196 lines
18 KiB
XML
196 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6086663" ID-GBIF-Dataset="f9566778-1df4-44d6-9a4c-849de39a4b6c" ID-GBIF-Taxon="124955173" ID-Zenodo-Dep="6086663" checkinTime="1480975261095" checkinUser="plazi" docAuthor="Vincent J. Kalkman" docDate="2016" docId="03938796FFC5FFF326F889C9FEFDFB93" docLanguage="en" docName="10.5281zenodo.163451_Kalkman.pdf" docOrigin="Odonatologica 45 (3)" docStyle="DocumentStyle{}" docTitle="Celebargiolestes Kennedy 1925" docType="treatment" docVersion="12" lastPageId="29" lastPageNumber="264" masterDocId="FFAAFFEEFFDEFFEE27258934FFCBFF99" masterDocTitle="Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)" masterLastPageNumber="269" masterPageNumber="235" pageId="27" pageNumber="262" updateTime="1644906037961" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vincent J. Kalkman</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>235</mods:start>
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<mods:end>269</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.163451</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f9566778-1df4-44d6-9a4c-849de39a4b6c</mods:identifier>
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<mods:identifier type="Zenodo-Dep">163451</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6086663" ID-GBIF-Taxon="124955173" ID-Zenodo-Dep="6086663" LSID="urn:lsid:plazi:treatment:03938796FFC5FFF326F889C9FEFDFB93" httpUri="http://treatment.plazi.org/id/03938796FFC5FFF326F889C9FEFDFB93" lastPageId="29" lastPageNumber="264" pageId="27" pageNumber="262">
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<subSubSection box="[477,718,253,281]" pageId="27" pageNumber="262" type="nomenclature">
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<paragraph blockId="27.[159,1035,253,417]" box="[477,718,253,281]" pageId="27" pageNumber="262">
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<heading bold="true" box="[477,718,253,281]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="262" reason="0">
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<emphasis bold="true" box="[477,718,253,281]" pageId="27" pageNumber="262">
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[477,676,253,281]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[477,676,253,281]" italics="true" pageId="27" pageNumber="262">Celebargiolestes</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="262" type="materials_examined">
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<paragraph blockId="27.[159,1035,253,417]" box="[159,360,299,325]" pageId="27" pageNumber="262">
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<heading bold="true" box="[159,360,299,325]" fontSize="11" level="3" pageId="27" pageNumber="262" reason="0">
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<emphasis bold="true" box="[159,360,299,325]" pageId="27" pageNumber="262">Material studied</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="27.[159,1035,253,417]" box="[159,363,331,356]" pageId="27" pageNumber="262">
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<heading bold="true" box="[159,363,331,356]" fontSize="10" level="6" pageId="27" pageNumber="262" reason="6">
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<emphasis bold="true" box="[159,363,331,356]" pageId="27" pageNumber="262">Sulawesi Tenggara</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="27.[159,1035,253,417]" pageId="27" pageNumber="262">2♀, Tenggara, Palau Kabaena, 1 km S of Tangkeno, 550m, 08−09-xi-1989, R. de Jong & J. Huisman leg.</paragraph>
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</subSubSection>
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<caption httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="27" pageNumber="262" targetBox="[156,1038,480,1372]" targetPageId="27">
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<paragraph blockId="27.[159,1037,1413,1555]" pageId="27" pageNumber="262">
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<emphasis bold="true" box="[159,311,1413,1438]" pageId="27" pageNumber="262">Figures 34–39</emphasis>
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. Appendages of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[494,670,1413,1437]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="genus">
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<emphasis box="[494,670,1413,1437]" italics="true" pageId="27" pageNumber="262">Celebargiolestes</emphasis>
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</taxonomicName>
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males. 34 –
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<taxonomicName authority=", Ranotongkor" authorityName="Ranotongkor" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[807,910,1413,1437]" italics="true" pageId="27" pageNumber="262">C. cinctus</emphasis>
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, Ranotongkor
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</taxonomicName>
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, 02-viii-1940, JvT6476, dorsal view; 35 –
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<taxonomicName authority=", Ranotongkor" authorityName="Ranotongkor" box="[636,892,1442,1467]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[636,738,1442,1466]" italics="true" pageId="27" pageNumber="262">C. cinctus</emphasis>
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, Ranotongkor
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</taxonomicName>
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, 02-viii-1940, JvT6476, lateral view; 36 – Ranotongkor, 02-viii-1940, JvT6476, ventral view; 37 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[159,259,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis box="[159,259,1501,1525]" italics="true" pageId="27" pageNumber="262">C. askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[265,351,1501,1526]" pageId="27" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, dorsal view; 38 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[657,757,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis box="[657,757,1501,1525]" italics="true" pageId="27" pageNumber="262">C. askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[763,850,1501,1526]" pageId="27" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, lateral view; 39 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[275,374,1530,1554]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="262" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis box="[275,374,1530,1554]" italics="true" pageId="27" pageNumber="262">C. askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[380,467,1530,1555]" pageId="27" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, ventral view.
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</paragraph>
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</caption>
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<subSubSection lastPageId="29" lastPageNumber="264" pageId="28" pageNumber="263" type="discussion">
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<paragraph blockId="28.[159,1036,253,492]" pageId="28" pageNumber="263">
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The two available females from this locality have a face resembling that of
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<taxonomicName authorityName="Vincent J. Kalkman" authorityYear="2016" box="[959,1030,254,277]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="orri">
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<emphasis box="[959,1030,254,277]" italics="true" pageId="28" pageNumber="263">C. orri</emphasis>
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</taxonomicName>
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: labrum and clypeus black, genae and mandibles dark brown to black grading to yellow-brown in the upper posterior corner, front of head tawny-brown with the centre of the frons with mottled dark brown to black in one specimen but largely tawny brown in the other specimen, top of head including sockets of antennae black. This locality is isolated from the main range of
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<taxonomicName authorityName="Vincent J. Kalkman" authorityYear="2016" box="[620,691,407,430]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="orri">
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<emphasis box="[620,691,407,430]" italics="true" pageId="28" pageNumber="263">C. orri</emphasis>
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</taxonomicName>
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and it is not unlikely that these specimens belong to a yet undescribed species. More material, including males, is needed to confirm this.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="28" pageNumber="263" targetBox="[156,1039,522,1404]" targetPageId="28">
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<paragraph blockId="28.[159,1037,1442,1555]" pageId="28" pageNumber="263">
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<emphasis bold="true" box="[159,319,1442,1467]" pageId="28" pageNumber="263">Figures 40–45.</emphasis>
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Appendages of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[497,673,1442,1466]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[497,673,1442,1466]" italics="true" pageId="28" pageNumber="263">Celebargiolestes</emphasis>
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</taxonomicName>
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males. 40 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[813,876,1442,1466]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[813,876,1442,1466]" italics="true" pageId="28" pageNumber="263">C. toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[882,970,1442,1467]" pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, dorsal view; 41 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[411,473,1472,1496]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[411,473,1472,1496]" italics="true" pageId="28" pageNumber="263">C. toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[480,566,1472,1497]" pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, lateral view; 42 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[878,940,1472,1496]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[878,940,1472,1496]" italics="true" pageId="28" pageNumber="263">C. toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[947,1033,1472,1497]" pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, ventral view; 43 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[467,532,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[467,532,1501,1525]" italics="true" pageId="28" pageNumber="263">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[539,625,1501,1526]" pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, dorsal view; 44 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[933,998,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[933,998,1501,1525]" italics="true" pageId="28" pageNumber="263">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, lateral view; 45 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[515,580,1530,1554]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="263" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[515,580,1530,1554]" italics="true" pageId="28" pageNumber="263">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[586,671,1532,1555]" pageId="28" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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, JvT6646, ventral view.
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</paragraph>
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</caption>
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<paragraph blockId="29.[159,1036,253,1034]" box="[533,661,253,281]" pageId="29" pageNumber="264">
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<heading bold="true" box="[533,661,253,281]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="264" reason="0">
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<emphasis bold="true" box="[533,661,253,281]" pageId="29" pageNumber="264">Discussion</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="29.[159,1036,253,1034]" box="[159,621,299,325]" pageId="29" pageNumber="264">
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<heading bold="true" box="[159,621,299,325]" fontSize="11" level="3" pageId="29" pageNumber="264" reason="0">
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<emphasis bold="true" box="[159,621,299,325]" pageId="29" pageNumber="264">Rationale for the taxonomic decisions</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="29.[159,1036,253,1034]" pageId="29" pageNumber="264">
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The species of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[331,510,336,362]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="genus">
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<emphasis box="[331,510,336,362]" italics="true" pageId="29" pageNumber="264">Celebargiolestes</emphasis>
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</taxonomicName>
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are all closely related and show no clear differences except for the pattern on the head, the shape of the head of the genital ligula and the shape of the inferior appendages. Based on these, four groups can easily be recognized. They are here described as species. Within three of these there is variability in the shape of the inferior appendages between localities sometimes coinciding with small variability in coloration. The differences in the shape of the inferior appendages are slight, difficult to judge without direct comparison of material and the use of this character for reliable identification is limited. Material from other locations might show that there is gradual change between regions. For this reason I refrained from giving names and regard them as regional differences within species.
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</paragraph>
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<paragraph blockId="29.[159,1036,253,1034]" pageId="29" pageNumber="264">
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It is remarkable that
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<taxonomicName box="[421,533,748,774]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[421,533,748,774]" italics="true" pageId="29" pageNumber="264">C. cinctus</emphasis>
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</taxonomicName>
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is found in two disjunct areas; however the populations in these disjunct regions share the same pattern on the face and have the same shape of the head of the genital ligula.
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[773,1035,822,848]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis box="[773,1035,822,848]" italics="true" pageId="29" pageNumber="264">Celebargiolestes askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[159,247,860,886]" pageId="29" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
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has the same pattern on the face and shape of the head of the genital ligula as
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[300,377,897,923]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[300,377,897,923]" italics="true" pageId="29" pageNumber="264">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[385,473,897,923]" pageId="29" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
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but differs from the latter by the clearly shorter lower appendages. It could be that
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<taxonomicName authorityName="Vincent J. Kalkman" authorityYear="2016" box="[562,671,934,960]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="species" species="askewi">
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<emphasis box="[562,671,934,960]" italics="true" pageId="29" pageNumber="264">C. askewi</emphasis>
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</taxonomicName>
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represents just one of the more clearly differentiated regional forms of
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<taxonomicName authorityName="Vincent J. Kalkman" authorityYear="2016" box="[616,693,972,998]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="264" phylum="Arthropoda" rank="species" species="orri">
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<emphasis box="[616,693,972,998]" italics="true" pageId="29" pageNumber="264">C. orri</emphasis>
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</taxonomicName>
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which would make the latter paraphyletic.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |