ADBA3F" box="[151,204,1372,1396]" pageId="1" pageNumber="552">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C34D3BCD860CC1FFEE3FAD7F7D1BA3F" authority="Tryzna & Banar" authorityName="Tryzna & Banar" authorityYear="2021" box="[298,688,1372,1396]" class="Insecta" family="Anthribidae" genus="Portentus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="marschneri" status="sp. nov.">
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<emphasis id="B940742DD860CC1FFEE3FAD7F767BA3F" box="[298,518,1372,1396]" italics="true" pageId="1" pageNumber="552">Portentus marschneri</emphasis>
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Trýzna & Baňař
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</taxonomicName>
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<emphasis id="B940742DD860CC1FFD7FFAD7F666BA3F" bold="true" box="[694,775,1372,1396]" pageId="1" pageNumber="552">
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<taxonomicNameLabel id="A273C956D860CC1FFD7FFAD7F666BA3F" box="[694,775,1372,1396]" pageId="1" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by original designation.
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</paragraph>
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<paragraph id="8B8BA83FD860CC1FFF5EFA29F7A0BAF7" blockId="1.[151,1437,1442,2009]" box="[151,705,1442,1468]" pageId="1" pageNumber="552">
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<emphasis id="B940742DD860CC1FFF5EFA29F47ABAF7" bold="true" box="[151,283,1442,1468]" pageId="1" pageNumber="552">Placement.</emphasis>
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Tribe
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<taxonomicName id="4C34D3BCD860CC1FFEADFA29F7A0BAF7" authority="Lacordaire, 1866" authorityName="Lacordaire" authorityYear="1866" box="[356,705,1442,1468]" class="Insecta" family="Anthribidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="tribe" tribe="Phloeophilini">
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Phloeophilini
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<bibRefCitation id="EFA5D5CED860CC1FFDCBFA29F7A0BAF7" author="Lacordaire, J. T." box="[514,705,1442,1468]" pageId="1" pageNumber="552" pagination="1 - 620" refId="ref5566" refString="Lacordaire, J. T. (1866) Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Roret, Paris, 7, 1 - 620." type="journal article" year="1866">Lacordaire, 1866</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8B8BA83FD860CC1FFF0EFA4DF1D4BAAA" blockId="1.[151,1437,1442,2009]" box="[199,1205,1477,1505]" pageId="1" pageNumber="552">
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<emphasis id="B940742DD860CC1FFF0EFA4DF4F7BA94" bold="true" box="[199,406,1477,1504]" pageId="1" pageNumber="552">Name derivation.</emphasis>
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<taxonomicName id="4C34D3BCD860CC1FFE54FA4CF76BBAAB" authorityName="Tryzna & Banar" authorityYear="2021" box="[413,522,1479,1504]" class="Insecta" family="Anthribidae" genus="Portentus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
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<emphasis id="B940742DD860CC1FFE54FA4CF76BBAAB" box="[413,522,1479,1504]" italics="true" pageId="1" pageNumber="552">Portentus</emphasis>
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</taxonomicName>
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, gender masculine, from Latin
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<emphasis id="B940742DD860CC1FFCA3FA4CF68CBAAB" box="[874,1005,1479,1504]" italics="true" pageId="1" pageNumber="552">portentosus</emphasis>
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(peculiar, weird).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EFBB4D860CC1FFF0EFA62F4D5B997" pageId="1" pageNumber="552" type="diagnosis">
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<paragraph id="8B8BA83FD860CC1FFF0EFA62F4D5B997" blockId="1.[151,1437,1442,2009]" pageId="1" pageNumber="552">
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<emphasis id="B940742DD860CC1FFF0EFA62F45BB948" bold="true" box="[199,314,1513,1539]" pageId="1" pageNumber="552">Diagnosis</emphasis>
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. Head elongate, eyes entire, not emarginate. Antennal scrobes visible from above. Antennae shorter, not reaching posterior margin of pronotum, segments of funicle distinctly thick, three-segmented club well-developed, pronotum narrower than elytral base, with two wide, blunt tubercles situated laterally. Dorsal transverse carina distinctly antebasal with very wide posterior pronotal declivity, shortly interrupted in middle, lateral carina of pronotum very short and almost straight, in contact with dorsal transverse carina forming obtuse angle. Elytra suboval, narrowed evenly in distal third of length, each elytron with two tubercles, one in sub-basal and second in median portion of length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EFBB4D860CC1CFF0EF96EF0EEBCED" lastPageId="2" lastPageNumber="553" pageId="1" pageNumber="552" type="description">
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<paragraph id="8B8BA83FD860CC1CFF0EF96EF719BE0A" blockId="1.[151,1437,1442,2009]" lastBlockId="2.[151,1437,150,1149]" lastPageId="2" lastPageNumber="553" pageId="1" pageNumber="552">
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<emphasis id="B940742DD860CC1FFF0EF96EF438B9B4" bold="true" box="[199,345,1765,1791]" pageId="1" pageNumber="552">Description.</emphasis>
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<emphasis id="B940742DD860CC1FFEA9F96DF4FCB84B" box="[352,413,1766,1792]" italics="true" pageId="1" pageNumber="552">Head</emphasis>
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(
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<figureCitation id="130FB4BAD860CC1FFE62F96DF76FB84B" box="[427,526,1766,1792]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="1" pageNumber="552">Figs 1, 3</figureCitation>
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,
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<figureCitation id="130FB4BAD860CC1FFDD2F96DF74BB84A" box="[539,554,1766,1793]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1565,1589]" captionTargetBox="[170,1432,183,1537]" captionTargetId="figure-18@4.[151,1435,181,1542]" captionTargetPageId="4" captionText="FIGURES 5–8. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 5, ventral habitus (all left tibiae and tarsi omitted); 6, head in ventral view; 7, right antenna in dorsal view; 8, abdomen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140806" httpUri="https://zenodo.org/record/5140806/files/figure.png" pageId="1" pageNumber="552">6</figureCitation>
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) elongate, eyes of medium size, situated laterally, hemispherical, convex laterally, not emarginate, well-separated from each other (dorsal ocular index ca. 2.00). Rostrum with two longitudinal lateral carinae reaching from posterior margins of scrobes to anterior margins of eyes, space between carinae depressed. Lateral margins of rostrum almost parallel, very slightly expanding anteriorly in dorsal view, anterior part moderately emarginate. Antennal scrobes visible in dorsal view (
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<figureCitation id="130FB4BAD860CC1FFCAFF8FDF6C9B8DB" box="[870,936,1910,1936]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="1" pageNumber="552">Fig. 3</figureCitation>
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), margins of scrobes not elevated, carinate at their dorsal margins. Scrobes situated in distal half of rostrum, almost reaching bases of mandibles anteriorly, distant from anterior margin of eyes. Lower side of scrobes (
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<figureCitation id="130FB4BAD860CC1FFCD1F835F63AB892" box="[792,859,1982,2009]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1565,1589]" captionTargetBox="[170,1432,183,1537]" captionTargetId="figure-18@4.[151,1435,181,1542]" captionTargetPageId="4" captionText="FIGURES 5–8. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 5, ventral habitus (all left tibiae and tarsi omitted); 6, head in ventral view; 7, right antenna in dorsal view; 8, abdomen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140806" httpUri="https://zenodo.org/record/5140806/files/figure.png" pageId="1" pageNumber="552">Fig. 6</figureCitation>
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) open posteriorly, this opening of scrobes distinctly visible in dorsal view, dorsal part of rostrum narrowest at this point.
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<emphasis id="B940742DD863CC1CFC60FF1CF172BFFA" box="[937,1043,151,177]" italics="true" pageId="2" pageNumber="553">Antennae</emphasis>
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(
|
||
<figureCitation id="130FB4BAD863CC1CFBEAFF1CF10DBFFA" box="[1059,1132,151,177]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFBB2FF1DF1C9BFFA" box="[1147,1192,150,177]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1565,1589]" captionTargetBox="[170,1432,183,1537]" captionTargetId="figure-18@4.[151,1435,181,1542]" captionTargetPageId="4" captionText="FIGURES 5–8. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 5, ventral habitus (all left tibiae and tarsi omitted); 6, head in ventral view; 7, right antenna in dorsal view; 8, abdomen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140806" httpUri="https://zenodo.org/record/5140806/files/figure.png" pageId="2" pageNumber="553">5, 7</figureCitation>
|
||
) short, antennomeres III–VIII of similar lengths and widths, IX triangular, slightly longer than wide, X distinctly wider than long, XI triangular, slightly longer than wide with widely rounded tip. Stalk (= basal-most part of scape) hidden in scrobe in dorsal view, visible part of scape almost spherical, pedicel longer than each of antennomeres III–VIII. Antennae with appressed setae, more visible on club.
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFEC0F438BD2A" blockId="2.[151,1437,150,1149]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFEC0F457BE2F" box="[199,310,331,356]" italics="true" pageId="2" pageNumber="553">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="130FB4BAD863CC1CFE8EFEC0F4CFBE2E" box="[327,430,331,357]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFE75FEC0F762BE2E" box="[444,515,331,357]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[154,1435,181,1456]" captionTargetId="figure-17@7.[151,1435,181,1469]" captionTargetPageId="7" captionText="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi="http://doi.org/10.5281/zenodo.5140812" httpUri="https://zenodo.org/record/5140812/files/figure.png" pageId="2" pageNumber="553">19–22</figureCitation>
|
||
) almost as long as wide, widest in middle. Dorsal and lateral parts of pronotum with very coarse irregular punctation. Disc (
|
||
<figureCitation id="130FB4BAD863CC1CFD48FEE4F785BEC2" box="[641,740,367,393]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 2, 4</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFD39FEE4F656BEC2" box="[752,823,367,393]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[154,1435,181,1456]" captionTargetId="figure-17@7.[151,1435,181,1469]" captionTargetPageId="7" captionText="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi="http://doi.org/10.5281/zenodo.5140812" httpUri="https://zenodo.org/record/5140812/files/figure.png" pageId="2" pageNumber="553">19–22</figureCitation>
|
||
) with two tubercles in lateral portions, tips of tubercles bluntly rounded. Dorsal transverse carina very slightly bisinuate, shortly, indistinctly interrupted at middle, lateral carinae short, straight, reaching approximately to middle of pronotum. Secondary carina (sensu
|
||
<bibRefCitation id="EFA5D5CED863CC1CFB29FE3CF5D2BEBE" author="Holloway, B. A." pageId="2" pageNumber="553" refId="ref5438" refString="Holloway, B. A. (1982) Anthribidae (Insecta: Coleoptera). Fauna of New Zealand. Vol. 3. Science Information Division, DSIR, Wellington, 264 pp." type="book" year="1982">Holloway, 1982: 21</bibRefCitation>
|
||
and 187,
|
||
<figureCitation id="130FB4BAD863CC1CFEEFFE50F419BEBE" box="[294,376,475,501]" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[159,1428,181,1915]" captionTargetId="figure-18@6.[159,1428,181,1918]" captionTargetPageId="6" captionText="FIGURES 12–18. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 12, aedeagus in ventral view; 13, apex of aedeagus in ventral view; 14, segment VIII in ventral view; 15, sternite of segment IX (= spiculum gastrale) in ventral view; 16, pygidium in dorsal view; 17, apex of tegmen in ventral view; 18, tegmen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140810" httpUri="https://zenodo.org/record/5140810/files/figure.png" pageId="2" pageNumber="553">Fig. 14</figureCitation>
|
||
) absent, with only poorly defined irregularly interrupted transverse callus. Basal longitudinal carinulae of pronotum (sensu
|
||
<bibRefCitation id="EFA5D5CED863CC1CFE24FE74F7DDBD52" author="Jordan, K." box="[493,700,510,537]" pageId="2" pageNumber="553" pagination="151 - 158" refId="ref5523" refString="Jordan, K. (1928) Some Anthribidae collected by R. E. Turner in South Africa. Novitates Zoologicae, 34, 151 - 158. https: // doi. org / 10.5962 / bhl. part. 11789" type="journal article" year="1928">
|
||
<collectingCountry id="F323E8AFD863CC1CFE24FE74F75ABD52" box="[493,571,511,537]" name="Jordan" pageId="2" pageNumber="553">Jordan</collectingCountry>
|
||
, 1928: 151
|
||
</bibRefCitation>
|
||
) (= two carinulae on lateral sides of pronotum, visible in lateral view between dorsal transverse carina and posterior margin of pronotum), present, well-developed, almost straight (see
|
||
<figureCitation id="130FB4BAD863CC1CFF02FDCCF42CBD2A" box="[203,333,583,609]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[154,1435,181,1456]" captionTargetId="figure-17@7.[151,1435,181,1469]" captionTargetPageId="7" captionText="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi="http://doi.org/10.5281/zenodo.5140812" httpUri="https://zenodo.org/record/5140812/files/figure.png" pageId="2" pageNumber="553">Figs 19–22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFDE0F1CDBDBA" blockId="2.[151,1437,150,1149]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFDE0F46CBDCE" box="[199,269,619,645]" italics="true" pageId="2" pageNumber="553">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="130FB4BAD863CC1CFED5FDE0F41EBDCE" box="[284,383,619,645]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 1, 4</figureCitation>
|
||
) suboval, sides parallel, narrowed even in distal third of length. Humeri well-developed. Each elytron with two tubercles, one in sub-basal area with apex on elytral interval I, and the second in median area with apex on elytral interval II. Cross-section of sub-basal tubercles rather spherical, that of median tubercles suboval. Elytral declivity decreasing posteriorly immediately from distal margins of median tubercles.
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFD70F0E6BC5D" blockId="2.[151,1437,150,1149]" box="[199,1415,763,790]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFD70F426BC5F" box="[199,327,763,788]" italics="true" pageId="2" pageNumber="553">Metanotum</emphasis>
|
||
(
|
||
<figureCitation id="130FB4BAD863CC1CFE9FFD70F4C6BC5E" box="[342,423,763,789]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1864,1888]" captionTargetBox="[151,1435,849,1841]" captionTargetId="figure-293@5.[151,1435,849,1841]" captionTargetPageId="5" captionText="FIGURES 9–11. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 9, right hindwing; 10, metanotum in dorsal view; 11, abdominal tergites in dorsal view. Scale bars in mm. Abbreviations: A1, A2, A3 = anal vein 1, 2, 3 respectively; Ac = anal cell; alc = alacrista; C = costa; Cu = cubitus; J = jugal vein; M = media; R = radius; Rc = radial cell; Rs = radial sector; Sc = subcosta; sc3 = metascutum; scl3 = metascutellum." figureDoi="http://doi.org/10.5281/zenodo.5140808" httpUri="https://zenodo.org/record/5140808/files/figure.png" pageId="2" pageNumber="553">Fig. 10</figureCitation>
|
||
) robust, wide. Hindwing (
|
||
<figureCitation id="130FB4BAD863CC1CFD07FD70F673BC5E" box="[718,786,763,789]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1864,1888]" captionTargetBox="[151,1435,849,1841]" captionTargetId="figure-293@5.[151,1435,849,1841]" captionTargetPageId="5" captionText="FIGURES 9–11. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 9, right hindwing; 10, metanotum in dorsal view; 11, abdominal tergites in dorsal view. Scale bars in mm. Abbreviations: A1, A2, A3 = anal vein 1, 2, 3 respectively; Ac = anal cell; alc = alacrista; C = costa; Cu = cubitus; J = jugal vein; M = media; R = radius; Rc = radial cell; Rs = radial sector; Sc = subcosta; sc3 = metascutum; scl3 = metascutellum." figureDoi="http://doi.org/10.5281/zenodo.5140808" httpUri="https://zenodo.org/record/5140808/files/figure.png" pageId="2" pageNumber="553">Fig. 9</figureCitation>
|
||
) with well-developed main veins (not studied in detail).
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFC94F149BC72" blockId="2.[151,1437,150,1149]" box="[199,1064,799,825]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFC94F59DBC73" box="[199,252,799,824]" italics="true" pageId="2" pageNumber="553">Legs</emphasis>
|
||
relatively robust, tarsomere I longer than II, inner tooth of claw present.
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFCC8F0EEBCED" blockId="2.[151,1437,150,1149]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFCC8F450BC16" box="[199,305,835,861]" italics="true" pageId="2" pageNumber="553">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="130FB4BAD863CC1CFE89FCC8F4E0BC16" box="[320,385,835,861]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1565,1589]" captionTargetBox="[170,1432,183,1537]" captionTargetId="figure-18@4.[151,1435,181,1542]" captionTargetPageId="4" captionText="FIGURES 5–8. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 5, ventral habitus (all left tibiae and tarsi omitted); 6, head in ventral view; 7, right antenna in dorsal view; 8, abdomen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140806" httpUri="https://zenodo.org/record/5140806/files/figure.png" pageId="2" pageNumber="553">Fig. 8</figureCitation>
|
||
) shorter than broad, sternites forming smooth concave arch in lateral view. Pygidium (
|
||
<figureCitation id="130FB4BAD863CC1CFA8AFCC8F0F5BC15" box="[1347,1428,835,862]" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[159,1428,181,1915]" captionTargetId="figure-18@6.[159,1428,181,1918]" captionTargetPageId="6" captionText="FIGURES 12–18. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 12, aedeagus in ventral view; 13, apex of aedeagus in ventral view; 14, segment VIII in ventral view; 15, sternite of segment IX (= spiculum gastrale) in ventral view; 16, pygidium in dorsal view; 17, apex of tegmen in ventral view; 18, tegmen in ventral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140810" httpUri="https://zenodo.org/record/5140810/files/figure.png" pageId="2" pageNumber="553">Fig. 16</figureCitation>
|
||
) wider than long. Abdominal tergites (with partly damaged metapostnotum in its anterior portion) (
|
||
<figureCitation id="130FB4BAD863CC1CFB1DFCECF043BCCA" box="[1236,1314,871,897]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1864,1888]" captionTargetBox="[151,1435,849,1841]" captionTargetId="figure-293@5.[151,1435,849,1841]" captionTargetPageId="5" captionText="FIGURES 9–11. Portentus marschneri Trýzna & Baňař sp. nov., male holotype; 9, right hindwing; 10, metanotum in dorsal view; 11, abdominal tergites in dorsal view. Scale bars in mm. Abbreviations: A1, A2, A3 = anal vein 1, 2, 3 respectively; Ac = anal cell; alc = alacrista; C = costa; Cu = cubitus; J = jugal vein; M = media; R = radius; Rc = radial cell; Rs = radial sector; Sc = subcosta; sc3 = metascutum; scl3 = metascutellum." figureDoi="http://doi.org/10.5281/zenodo.5140808" httpUri="https://zenodo.org/record/5140808/files/figure.png" pageId="2" pageNumber="553">Fig. 11</figureCitation>
|
||
) I–IV subequal in size, tergites V–VI smaller, spiracula on tergite I much larger than in other tergites (not studied in detail).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EFBB4D863CC1CFF0EFC25F78ABB36" pageId="2" pageNumber="553" type="diagnosis">
|
||
<paragraph id="8B8BA83FD863CC1CFF0EFC25F78ABB36" blockId="2.[151,1437,150,1149]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFF0EFC25F4ACBC83" bold="true" box="[199,461,942,968]" pageId="2" pageNumber="553">Differential diagnosis.</emphasis>
|
||
We place
|
||
<taxonomicName id="4C34D3BCD863CC1CFDF6FC24F7CDBC83" authority="Trýzna & Baňař, 2021" authorityName="Trýzna & Baňař" authorityYear="2021" box="[575,684,943,968]" class="Insecta" family="Anthribidae" genus="Portentus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940742DD863CC1CFDF6FC24F7CDBC83" box="[575,684,943,968]" italics="true" pageId="2" pageNumber="553">Portentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940742DD863CC1CFD7BFC24F67ABC82" bold="true" box="[690,795,943,969]" pageId="2" pageNumber="553">
|
||
<taxonomicNameLabel id="A273C956D863CC1CFD7BFC24F67ABC82" box="[690,795,943,969]" pageId="2" pageNumber="553" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the tribe
|
||
<taxonomicName id="4C34D3BCD863CC1CFC56FC24F19FBC82" authority="Lacordaire, 1866" authorityName="Lacordaire" authorityYear="1866" box="[927,1278,943,969]" class="Insecta" family="Anthribidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="tribe" tribe="Phloeophilini">
|
||
Phloeophilini
|
||
<bibRefCitation id="EFA5D5CED863CC1CFBF5FC24F19FBC82" author="Lacordaire, J. T." box="[1084,1278,943,969]" pageId="2" pageNumber="553" pagination="1 - 620" refId="ref5566" refString="Lacordaire, J. T. (1866) Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Roret, Paris, 7, 1 - 620." type="journal article" year="1866">Lacordaire, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
, based mainly on the following morphological characters: rostrum flat, grooved or with one or two longitudinal carinae; eyes situated lateraly, oval or suboval, not emarginate; dorsal transverse carina of pronotum conspicuous, antebasal, i.e. markedly distant from posterior margin of pronotum, not forming a lateral lobe, in contact with lateral carina forming obtuse angle or wide arch. The new genus can be distinguished from other Palaearctic and Oriental genera by the morphological characters given in the key below:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EFBB4D863CC1CFF5EFB23F0FDB814" pageId="2" pageNumber="553" type="key">
|
||
<key id="FFA5FF0ED863CC1CFF5EFB23F0FDB814" pageId="2" pageNumber="553">
|
||
<keyStep id="30C0B09AD863CC1CFF5EFB23F0FDBA5E" pageId="2" pageNumber="553">
|
||
<paragraph id="8B8BA83FD863CC1CFF5EFB23F0FDBB91" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EFB23F0FDBB91" pageId="2" pageNumber="553">
|
||
1 Antennal scrobes distinctly visible in dorsal view. Rostrum without medial groove, only more or less depressed in median area (
|
||
<figureCitation id="130FB4BAD863CC1CFF20FB4EF447BB91" box="[233,294,1221,1243]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFEF8FB4EF42BBB90" box="[305,330,1221,1243]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">25</figureCitation>
|
||
).......................................................................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF5EFB68F0FDBA5E" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EFB68F0FDBA5E" pageId="2" pageNumber="553">
|
||
- Antennal scrobes not distinctly visible in dorsal view. Rostrum grooved in basal part, medial longitudinal groove reaching from posterior margin of scrobes to line of middle of eyes or to posterior margin of eyes (
|
||
<figureCitation id="130FB4BAD863CC1CFBCEFA8BF133BA5E" box="[1031,1106,1280,1302]" captionStart="FIGURES 28–32" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[205,1383,208,1912]" captionTargetId="figure-17@9.[203,1385,181,1920]" captionTargetPageId="9" captionText="FIGURES 28–32. Phloeophilus oculatus (Jordan, 1904), male from Laos, Phongsaly prov., Phongsaly env.; 28, dorsal habitus; 29, pronotum in dorsal view; 30, head in dorsal view; 31, right antenna in dorsal view; 32, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140816" httpUri="https://zenodo.org/record/5140816/files/figure.png" pageId="2" pageNumber="553">Figs 30</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFB95FB74F114BA5E" box="[1116,1141,1279,1301]" captionStart="FIGURES 33–37" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[187,1406,189,1919]" captionTargetId="figure-18@10.[178,1409,181,1919]" captionTargetPageId="10" captionText="FIGURES 33–37. Bothrus aspersus (Jordan, 1923), male from Thailand, Chiang Mai env.; 33, dorsal habitus; 34, pronotum in dorsal view; 35, head in dorsal view; 36, right antenna in dorsal view; 37, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140818" httpUri="https://zenodo.org/record/5140818/files/figure.png" pageId="2" pageNumber="553">35</figureCitation>
|
||
)...................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C0B09AD863CC1CFF5EFA96F0FDB987" pageId="2" pageNumber="553">
|
||
<paragraph id="8B8BA83FD863CC1CFF5EFA96F0FDBAA9" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EFA96F0FDBAA9" pageId="2" pageNumber="553">
|
||
2 Dorsal part of antennal scrobes not reflexed and carinate (
|
||
<figureCitation id="130FB4BAD863CC1CFCC3FA96F63DBA78" box="[778,860,1309,1331]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Fig. 3, 4</figureCitation>
|
||
). Rostrum between margins of scrobes flat and with two lateral longitudinal carinae reaching from anterior margins of eyes to posterior margins of scrobes (
|
||
<figureCitation id="130FB4BAD863CC1CFBBBFAB1F1CABA1B" box="[1138,1195,1338,1360]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Fig. 3</figureCitation>
|
||
). Scape almost spherical, distinctly wider than pedicel, antennae forming a distinctly 3-segmented separate antennal club (in male) (
|
||
<figureCitation id="130FB4BAD863CC1CFB1FFAD3F04ABA25" box="[1238,1323,1368,1390]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 1, 3</figureCitation>
|
||
). Pronotum with two lateral tubercles on disc, each elytron with two tubercles on the second elytral interval in sub-basal and median areas (
|
||
<figureCitation id="130FB4BAD863CC1CFF20FA19F45EBAEC" box="[233,319,1426,1448]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Figs 2, 4</figureCitation>
|
||
,
|
||
<figureCitation id="130FB4BAD863CC1CFE81FA19F4E9BAE3" box="[328,392,1426,1448]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[154,1435,181,1456]" captionTargetId="figure-17@7.[151,1435,181,1469]" captionTargetPageId="7" captionText="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi="http://doi.org/10.5281/zenodo.5140812" httpUri="https://zenodo.org/record/5140812/files/figure.png" pageId="2" pageNumber="553">19–20</figureCitation>
|
||
). Upper side of body with coarse punctation (
|
||
<figureCitation id="130FB4BAD863CC1CFCF2FA19F6ECBAE3" box="[827,909,1426,1448]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-17@3.[178,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Portentus marschneri Trýzna & Baňař sp. nov., male holotype from Malaysia, Penang National Park; 1, dorsal habitus; 2, pronotum in dorsal view; 3, head in dorsal view; 4, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140804" httpUri="https://zenodo.org/record/5140804/files/figure.png" pageId="2" pageNumber="553">Fig. 1, 2</figureCitation>
|
||
). Dorsal transverse carina in contact with lateral carina forming obtuse angle, basal longitudinal carinulae of pronotum distinctly developed (
|
||
<figureCitation id="130FB4BAD863CC1CFBDFFA3BF13ABA8D" box="[1046,1115,1456,1478]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[154,1435,181,1456]" captionTargetId="figure-17@7.[151,1435,181,1469]" captionTargetPageId="7" captionText="FIGURES 19–22. Portentus marschneri Trýzna & Baňař sp. nov., male holotype, pronotum; 19, 21, dorsal view; 20, 22, left side in lateral view; 21–22, schematic; BLC = basal longitudinal carinula; BR = basal rim; DTC = dorsal transverse carina; LC = lateral carina; PD = pronotal declivity; TC = transverse callus." figureDoi="http://doi.org/10.5281/zenodo.5140812" httpUri="https://zenodo.org/record/5140812/files/figure.png" pageId="2" pageNumber="553">Fig. 20</figureCitation>
|
||
)..............................................................................................
|
||
<emphasis id="B940742DD863CC1CFBE7FA46F0FDBAA9" bold="true" box="[1070,1436,1484,1507]" pageId="2" pageNumber="553">
|
||
<taxonomicName id="4C34D3BCD863CC1CFBE7FA46F05DBAA8" authority="Tryzna & Banar" authorityName="Tryzna & Banar" authorityYear="2021" box="[1070,1340,1484,1507]" class="Insecta" family="Anthribidae" genus="Portentus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940742DD863CC1CFBE7FA46F1EFBAA9" bold="true" box="[1070,1166,1485,1506]" italics="true" pageId="2" pageNumber="553">Portentus</emphasis>
|
||
Trýzna & Baňař
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273C956D863CC1CFA8BFA46F0FDBAA9" box="[1346,1436,1485,1506]" pageId="2" pageNumber="553" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF5EFA60F0FDB987" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EFA60F0FDB987" pageId="2" pageNumber="553">
|
||
- Dorsal part of antennal scrobes reflexed and carinate, forming a more or less top-shaped pattern, higher than the apical part of the rostrum (
|
||
<figureCitation id="130FB4BAD863CC1CFE95F983F4A6B955" box="[348,455,1543,1566]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Fig. 25, 27</figureCitation>
|
||
). Rostrum between elevated margins of scrobes depressed, and with one indistinct median longitudinal carina reaching from anterior margins of eyes to area between scrobes (
|
||
<figureCitation id="130FB4BAD863CC1CFC5EF9AEF6BEB970" box="[919,991,1573,1595]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Fig. 25</figureCitation>
|
||
). Scape and pedicel of the same width, antennae from antennomere VIII continuously extended apically, not forming a sharply distinct and separate antennal club (in male) (
|
||
<figureCitation id="130FB4BAD863CC1CFF20F9EBF439B93D" box="[233,344,1632,1654]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Figs 23, 26</figureCitation>
|
||
), or only weakly formed (in female). Pronotum and elytra without tubercles (
|
||
<figureCitation id="130FB4BAD863CC1CFBF7F9EBF1B3B93D" box="[1086,1234,1632,1654]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Figs 23–24, 27</figureCitation>
|
||
). Upper side of body with only fine punctation (
|
||
<figureCitation id="130FB4BAD863CC1CFE2BF9F6F74BB9D8" box="[482,554,1661,1683]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Fig. 23</figureCitation>
|
||
). Dorsal transverse carina roundly connected with lateral carina, formed in wide arch, basal longitudinal carinulae of pronotum lacking or very weakly developed (
|
||
<figureCitation id="130FB4BAD863CC1CFC40F911F696B9FB" box="[905,1015,1690,1712]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[180,1405,188,1914]" captionTargetId="figure-18@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURES 23–27. Morimotanthribus leigongshani Li, Trýzna & Guo, 2019, male paratype from China, W Guizhou prov., Leigongshan, Xijiang; 23, dorsal habitus; 24, pronotum in dorsal view; 25, head in dorsal view; 26, right antenna in dorsal view; 27, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140814" httpUri="https://zenodo.org/record/5140814/files/figure.png" pageId="2" pageNumber="553">Figs 24, 27</figureCitation>
|
||
)..................................................................................................
|
||
<taxonomicName id="4C34D3BCD863CC1CFC35F93CF0FDB987" authority="Senoh & Tryzna, 2006" authorityName="Senoh & Tryzna" authorityYear="2006" box="[1020,1436,1719,1741]" class="Insecta" family="Anthribidae" genus="Morimotanthribus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940742DD863CC1CFC35F93CF0FDB987" bold="true" box="[1020,1436,1719,1741]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFC35F93CF1D2B986" bold="true" box="[1020,1203,1719,1741]" italics="true" pageId="2" pageNumber="553">Morimotanthribus</emphasis>
|
||
<bibRefCitation id="EFA5D5CED863CC1CFB70F93CF0FDB987" author="Senoh, T. & Tryzna, M." box="[1209,1436,1719,1741]" pageId="2" pageNumber="553" pagination="397 - 401" refId="ref5806" refString="Senoh, T. & Tryzna, M. (2006) Morimotanthribus chinensis (Coleoptera, Anthribidae), a new genus and species from China. Elytra, 34 (2), 397 - 401." type="journal article" year="2006">Senoh & Trýzna, 2006</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C0B09AD863CC1CFF5EF95EF0FDB814" pageId="2" pageNumber="553">
|
||
<paragraph id="8B8BA83FD863CC1CFF5EF95EF0FDB84C" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EF95EF0FDB84C" pageId="2" pageNumber="553">
|
||
3 Two dorso-lateral carinae of rostrum distinct, long, parallel or divergent posteriorly. Central longitudinal groove of rostrum narrow and shallow, not extended apically (
|
||
<figureCitation id="130FB4BAD863CC1CFD4DF979F7ABB843" box="[644,714,1778,1800]" captionStart="FIGURES 28–32" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[205,1383,208,1912]" captionTargetId="figure-17@9.[203,1385,181,1920]" captionTargetPageId="9" captionText="FIGURES 28–32. Phloeophilus oculatus (Jordan, 1904), male from Laos, Phongsaly prov., Phongsaly env.; 28, dorsal habitus; 29, pronotum in dorsal view; 30, head in dorsal view; 31, right antenna in dorsal view; 32, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140816" httpUri="https://zenodo.org/record/5140816/files/figure.png" pageId="2" pageNumber="553">Fig. 30</figureCitation>
|
||
).................................
|
||
<taxonomicName id="4C34D3BCD863CC1CFBAAF979F0FDB84C" ID-CoL="8HPFL" authority="Schoenherr, 1833" authorityName="Schoenherr" authorityYear="1833" box="[1123,1436,1778,1800]" class="Insecta" family="Anthribidae" genus="Phloeophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940742DD863CC1CFBAAF979F0FDB84C" bold="true" box="[1123,1436,1778,1800]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFBAAF979F187B843" bold="true" box="[1123,1254,1778,1800]" italics="true" pageId="2" pageNumber="553">Phloeophilus</emphasis>
|
||
Schoenherr, 1833
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8BA83FD863CC1CFF5EF89BF0FDB814" blockId="2.[151,1437,1192,1888]" pageId="2" pageNumber="553">
|
||
<keyLead id="30C50B0AD863CC1CFF5EF89BF0FDB814" pageId="2" pageNumber="553">
|
||
- Two dorso-lateral carinae of rostrum indistinct, short, reaching from inner edge of eyes obliquely inwards to central longitudinal groove. Central longitudinal groove wide and very deep, extended apically in shape of letter Y or T (
|
||
<figureCitation id="130FB4BAD863CC1CFB03F8A6F071B808" box="[1226,1296,1837,1859]" captionStart="FIGURES 33–37" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[187,1406,189,1919]" captionTargetId="figure-18@10.[178,1409,181,1919]" captionTargetPageId="10" captionText="FIGURES 33–37. Bothrus aspersus (Jordan, 1923), male from Thailand, Chiang Mai env.; 33, dorsal habitus; 34, pronotum in dorsal view; 35, head in dorsal view; 36, right antenna in dorsal view; 37, lateral habitus. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5140818" httpUri="https://zenodo.org/record/5140818/files/figure.png" pageId="2" pageNumber="553">Fig. 35</figureCitation>
|
||
)..........................................................................................
|
||
<taxonomicName id="4C34D3BCD863CC1CFB64F8C1F0FDB814" ID-CoL="8HF2L" authority="Wolfrum, 1956" authorityName="Wolfrum" authorityYear="1956" box="[1197,1436,1866,1888]" class="Insecta" family="Anthribidae" genus="Bothrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940742DD863CC1CFB64F8C1F0FDB814" bold="true" box="[1197,1436,1866,1888]" pageId="2" pageNumber="553">
|
||
<emphasis id="B940742DD863CC1CFB64F8C1F19CB82B" bold="true" box="[1197,1277,1866,1888]" italics="true" pageId="2" pageNumber="553">Bothrus</emphasis>
|
||
Wolfrum, 1956
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |