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<document id="A25C5445D569064338F6FC56675FBE89" ID-DOI="10.5281/zenodo.5170031" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170031" ID-ZooBank="A5348E25-CC3E-476B-9AD2-0A6C3A37A61A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628280835588" checkinUser="felipe" docAuthor="Gordon, Robert D. &amp; Hanley, Guy A." docDate="2017" docId="038E8B10D843FFD7BF96E4A62D86FB71" docLanguage="en" docName="InsectaMundi.0601.1-158.pdf" docOrigin="Insecta Mundi 2017 (601)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Prodilis alison Gordon and Hanley 2017, new species" docType="treatment" docVersion="6" lastPageNumber="86" masterDocId="FFB7F368D814FF80BF56E4652816FFCA" masterDocTitle="South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="86" updateTime="1699039770044" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="1786228E9D9F5F40567A2B97EF493676" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F52836C8C1560DB0047A9179E215FD0C">
<mods:title id="34914B2D994D993E6A2AE314056CABEF">South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)</mods:title>
</mods:titleInfo>
<mods:name id="F5BD125C99C716C4F4FFCB6720AC4E66" type="personal">
<mods:role id="B72C1A877E9B046A486F4395C7F6E15E">
<mods:roleTerm id="A0153EACC917637E134D38267F161F4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A487A6FD7094D4D92B3D60B5742183D">Gordon, Robert D.</mods:namePart>
</mods:name>
<mods:name id="98484356AB9DD899DD94CC28CFE3F309" type="personal">
<mods:role id="AB5760563BD985728F78C8A1CFEC583B">
<mods:roleTerm id="7C9C970FB1D58D8B0BEE7B3C026C743F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D38E6C20A30D675F5FBA707ABE64320A">Hanley, Guy A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1345A263A76382B5B7D52B5CE70F4C07">text</mods:typeOfResource>
<mods:relatedItem id="7F681B1417ACC54E81B8FDD6DF181535" type="host">
<mods:titleInfo id="A8D73718FBE596E38B8B426DBE8268AC">
<mods:title id="D94ADFF85A635BABE639C39A782E01F5">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="3471ED3E2AA01510DE784FCF813D9CE8">
<mods:date id="CC6EB14A277880581D47B342D9D4A00E">2017</mods:date>
<mods:detail id="169816D59DF87362E37B92E70BEAC09E" type="pubDate">
<mods:number id="04F3DEA11F0F67B35F54EE2DD22503AE">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="87C63B27ABC51C7D4E315F789DFDBC1C" type="volume">
<mods:number id="622D3E338CC8F86077E08D5421A0A951">2017</mods:number>
</mods:detail>
<mods:detail id="0BB04ABB87765C05142AA0F2575026E2" type="issue">
<mods:number id="D01D03778BAC66D8D68600633D3371A8">601</mods:number>
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<mods:start id="BD29CFFA436E11BC4B89A535EADDA685">1</mods:start>
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<mods:classification id="A100CF483C0457EC34CA78B785DE51CB">journal article</mods:classification>
<mods:identifier id="C1A925E40041CC005B2BC1E49BCC5E5A" type="DOI">10.5281/zenodo.5170031</mods:identifier>
<mods:identifier id="49B2EF030494615E203D022DF7856F60" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="43E712F9CB0A3046F950EB3576338EEC" type="Zenodo-Dep">5170031</mods:identifier>
<mods:identifier id="E402841AAAAD13C228C6D9D8B184C325" type="ZooBank">A5348E25-CC3E-476B-9AD2-0A6C3A37A61A</mods:identifier>
</mods:mods>
<treatment id="038E8B10D843FFD7BF96E4A62D86FB71" ID-DOI="http://doi.org/10.5281/zenodo.6485184" ID-Zenodo-Dep="6485184" LSID="urn:lsid:plazi:treatment:038E8B10D843FFD7BF96E4A62D86FB71" httpUri="http://treatment.plazi.org/id/038E8B10D843FFD7BF96E4A62D86FB71" lastPageNumber="86" pageId="87" pageNumber="86">
<subSubSection id="C33D698DD843FFD7BF96E4A62B96FF11" box="[192,896,195,219]" pageId="87" pageNumber="86" type="nomenclature">
<paragraph id="8B983A06D843FFD7BF96E4A62B96FF11" blockId="87.[192,896,195,219]" box="[192,896,195,219]" pageId="87" pageNumber="86">
<heading id="D0D08D6AD843FFD7BF96E4A62B96FF11" bold="true" box="[192,896,195,219]" centered="true" fontSize="10" level="2" pageId="87" pageNumber="86" reason="3">
<emphasis id="B953E614D843FFD7BF96E4A62B96FF11" bold="true" box="[192,896,195,219]" pageId="87" pageNumber="86">
41.
<taxonomicName id="4C274185D843FFD7BFB8E4A62AC6FF11" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2017" box="[238,720,195,219]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="86" phylum="Arthropoda" rank="species" species="alison" status="sp. nov.">
<emphasis id="B953E614D843FFD7BFB8E4A629AFFF11" bold="true" box="[238,441,195,219]" italics="true" pageId="87" pageNumber="86">Prodilis alison</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A2605B6FD843FFD7BD88E4A62B96FF11" box="[734,896,195,219]" pageId="87" pageNumber="86" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD843FFD7BF96E5662B00FBD1" pageId="87" pageNumber="86" type="description">
<paragraph id="8B983A06D843FFD7BF96E5662C89FCB1" blockId="87.[192,1441,259,891]" pageId="87" pageNumber="86">
<emphasis id="B953E614D843FFD7BF96E56629AEFED1" bold="true" box="[192,440,259,283]" pageId="87" pageNumber="86">Description. Male</emphasis>
<typeStatus id="549C84A4D843FFD7BE94E5662A39FED1" box="[450,559,259,283]" pageId="87" pageNumber="86" type="holotype">holotype</typeStatus>
. Length
<quantity id="4CDF97E3D843FFD7BDC9E5662AE9FED1" box="[671,767,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="87" pageNumber="86" unit="mm" value="2.2">2.2 mm</quantity>
, width
<quantity id="4CDF97E3D843FFD7BC08E5662BD6FED1" box="[862,960,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="87" pageNumber="86" unit="mm" value="1.5">1.5 mm</quantity>
; body oval, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface shiny except elytron with trace of microsculpture. Color black; elytron with median, oval, yellow macula (
<figureCitation id="131C2683D843FFD7BB66E5262C8FFE91" box="[1072,1177,323,347]" captionStart="Figures 541–558" captionStartId="148.[192,287,1632,1654]" captionTargetBox="[212,1432,211,1586]" captionTargetId="figure-20@148.[198,1445,197,1626]" captionTargetPageId="148" captionText="Figures 541–558. Prodilis spp. 541–546) Prodilis jan 541) Habitus. 542) Basal abdominal ventrites. 543) Frons. 544) Phallobase ventral. 545 Phallobase lateral. 546) Sipho 547–552) Prodilis alison 547) Habitus. 548) Basal abdominal ventrites. 549) Frons. 550) Phallobase ventral. 551) Phallobase lateral. 552) Sipho. 553–558) Prodilis yvette 553) Habitus. 554) Basal abdominal ventrite. 555) Frons. 556) Phallobase ventral. 557) Phallobase lateral. 558) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170093" httpUri="https://zenodo.org/record/5170093/files/figure.png" pageId="87" pageNumber="86">Fig. 547</figureCitation>
); head (
<figureCitation id="131C2683D843FFD7BBACE5262D75FE91" box="[1274,1379,323,347]" captionStart="Figures 541–558" captionStartId="148.[192,287,1632,1654]" captionTargetBox="[212,1432,211,1586]" captionTargetId="figure-20@148.[198,1445,197,1626]" captionTargetPageId="148" captionText="Figures 541–558. Prodilis spp. 541–546) Prodilis jan 541) Habitus. 542) Basal abdominal ventrites. 543) Frons. 544) Phallobase ventral. 545 Phallobase lateral. 546) Sipho 547–552) Prodilis alison 547) Habitus. 548) Basal abdominal ventrites. 549) Frons. 550) Phallobase ventral. 551) Phallobase lateral. 552) Sipho. 553–558) Prodilis yvette 553) Habitus. 554) Basal abdominal ventrite. 555) Frons. 556) Phallobase ventral. 557) Phallobase lateral. 558) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170093" httpUri="https://zenodo.org/record/5170093/files/figure.png" pageId="87" pageNumber="86">Fig. 549</figureCitation>
); antenna, mouthparts, legs yellow. Head punctures small, separated by a diameter or less; pronotal punctures larger than on head, separated by a diameter or less; elytral punctures as large as on pronotum, separated by less than twice a diameter; prosternal punctures small, separated by a diameter or less; mesosternal punctures large, separated by less than a diameter; metasternal punctures basally and laterally as large as on mesosternum, separated by a diameter or less, median punctures small, separated by less than 3 times a diameter; abdomen with punctures on ventrites 1–3 large, separated by a diameter or less, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons densely pubescent, not widened from vertex to clypeus, sides parallel, slightly wider than eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin narrow, equal in width from base to apex. Epipleuron flat, wide in basal ½, as wide as pronotal hypomeron, without feeble depressions for reception of femoral apices. Prosternum longer than wide, longer than mesosternum, base weakly emarginate, lateral carina slender, extended anterior to procoxa. Postcoxal line on ventrite 1 short, rounded, extended slightly beyond middle of ventrite (
<figureCitation id="131C2683D843FFD7BD52E7662A7CFCD1" box="[516,618,771,795]" captionStart="Figures 541–558" captionStartId="148.[192,287,1632,1654]" captionTargetBox="[212,1432,211,1586]" captionTargetId="figure-20@148.[198,1445,197,1626]" captionTargetPageId="148" captionText="Figures 541–558. Prodilis spp. 541–546) Prodilis jan 541) Habitus. 542) Basal abdominal ventrites. 543) Frons. 544) Phallobase ventral. 545 Phallobase lateral. 546) Sipho 547–552) Prodilis alison 547) Habitus. 548) Basal abdominal ventrites. 549) Frons. 550) Phallobase ventral. 551) Phallobase lateral. 552) Sipho. 553–558) Prodilis yvette 553) Habitus. 554) Basal abdominal ventrite. 555) Frons. 556) Phallobase ventral. 557) Phallobase lateral. 558) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170093" httpUri="https://zenodo.org/record/5170093/files/figure.png" pageId="87" pageNumber="86">Fig. 548</figureCitation>
). Apex of ventrite 5 arcuate. Genitalia with phallobase long, slender, basal lobe longer than paramere, widened from base to 3/4, slightly more strongly widened before abruptly rounded apex, apex not emarginate; paramere slender, straight, apex rounded, without marginal serrations (
<figureCitation id="131C2683D843FFD7BE5BE70629B9FCB1" box="[269,431,867,891]" captionStart="Figures 541–558" captionStartId="148.[192,287,1632,1654]" captionTargetBox="[212,1432,211,1586]" captionTargetId="figure-20@148.[198,1445,197,1626]" captionTargetPageId="148" captionText="Figures 541–558. Prodilis spp. 541–546) Prodilis jan 541) Habitus. 542) Basal abdominal ventrites. 543) Frons. 544) Phallobase ventral. 545 Phallobase lateral. 546) Sipho 547–552) Prodilis alison 547) Habitus. 548) Basal abdominal ventrites. 549) Frons. 550) Phallobase ventral. 551) Phallobase lateral. 552) Sipho. 553–558) Prodilis yvette 553) Habitus. 554) Basal abdominal ventrite. 555) Frons. 556) Phallobase ventral. 557) Phallobase lateral. 558) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170093" httpUri="https://zenodo.org/record/5170093/files/figure.png" pageId="87" pageNumber="86">Fig. 550, 551</figureCitation>
); sipho long, slender, apical 1/3 almost filamentous (
<figureCitation id="131C2683D843FFD7BB7CE7062C84FCB1" box="[1066,1170,867,891]" captionStart="Figures 541–558" captionStartId="148.[192,287,1632,1654]" captionTargetBox="[212,1432,211,1586]" captionTargetId="figure-20@148.[198,1445,197,1626]" captionTargetPageId="148" captionText="Figures 541–558. Prodilis spp. 541–546) Prodilis jan 541) Habitus. 542) Basal abdominal ventrites. 543) Frons. 544) Phallobase ventral. 545 Phallobase lateral. 546) Sipho 547–552) Prodilis alison 547) Habitus. 548) Basal abdominal ventrites. 549) Frons. 550) Phallobase ventral. 551) Phallobase lateral. 552) Sipho. 553–558) Prodilis yvette 553) Habitus. 554) Basal abdominal ventrite. 555) Frons. 556) Phallobase ventral. 557) Phallobase lateral. 558) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170093" httpUri="https://zenodo.org/record/5170093/files/figure.png" pageId="87" pageNumber="86">Fig. 552</figureCitation>
).
</paragraph>
<paragraph id="8B983A06D843FFD7BF96E7C62BE4FC11" blockId="87.[192,1440,931,1051]" pageId="87" pageNumber="86">
<emphasis id="B953E614D843FFD7BF96E7C6293FFC71" bold="true" box="[192,297,931,955]" pageId="87" pageNumber="86">Female.</emphasis>
Similar to male except head not densely pubescent. Female genitalia with ramus of spermathecal capsule enlarged, slender to cornu, apex of cornu with forked beak.
</paragraph>
<paragraph id="8B983A06D843FFD7BF96E0662B00FBD1" blockId="87.[192,1440,931,1051]" box="[192,790,1027,1051]" pageId="87" pageNumber="86">
<emphasis id="B953E614D843FFD7BF96E0662954FBD1" bold="true" box="[192,322,1027,1051]" pageId="87" pageNumber="86">Variation</emphasis>
. Length 2.0 to
<quantity id="4CDF97E3D843FFD7BEAAE0662A4CFBD1" box="[508,602,1027,1051]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="87" pageNumber="86" unit="mm" value="2.2">2.2 mm</quantity>
, width
<quantity id="4CDF97E3D843FFD7BDE5E0662B04FBD1" box="[691,786,1027,1051]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="87" pageNumber="86" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD843FFD7BF96E0262C00FBB1" pageId="87" pageNumber="86" type="materials_examined">
<paragraph id="8B983A06D843FFD7BF96E0262C00FBB1" blockId="87.[192,1440,1091,1211]" pageId="87" pageNumber="86">
<emphasis id="B953E614D843FFD7BF96E026299EFB91" bold="true" box="[192,392,1091,1115]" pageId="87" pageNumber="86">Type material.</emphasis>
<materialsCitation id="3B4F305BD843FFD7BED8E0262C4DFB91" box="[398,1115,1091,1115]" collectionCode="CMNH" country="Brazil" location="Santarem" pageId="87" pageNumber="86" specimenCount="1" typeStatus="holotype">
<typeStatus id="549C84A4D843FFD7BED8E02629EBFB91" box="[398,509,1091,1115]" pageId="87" pageNumber="86" type="holotype">Holotype</typeStatus>
male;
<collectingCountry id="F3307A96D843FFD7BD1DE0262AA3FB91" box="[587,693,1091,1115]" name="Brazil" pageId="87" pageNumber="86">BRAZIL</collectingCountry>
:
<location id="8EF86CDDD843FFD7BD92E0262B29FB91" LSID="urn:lsid:plazi:treatment:038E8B10D843FFD7BF96E4A62D86FB71:8EF86CDDD843FFD7BD92E0262B29FB91" box="[708,831,1091,1115]" country="Brazil" name="Santarem" pageId="87" pageNumber="86">Santarem</location>
, AccNo. 2966. (
<collectionCode id="ED36A2C3D843FFD7BCA2E0262C44FB91" box="[1012,1106,1091,1115]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="87" pageNumber="86" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F305BD843FFD7BB32E0212988FBB1" collectionCode="CMNH" country="Brazil" location="Santarem" pageId="87" pageNumber="86" specimenCount="1" typeStatus="paratype">
<typeStatus id="549C84A4D843FFD7BB32E0212CF7FB96" box="[1124,1249,1092,1116]" pageId="87" pageNumber="86" type="paratype">Paratypes</typeStatus>
; 1, same data as holotype (
<collectionCode id="ED36A2C3D843FFD7BE6EE0062980FBB1" box="[312,406,1123,1147]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="87" pageNumber="86" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F305BD843FFD7BEFCE0062C04FBB1" box="[426,1042,1123,1148]" collectionCode="BMNH" collectorName="Cl. Fry" country="Brazil" location="Brazil" pageId="87" pageNumber="86" specimenCount="1" stateProvince="Pernambuco" typeStatus="paratype">
2,
<collectorName id="26D25FD0D843FFD7BE9EE0062A0AFBB6" box="[456,540,1123,1148]" pageId="87" pageNumber="86">Cl. Fry</collectorName>
, Pernam. (
<collectingRegion id="49E3F4E4D843FFD7BDC9E0062B2BFBB1" box="[671,829,1123,1147]" country="Brazil" name="Pernambuco" pageId="87" pageNumber="86">Pernambuco</collectingRegion>
,
<location id="8EF86CDDD843FFD7BC1EE0062B82FBB1" LSID="urn:lsid:plazi:treatment:038E8B10D843FFD7BF96E4A62D86FB71:8EF86CDDD843FFD7BC1EE0062B82FBB1" box="[840,916,1123,1147]" country="Brazil" name="Brazil" pageId="87" pageNumber="86" stateProvince="Pernambuco">Brazil</location>
) (
<collectionCode id="ED36A2C3D843FFD7BCFDE0062C1FFBB1" box="[939,1033,1123,1147]" pageId="87" pageNumber="86">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD843FFD7BF96E0C62D86FB71" box="[192,1424,1187,1211]" pageId="87" pageNumber="86" type="discussion">
<paragraph id="8B983A06D843FFD7BF96E0C62D86FB71" blockId="87.[192,1440,1091,1211]" box="[192,1424,1187,1211]" pageId="87" pageNumber="86">
<emphasis id="B953E614D843FFD7BF96E0C62952FB71" bold="true" box="[192,324,1187,1211]" pageId="87" pageNumber="86">Remarks.</emphasis>
This species has a distinctive elytral color pattern as well as characteristic male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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