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<document id="EC5D7AF3CC3EAC81529CAB8054FD1354" ID-DOI="10.5281/zenodo.185015" ID-GBIF-Dataset="0915c7c5-1af7-49ff-92f5-c7b11fbbb440" ID-ISSN="1175-5326" ID-Zenodo-Dep="185015" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459948681130" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2009" docId="03867469FFF1FFB2FF40954098F1A296" docLanguage="en" docName="zt01970p068.pdf" docOrigin="Zootaxa 1970" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crossopriza lyoni Blackwall 1867" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFBF0C11FFF0FFB3FFD794629B2DA52F" masterDocTitle="Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae)" masterLastPageNumber="68" masterPageNumber="64" pageNumber="65" updateTime="1698230759048" updateUser="plazi">
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<mods:title id="68D97ABC138110DFAA6EE4E15BAB2A7B">Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae)</mods:title>
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<mods:namePart id="D838E9BC4C60C07FB7446C888A13CE0C">Huber, Bernhard A.</mods:namePart>
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<mods:date id="BFF844D63A14689E894D01B979E6090F">2009</mods:date>
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<treatment id="03867469FFF1FFB2FF40954098F1A296" ID-DOI="http://doi.org/10.5281/zenodo.6213098" ID-GBIF-Taxon="119376403" ID-Zenodo-Dep="6213098" LSID="urn:lsid:plazi:treatment:03867469FFF1FFB2FF40954098F1A296" httpUri="http://treatment.plazi.org/id/03867469FFF1FFB2FF40954098F1A296" lastPageNumber="65" pageId="1" pageNumber="65">
<subSubSection id="C33596F4FFF1FFB2FF409540990CA415" box="[151,545,290,314]" pageId="1" pageNumber="65" type="nomenclature">
<paragraph id="8B90C57FFFF1FFB2FF409540990CA415" blockId="1.[151,545,290,314]" box="[151,545,290,314]" pageId="1" pageNumber="65">
<heading id="D0D87213FFF1FFB2FF409540990CA415" bold="true" box="[151,545,290,314]" fontSize="10" level="2" pageId="1" pageNumber="65" reason="2">
<taxonomicName id="4C2FBEFCFFF1FFB2FF409540990CA415" ID-CoL="ZMLK" authority="Blackwall, 1867" authorityName="Blackwall" authorityYear="1867" box="[151,545,290,314]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FF409540990CA415" bold="true" box="[151,545,290,314]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF4095409A7AA415" bold="true" box="[151,343,290,314]" italics="true" pageId="1" pageNumber="65">Crossopriza lyoni</emphasis>
(Blackwall, 1867)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33596F4FFF1FFB2FF40950A989BA728" pageId="1" pageNumber="65" type="reference_group">
<paragraph id="8B90C57FFFF1FFB2FF40950A988CA4ED" blockId="1.[151,991,359,519]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF40950A9A67A452" box="[151,330,360,381]" italics="true" pageId="1" pageNumber="65">
Tibiosa
<taxonomicName id="4C2FBEFCFFF1FFB2FF33950A9A67A452" authority="Gonzalez-Sponga, 2006: 11" authorityName="Gonzalez-Sponga" authorityPageNumber="11" authorityYear="2006" box="[228,330,360,381]" pageId="1" pageNumber="65" rank="species" species="caracensis">caracensis</taxonomicName>
</emphasis>
González-Sponga, 2006: 1113; pl. 1, figs. 110; new synonymy.
<emphasis id="B95B196DFFF1FFB2FF4095E89A5DA4B0" box="[151,368,394,415]" italics="true" pageId="1" pageNumber="65">Tibiosa casanaimensis</emphasis>
González-Sponga, 2006: 1416; pl. 2, figs. 19; new synonymy.
<emphasis id="B95B196DFFF1FFB2FF4095CF9A1DA4ED" box="[151,304,429,450]" italics="true" pageId="1" pageNumber="65">Tibiosa coreana</emphasis>
González-Sponga, 2006: 1719; pl. 3, figs. 19; new synonymy.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF4095B2989BA728" blockId="1.[151,991,359,519]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF4095B29A67A4CA" box="[151,330,464,485]" italics="true" pageId="1" pageNumber="65">Tibiosa guayanesa</emphasis>
González-Sponga, 2006: 2022; pl. 4, figs. 19; new synonymy.
<emphasis id="B95B196DFFF1FFB2FF4095909A6BA728" box="[151,326,498,519]" italics="true" pageId="1" pageNumber="65">Tibiosa moraensis</emphasis>
González-Sponga, 2006: 2325; pl. 5, figs. 19; new synonymy.
</paragraph>
</subSubSection>
<subSubSection id="C33596F4FFF1FFB2FF40965A98F1A296" pageId="1" pageNumber="65" type="description">
<paragraph id="8B90C57FFFF1FFB2FF40965A9E54A75E" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF40965A9A94A760" box="[151,441,568,591]" italics="true" pageId="1" pageNumber="65">Justification of synonymies</emphasis>
. Three lines of evidence strongly support the synonymies: the characters used by González-
<bibRefCitation id="EFBEB88EFFF1FFB2FF4096389A19A75E" author="Sponga" box="[151,308,602,625]" pageId="1" pageNumber="68" refString="Gonzalez - Sponga, M. A. (2006) Aracnidos de Venezuela. Un nuevo genero y cinco nuevas especies de la familia Pholcidae (Araneae). Sapiens, Revista Universitaria de Investigacion, 7 (1), 9 - 27." type="journal article" year="2006">Sponga (2006)</bibRefCitation>
to diagnose the species, the biogeography of
<taxonomicName id="4C2FBEFCFFF1FFB2FCCC963898B1A75E" box="[795,924,602,625]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B95B196DFFF1FFB2FCCC963898B1A75E" box="[795,924,602,625]" italics="true" pageId="1" pageNumber="65">Crossopriza</emphasis>
</taxonomicName>
, and the microhabitats of the “new” species.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF11961F9F13A64B" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF11961F9A13A7BB" box="[198,318,637,660]" italics="true" pageId="1" pageNumber="65">Characters</emphasis>
.
<collectingCountry id="F33885EFFFF1FFB2FE99961F9AA8A7BB" box="[334,389,637,660]" name="Malta" pageId="1" pageNumber="65">Male</collectingCountry>
palps, male chelicerae, and the female epigynum are the most important characters to diagnose species in
<taxonomicName id="4C2FBEFCFFF1FFB2FED196C29A5EA798" box="[262,371,672,695]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
. Most of González-Spongas (2006) drawings of these characters are fairly good and clearly indicative of
<taxonomicName id="4C2FBEFCFFF1FFB2FF3296A09A17A7F6" box="[229,314,706,729]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FF3296A09A17A7F6" box="[229,314,706,729]" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
(for drawings, see
<bibRefCitation id="EFBEB88EFFF1FFB2FDDA96A099A2A7F6" author="Millot" box="[525,655,706,729]" pageId="1" pageNumber="68" refString="Millot, J. (1946) Les pholcides de Madagascar (Araneides). Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie, 22 (3), 127 - 158." type="journal article" year="1946">Millot 1946</bibRefCitation>
,
<bibRefCitation id="EFBEB88EFFF1FFB2FD4C96A0984DA7F6" author="Beatty" box="[667,864,706,729]" pageId="1" pageNumber="68" refString="Beatty, J. A., Berry J. W. &amp; Huber, B. A. (2008) The pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae). Journal of Arachnology, 36, 1 - 25." type="journal article" year="2008">Beatty et al. 2008</bibRefCitation>
). However, in only one palp is the bulb in its natural position (plate 3); in the others it is slightly to strongly (plates 1, 4) rotated which produces significantly different views. By manipulating a bulb under a dissecting microscope, all these views can easily be reproduced. All drawings of male chelicerae clearly show the two pairs of very distinctive apophyses that characterize
<taxonomicName id="4C2FBEFCFFF1FFB2FBC097489F41A66E" box="[1047,1132,810,833]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FBC097489F41A66E" box="[1047,1132,810,833]" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
but do not occur in this conformation in any other described or undescribed species seen by the author (see below).
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF119712984BA11B" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
Two of the epigyna in González-Sponga (2006, plates 1, 5) are fairly good representations of what is seen in adult
<taxonomicName id="4C2FBEFCFFF1FFB2FA5397129BE0A686" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FA5397129BE0A686" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
females. Plate 2 probably shows a male genital area. The anterior structures shown are light spots in the otherwise black area in front of the epigastric furrow; they occur in variable shape and size in juveniles and adults of both sexes. The posterior structure probably represents the small brown plate in front of the male gonopore. The “epigynum” in plate 4 is probably from a juvenile specimen. In plate 3, figs. 8 and 9 are almost certainly from
<taxonomicName id="4C2FBEFCFFF1FFB2FB9897989FDDA13E" box="[1103,1264,1018,1041]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="atlanta">
<emphasis id="B95B196DFFF1FFB2FB9897989FDDA13E" box="[1103,1264,1018,1041]" italics="true" pageId="1" pageNumber="65">Artema atlanta</emphasis>
</taxonomicName>
. Both the round abdomen and the distinctive epigynum strongly hint at that species.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF119022999FA1B3" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">Most drawings of palpal endites are appropriate and show the usual variation (both true and artificial) seen in these—in pholcids otherwise taxonomically barely informative—structures. On plate 2, the endites are probably covered by a piece of dirt or an extrusion of internal fluids.</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF1190CA9BD8A008" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">Carapace shape is rarely a distinctive character within pholcid genera, and all the variation shown in González- Spongas (2006) drawings can easily be attributed to small differences in the angle of view and to slight deformations of the soft lateral margins. Differences in the angle of view also explain the minimal differences shown in the positions of the eyes.</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF1191509A85A0FB" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
The sternum in
<taxonomicName id="4C2FBEFCFFF1FFB2FEA791519903A065" box="[368,558,1331,1354]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FEA791519903A065" box="[368,558,1331,1354]" italics="true" pageId="1" pageNumber="65">Crossopriza lyoni</emphasis>
</taxonomicName>
has a straight posterior border between the joints of the fourth leg coxae. The lateral extensions shown by González-Sponga (2006) are in fact extensions of the carapace that reach ventrally on both sides of the petiole and almost contact the sternum. The median extension is in fact an unpaired sclerite on the petiolus. The borders of all these sclerites are often poorly visible due to underlying patterns of dark pigment in the soft tissue directly under the cuticle.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF1191829BF3A3AE" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
Finally, size measures are a difficult character to separate species. Except for one species, all male tibia 1 measures (
<quantity id="4CD7689AFFF1FFB2FF4892609A16A336" box="[159,315,1538,1561]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.58" metricValueMin="1.01" pageId="1" pageNumber="65" unit="mm" value="12.95" valueMax="15.8" valueMin="10.1">10.115.8 mm</quantity>
) fall within the range of
<taxonomicName id="4C2FBEFCFFF1FFB2FD9F926099B0A336" box="[584,669,1538,1561]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FD9F926099B0A336" box="[584,669,1538,1561]" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
. The only exception (table 2) is clearly an error: in this male, leg 1 has the shortest tibia of all legs, while in fact it should have the longest (like the female and like all other “species”). In at least two cases (tables 2, 3, male legs 1) leg length calculations are wrong, further eroding confidence in these measurements.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF1192EF9F40A27E" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF1192EF9A76A38B" box="[198,347,1677,1700]" italics="true" pageId="1" pageNumber="65">Biogeography</emphasis>
.
<taxonomicName id="4C2FBEFCFFF1FFB2FEBA92EF9902A38B" box="[365,559,1677,1700]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FEBA92EF9902A38B" box="[365,559,1677,1700]" italics="true" pageId="1" pageNumber="65">Crossopriza lyoni</emphasis>
</taxonomicName>
is a cosmotropical spider that has been collected in many countries including North, Central, and South
<collectingCountry id="F33885EFFFF1FFB2FE6592D2993FA3E8" box="[434,530,1712,1735]" name="United States of America" pageId="1" pageNumber="65">America</collectingCountry>
. I have seen specimens from many countries, including
<collectingCountry id="F33885EFFFF1FFB2FBBB92D29FF4A3E8" box="[1132,1241,1712,1735]" name="Argentina" pageId="1" pageNumber="65">Argentina</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FB3292D29E64A3E8" box="[1253,1353,1712,1735]" name="Paraguay" pageId="1" pageNumber="65">Paraguay</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FA8392D29EB5A3E8" box="[1364,1432,1712,1735]" name="Brazil" pageId="1" pageNumber="65">Brazil</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FF4092B09A2AA3C6" box="[151,263,1746,1769]" name="Nicaragua" pageId="1" pageNumber="65">Nicaragua</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FEC592B09A6AA3C6" box="[274,327,1746,1769]" name="United States of America" pageId="1" pageNumber="65">USA</collectingCountry>
(
<bibRefCitation id="EFBEB88EFFF1FFB2FE8292B0993EA3C6" author="Huber" box="[341,531,1746,1769]" pageId="1" pageNumber="68" refString="Huber, B. A., Deeleman - Reinhold, C. L. &amp; Perez G., A. (1999) The spider genus Crossopriza (Araneae, Pholcidae) in the New World. American Museum Novitates, 3262, 1 - 10." type="journal article" year="1999">Huber et al. 1999</bibRefCitation>
),
<collectingCountry id="F33885EFFFF1FFB2FDF192B099B9A3C6" box="[550,660,1746,1769]" name="Venezuela" pageId="1" pageNumber="65">Venezuela</collectingCountry>
(listed in Colmenares-García 2008), and
<collectingCountry id="F33885EFFFF1FFB2FB9992B09FA9A3C6" box="[1102,1156,1746,1769]" name="Haiti" pageId="1" pageNumber="65">Haiti</collectingCountry>
(unpublished). Except for
<taxonomicName id="4C2FBEFCFFF1FFB2FF4092979BC6A223" box="[151,235,1781,1804]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FF4092979BC6A223" box="[151,235,1781,1804]" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2FBEFCFFF1FFB2FF2E92979A56A223" box="[249,379,1781,1804]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B95B196DFFF1FFB2FF2E92979A56A223" box="[249,379,1781,1804]" italics="true" pageId="1" pageNumber="65">Crossopriza</emphasis>
</taxonomicName>
is restricted to Africa north of the Equator and the Arabian Peninsula. I have seen congeneric species (described and undescribed) from
<collectingCountry id="F33885EFFFF1FFB2FDE1937A99ABA200" box="[566,646,1816,1839]" name="Guinea-Bissau" pageId="1" pageNumber="65">Guinea</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FD46937A983CA200" box="[657,785,1816,1839]" name="Ivory Coast" pageId="1" pageNumber="65">Ivory Coast</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FCCB937A9883A200" box="[796,942,1816,1839]" name="Burkina Faso" pageId="1" pageNumber="65">Burkina Faso</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FC6E937A9F2CA200" box="[953,1025,1816,1839]" name="Ghana" pageId="1" pageNumber="65">Ghana</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FBDB937A9FFFA200" box="[1036,1234,1816,1839]" name="Equatorial Guinea" pageId="1" pageNumber="65">Equatorial Guinea</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FB0B937A9E3AA200" box="[1244,1303,1816,1839]" name="Chad" pageId="1" pageNumber="65">Chad</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FAF5937A9EB5A200" box="[1314,1432,1816,1839]" name="Democratic Republic of the Congo" pageId="1" pageNumber="65">Congo DR</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FF4093589BF1A27E" box="[151,220,1850,1873]" name="Sudan" pageId="1" pageNumber="65">Sudan</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FF3093589A69A27E" box="[231,324,1850,1873]" name="Ethiopia" pageId="1" pageNumber="65">Ethiopia</collectingCountry>
,
<collectingCountry id="F33885EFFFF1FFB2FE9993589AB6A27E" box="[334,411,1850,1873]" name="Yemen" pageId="1" pageNumber="65">Yemen</collectingCountry>
, and
<collectingCountry id="F33885EFFFF1FFB2FE049358993AA27E" box="[467,535,1850,1873]" name="Oman" pageId="1" pageNumber="65">Oman</collectingCountry>
. A taxonomic revision of this material is in preparation.
</paragraph>
<paragraph id="8B90C57FFFF1FFB2FF11933F98F1A296" blockId="1.[151,1437,568,1977]" pageId="1" pageNumber="65">
<emphasis id="B95B196DFFF1FFB2FF11933F9A7FA25B" box="[198,338,1885,1908]" italics="true" pageId="1" pageNumber="65">Microhabitat</emphasis>
. Four of González-Spongas (2006) “new species” where collected in human buildings. One was additionally found under waist, one on the walls of a dry ravine. For the fifth species, no data on microhabitat are given.
<collectingCountry id="F33885EFFFF1FFB2FF4093C09BC8A296" box="[151,229,1954,1977]" name="Oman" pageId="1" pageNumber="65">Human</collectingCountry>
constructions and their surroundings are the typical habitat of
<taxonomicName id="4C2FBEFCFFF1FFB2FC5693C098F8A296" box="[897,981,1954,1977]" class="Arachnida" family="Pholcidae" genus="Crossopriza" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="lyoni">
<emphasis id="B95B196DFFF1FFB2FC5693C098F8A296" box="[897,981,1954,1977]" italics="true" pageId="1" pageNumber="65">C. lyoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>