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<mods:title id="9DFB93C4C53380D631D54E7A7DBB0443">Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key</mods:title>
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<mods:affiliation id="295FDDA11199E2255802487ABC6B2CF1">Department of Life Sciences, Natural History Museum, SW 7 5 BD, London, United Kingdom. &amp; Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401, Daugavpils, Latvia &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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<treatment id="038E8D48617DA842FF73FC64F558F827" ID-DOI="http://doi.org/10.5281/zenodo.7415024" ID-GBIF-Taxon="205926627" ID-Zenodo-Dep="7415024" LSID="urn:lsid:plazi:treatment:038E8D48617DA842FF73FC64F558F827" httpUri="http://treatment.plazi.org/id/038E8D48617DA842FF73FC64F558F827" lastPageNumber="122" pageId="1" pageNumber="122">
<subSubSection id="C33D6FD5617DA842FF73FC64F70BFBB9" pageId="1" pageNumber="122" type="nomenclature">
<paragraph id="8B983C5E617DA842FF73FC64F7A2FC64" blockId="1.[151,437,979,1006]" box="[151,437,979,1006]" pageId="1" pageNumber="122">
<heading id="D0D08B32617DA842FF73FC64F7A2FC64" bold="true" box="[151,437,979,1006]" fontSize="11" level="1" pageId="1" pageNumber="122" reason="1">
<taxonomicName id="4C2747DD617DA842FF73FC64F7A2FC64" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[151,437,979,1006]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FF73FC64F7A2FC64" bold="true" box="[151,437,979,1006]" pageId="1" pageNumber="122">
<emphasis id="B953E04C617DA842FF73FC64F72AFC67" bold="true" box="[151,317,979,1006]" italics="true" pageId="1" pageNumber="122">Physciolagria</emphasis>
<bibRefCitation id="EFB641AF617DA842FEA1FC64F7A2FC64" author="Pic, M." box="[325,437,979,1005]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B983C5E617DA842FF73FBAFF70BFBB9" blockId="1.[151,742,1048,1103]" box="[151,284,1048,1072]" pageId="1" pageNumber="122">
<bibRefCitation id="EFB641AF617DA842FF73FBAFF70BFBB9" author="Pic, M." box="[151,284,1048,1072]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic (1930: 7)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD5617DA842FF73FB80F4F1FBC6" box="[151,742,1078,1103]" pageId="1" pageNumber="122" type="type_taxon">
<paragraph id="8B983C5E617DA842FF73FB80F4F1FBC6" blockId="1.[151,742,1048,1103]" box="[151,742,1078,1103]" pageId="1" pageNumber="122">
<typeStatus id="549C82FC617DA842FF73FB80F6DEFBC6" box="[151,201,1079,1103]" pageId="1" pageNumber="122">Type</typeStatus>
species:
<taxonomicName id="4C2747DD617DA842FEC2FB81F478FBC7" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[294,623,1078,1102]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C617DA842FEC2FB81F410FBC7" box="[294,519,1078,1102]" italics="true" pageId="1" pageNumber="122">Physciolagria liturata</emphasis>
<bibRefCitation id="EFB641AF617DA842FDE9FB81F478FBC7" author="Pic, M." box="[525,623,1078,1102]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
, monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD5617DA842FF73FBCAF7A2FA8B" pageId="1" pageNumber="122" type="reference_group">
<paragraph id="8B983C5E617DA842FF73FBCAF7A2FA8B" blockId="1.[151,1437,1148,1967]" pageId="1" pageNumber="122">
<emphasis id="B953E04C617DA842FF73FBCAF755FB1F" bold="true" box="[151,322,1148,1175]" pageId="1" pageNumber="122">Type material</emphasis>
(
<figureCitation id="131C20DB617DA842FEB1FBCBF7A9FB1E" box="[341,446,1148,1175]" captionStart="FIGURES 14" captionStartId="3.[152,270,1959,1983]" captionTargetBox="[164,1406,433,1933]" captionTargetId="figure-116@3.[162,1426,417,1936]" captionTargetPageId="3" captionText="FIGURES 14. Physciolagria liturata Pic, 1930, holotype ♀. 1—Habitus, dorsal view; 2—ditto, lateral view; 3—Forebody, anterodorsal view; 4—Labels [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413350" httpUri="https://zenodo.org/record/7413350/files/figure.png" pageId="1" pageNumber="122">Figs 14</figureCitation>
),
<materialsCitation id="3B4F3603617DA842FE33FBCBF7A5FA8B" ID-GBIF-Occurrence="3988003323" collectingDate="1928-09" collectionCode="BMNH" pageId="1" pageNumber="122" specimenCode="NHMUK014663499" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="549C82FC617DA842FE33FBCBF42EFB1F" box="[471,569,1148,1174]" pageId="1" pageNumber="122" type="holotype">holotype</typeStatus>
<specimenCount id="9D21F7D7617DA842FDA0FBCAF44EFB1E" box="[580,601,1149,1175]" pageId="1" pageNumber="122" type="female"></specimenCount>
[
<collectionCode id="ED36A49B617DA842FD8FFBCAF4DFFB1E" box="[619,712,1149,1175]" pageId="1" pageNumber="122">BMNH</collectionCode>
]:
<collectorName id="26D25988617DA842FD39FBCAF502FB1E" box="[733,789,1149,1175]" pageId="1" pageNumber="122">Type</collectorName>
[printed, label circular, red frame] //
<collectorName id="26D25988617DA842FB35FBCAF33CFB1F" box="[1233,1323,1148,1175]" pageId="1" pageNumber="122">W. Nile</collectorName>
<date id="FF991A9E617DA842FAD2FBCAF38FFB1E" box="[1334,1432,1149,1175]" pageId="1" pageNumber="122" value="1928-09">
<collectingDate id="EFDDE376617DA842FAD2FBCAF38FFB1E" box="[1334,1432,1149,1175]" pageId="1" pageNumber="122" value="1928-09">IX-1928</collectingDate>
</date>
. G.D.H.C. handwritten] //
<collectorName id="26D25988617DA842FE4BFB17F7F1FB33" box="[431,486,1184,1210]" pageId="1" pageNumber="122">Pres.</collectorName>
by
<collectorName id="26D25988617DA842FDE3FB16F51DFB33" box="[519,778,1184,1211]" pageId="1" pageNumber="122">Imp.Inst.Ent. Brit.Mus.</collectorName>
1930298. [printed] //
<collectorName id="26D25988617DA842FBE0FB16F222FB32" box="[1028,1077,1185,1211]" pageId="1" pageNumber="122">Très</collectorName>
désiré [handwritten, underlined] //
<taxonomicName id="4C2747DD617DA842FF54FB73F75FFB57" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[176,328,1220,1246]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus" status="gen. nov.">Physciolagria</taxonomicName>
<taxonomicNameLabel id="A2605D37617DA842FEB5FB72F78FFB56" box="[337,408,1221,1247]" pageId="1" pageNumber="122" rank="genus">n gen.</taxonomicNameLabel>
liturata
<taxonomicNameLabel id="A2605D37617DA842FE1DFB72F427FB56" box="[505,560,1221,1247]" pageId="1" pageNumber="122" rank="species">n sp.</taxonomicNameLabel>
[handwritten] //
<specimenCode id="DB819425617DA842FD17FB73F5CDFB57" box="[755,986,1220,1246]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="122" type="Museum">NHMUK014663499</specimenCode>
[printed, provided with QR code].
<collectorName id="26D25988617DA842FA95FB73F38AFB57" box="[1393,1437,1220,1246]" pageId="1" pageNumber="122">The</collectorName>
type is in good condition
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD5617DA842FF23FABBF558F827" pageId="1" pageNumber="122" type="description">
<paragraph id="8B983C5E617DA842FF23FABBF351F9E2" blockId="1.[151,1437,1148,1967]" pageId="1" pageNumber="122">
<taxonomicName id="4C2747DD617DA842FF23FABBF773FAAF" authorityName="Telnov" authorityYear="2022" box="[199,356,1292,1318]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FF23FABBF773FAAF" box="[199,356,1292,1318]" italics="true" pageId="1" pageNumber="122">Physciolagria</emphasis>
</taxonomicName>
was originally erected to hold one African species,
<taxonomicName id="4C2747DD617DA842FC23FABAF2A3FAAE" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[967,1204,1292,1319]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C617DA842FC23FABAF22DFAAF" box="[967,1082,1292,1318]" italics="true" pageId="1" pageNumber="122">P. liturata</emphasis>
<bibRefCitation id="EFB641AF617DA842FBA0FABBF2A3FAAE" author="Pic, M." box="[1092,1204,1292,1319]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
, on the basis of the following features (translated from Latin and annotated by the author for clarity in accordance with the present situation (as in the studied
<typeStatus id="549C82FC617DA842FE5AFAE2F7F9FAE6" box="[446,494,1365,1391]" pageId="1" pageNumber="122">type</typeStatus>
specimen); authors additions in square brackets): “Body elongated; head short, eyes strong, strongly convex, not emarginate, setose; terminal [maxillary] palpomere elongate, truncate at apex; antennae short, strongly widened, second antennomere short, 3
<superScript id="7C529116617DA842FD05FA2BF4F8FA23" attach="left" box="[737,751,1436,1450]" fontSize="6" pageId="1" pageNumber="122">rd</superScript>
antennomere triangular, distally widened, antennomeres 410 short and strongly transverse, terminal antennomere elongate, rather wide and depressed [possible meant flattened dorso-ventrally], subtruncate at apex; pronotum laterally emarginated, [pronotal hypomeron] impressed at base, [pronotal] base beaded and [slightly] impressed, anterolateral angles obsolete, posterolateral angles nearly rightangled; elytra wide, elongated, epipleuron narrow, posteriorly indistinct; legs slender, tarsi elongated, penultimate tarsomeres lobate, tarsal claws [basally] subdentate; pro- and mesocoxa conical, contiguous” (
<bibRefCitation id="EFB641AF617DA842FB51F9E7F32FF9E2" author="Pic, M." box="[1205,1336,1616,1643]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic 1930: 7</bibRefCitation>
).
</paragraph>
<paragraph id="8B983C5E617DA842FF23F9C3F46AF972" blockId="1.[151,1437,1148,1967]" pageId="1" pageNumber="122">
The new genus was compared with
<taxonomicName id="4C2747DD617DA842FD89F9C3F58EF906" authority="Champion, 1889" authorityName="Champion" authorityYear="1889" box="[621,921,1652,1679]" class="Insecta" family="Mycteridae" genus="Physcius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FD89F9C3F4C6F907" box="[621,721,1652,1678]" italics="true" pageId="1" pageNumber="122">Physcius</emphasis>
Champion, 1889
</taxonomicName>
(now
<taxonomicName id="4C2747DD617DA842FC03F9C3F27CF907" authorityName="Blanchard" authorityYear="1845" box="[999,1131,1652,1678]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Mycteridae</taxonomicName>
:
<taxonomicName id="4C2747DD617DA842FB9DF9C3F2F8F907" authorityName="Thomson" authorityYear="1860" box="[1145,1263,1652,1678]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
, at the time of description in
<taxonomicName id="4C2747DD617DA842FED6F92FF7DCF93B" authorityName="LeConte" authorityYear="1862" box="[306,459,1688,1714]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Lacconotinae">Lacconotinae</taxonomicName>
) but
<bibRefCitation id="EFB641AF617DA842FE1BF92FF466F93A" author="Pic, M." box="[511,625,1688,1715]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic (1930)</bibRefCitation>
informally placed
<taxonomicName id="4C2747DD617DA842FCD9F92FF5CDF93B" authorityName="Telnov" authorityYear="2022" box="[829,986,1688,1714]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FCD9F92FF5CDF93B" box="[829,986,1688,1714]" italics="true" pageId="1" pageNumber="122">Physciolagria</emphasis>
</taxonomicName>
in
<taxonomicName id="4C2747DD617DA842FC1DF92FF273F93B" baseAuthorityName="Borchmann" baseAuthorityYear="1936" box="[1017,1124,1688,1714]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Lagriinae">Lagriinae</taxonomicName>
of
<taxonomicName id="4C2747DD617DA842FB60F92FF333F93B" authorityName="Latreille" authorityYear="1802" box="[1156,1316,1688,1714]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
(
<taxonomicName id="4C2747DD617DA842FACBF92FF38BF93B" baseAuthorityName="Latreille" baseAuthorityYear="1825" box="[1327,1436,1688,1714]" class="Insecta" family="Lagriidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Lagriidae</taxonomicName>
at the time of description) (translated from French): “but seems to be that it needs to be placed inside the
<taxonomicName id="4C2747DD617DA842FAD4F90BF38FF95F" baseAuthorityName="Latreille" baseAuthorityYear="1825" box="[1328,1432,1724,1750]" class="Insecta" family="Lagriidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Lagriidae</taxonomicName>
, rather than the
<taxonomicName id="4C2747DD617DA842FEA4F957F7C3F973" authorityName="LeConte" authorityYear="1862" box="[320,468,1760,1786]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Lacconotinae">Lacconotinae</taxonomicName>
” (
<bibRefCitation id="EFB641AF617DA842FE08F957F478F972" author="Pic, M." box="[492,623,1760,1787]" pageId="1" pageNumber="122" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic 1930: 7</bibRefCitation>
).
</paragraph>
<paragraph id="8B983C5E617DA842FF23F8B3F558F827" blockId="1.[151,1437,1148,1967]" pageId="1" pageNumber="122">
<bibRefCitation id="EFB641AF617DA842FF23F8B3F78CF897" author="Borchmann, F." box="[199,411,1796,1822]" pageId="1" pageNumber="122" pagination="1 - 561" refId="ref8102" refString="Borchmann, F. (1936) Coleoptera Heteromera Fam. Lagriidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 204. Brussels, Louis Desmet-Verteneuil, pp. 1 - 561, 9 pls." type="journal article" year="1936">Borchmann (1936)</bibRefCitation>
listed
<taxonomicName id="4C2747DD617DA842FE01F8B3F495F897" authorityName="Telnov" authorityYear="2022" box="[485,642,1796,1822]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FE01F8B3F495F897" box="[485,642,1796,1822]" italics="true" pageId="1" pageNumber="122">Physciolagria</emphasis>
</taxonomicName>
along other
<taxonomicName id="4C2747DD617DA842FCE9F8B3F56FF897" baseAuthorityName="Borchmann" baseAuthorityYear="1936" box="[781,888,1796,1822]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Lagriinae">Lagriinae</taxonomicName>
(
<taxonomicName id="4C2747DD617DA842FC63F8B3F5E3F897" baseAuthorityName="Latreille" baseAuthorityYear="1825" box="[903,1012,1796,1822]" class="Insecta" family="Lagriidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Lagriidae</taxonomicName>
at the time of the publication) genera, but did not include it in his key to
<taxonomicName id="4C2747DD617DA842FDC6F89FF49AF8CB" baseAuthorityName="Borchmann" baseAuthorityYear="1936" box="[546,653,1832,1858]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Lagriinae">Lagriinae</taxonomicName>
genera, stating that (translated from German, my additions in square brackets) “inadequate description prevented [from] including [it] in the key if [it] belongs in the family at all” (
<bibRefCitation id="EFB641AF617DA842FF7BF8C7F780F802" author="Borchmann, F." box="[159,407,1904,1931]" pageId="1" pageNumber="122" pagination="1 - 561" refId="ref8102" refString="Borchmann, F. (1936) Coleoptera Heteromera Fam. Lagriidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 204. Brussels, Louis Desmet-Verteneuil, pp. 1 - 561, 9 pls." type="journal article" year="1936">Borchmann 1936: 23</bibRefCitation>
).
<taxonomicName id="4C2747DD617DA842FE49F8C7F45DF803" authorityName="Telnov" authorityYear="2022" box="[429,586,1904,1930]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617DA842FE49F8C7F45DF803" box="[429,586,1904,1930]" italics="true" pageId="1" pageNumber="122">Physciolagria</emphasis>
</taxonomicName>
had not been rediscovered and redescribed since that, so
<bibRefCitation id="EFB641AF617DA842FB0FF8C7F6F5F826" author="Bouchard, P. &amp; Bousquet, Y. &amp; Aalbu, R. L. &amp; Alonso-Zarazaga, M. A. &amp; Merkl, O. &amp; Davies, A. E." pageId="1" pageNumber="122" pagination="1 - 633" refId="ref8147" refString="Bouchard, P., Bousquet, Y., Aalbu, R. L., Alonso-Zarazaga, M. A., Merkl, O. &amp; Davies, A. E. (2021) Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera). ZooKeys 1050, 1 - 633. https: // doi. org / 10.3897 / zookeys. 1050.64217" type="journal article" year="2021">
Bouchard
<emphasis id="B953E04C617DA842FA85F8C6F38AF803" box="[1377,1437,1904,1930]" italics="true" pageId="1" pageNumber="122">et al.</emphasis>
(2021)
</bibRefCitation>
listed it among
<emphasis id="B953E04C617DA842FE72F822F426F827" box="[406,561,1940,1966]" italics="true" pageId="1" pageNumber="122">incertae sedis</emphasis>
genera of
<taxonomicName id="4C2747DD617DA842FD43F823F55CF827" authorityName="Latreille" authorityYear="1802" box="[679,843,1940,1966]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>