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<document id="BCE3AA67E6E9AE1D3D2B7234CFFB0FB2" ID-DOI="10.11646/zootaxa.3889.3.3" ID-GBIF-Dataset="0f472ebf-c9c3-441b-96cc-510a76de5b4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="224919" ID-ZooBank="F5294390-C1C6-4011-89F7-76BBCE641919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989340508" checkinUser="plazi" docAuthor="Lewis, John G. E." docDate="2014" docId="C95387E94066F14EB8D70ABEC422FCA2" docLanguage="en" docName="zt03889p413.pdf" docOrigin="Zootaxa 3889 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus ruficeps Pocock 1890" docType="treatment" docVersion="8" lastPageNumber="406" masterDocId="356AFF914074F15CB8400A29C734FFBF" masterDocTitle="A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)" masterLastPageNumber="413" masterPageNumber="388" pageNumber="406" updateTime="1698561154455" updateUser="plazi">
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<mods:title id="FC4EBF1A83A0270579C074948A5D0FBA">A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="B142C579FE2C5A3740DD8092AD499E14">Lewis, John G. E.</mods:namePart>
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<mods:date id="718112DA086E6A5EF89285E0A9ECA7B9">2014</mods:date>
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<treatment id="C95387E94066F14EB8D70ABEC422FCA2" ID-DOI="http://doi.org/10.5281/zenodo.5661031" ID-GBIF-Taxon="119596227" ID-Zenodo-Dep="5661031" LSID="urn:lsid:plazi:treatment:C95387E94066F14EB8D70ABEC422FCA2" httpUri="http://treatment.plazi.org/id/C95387E94066F14EB8D70ABEC422FCA2" lastPageNumber="406" pageId="18" pageNumber="406">
<subSubSection id="09E065744066F14EB8D70ABEC507FF0D" box="[151,563,151,178]" pageId="18" pageNumber="406" type="nomenclature">
<paragraph id="414536FF4066F14EB8D70ABEC507FF0D" blockId="18.[151,563,151,178]" box="[151,563,151,178]" pageId="18" pageNumber="406">
<heading id="1A0D81934066F14EB8D70ABEC507FF0D" bold="true" box="[151,563,151,178]" fontSize="11" level="1" pageId="18" pageNumber="406" reason="1">
<taxonomicName id="86FA4D7C4066F14EB8D70ABEC507FF0D" ID-CoL="4B7ZC" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[151,563,151,178]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="738EEAED4066F14EB8D70ABEC507FF0D" bold="true" box="[151,563,151,178]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8D70ABEC6BDFF0E" bold="true" box="[151,393,151,177]" italics="true" pageId="18" pageNumber="406">Otostigmus ruficeps</emphasis>
<bibRefCitation id="256B4B0E4066F14EB9D00AB1C507FF0D" author="Pocock" box="[400,563,152,178]" pageId="18" pageNumber="406" refString="Pocock, R. I. (1890) Report on a small collection of scorpions and centipedes sent from Madras by Mr Edgar Thurston. Annals and Magazine of Natural History, 5 (6), 236 - 250. [+ plate 4]" type="journal article" year="1890">Pocock, 1890</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14EB8D70AF6C224FED0" pageId="18" pageNumber="406" type="reference_group">
<paragraph id="414536FF4066F14EB8D70AF6C532FEAC" blockId="18.[151,1296,222,367]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D70AF6C6DFFF4B" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[151,491,222,244]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="738EEAED4066F14EB8D70AF6C66DFF4B" box="[151,345,222,244]" italics="true" pageId="18" pageNumber="406">Otostigma ruficeps</emphasis>
<bibRefCitation id="256B4B0E4066F14EB91E0AF7C6DFFF4B" author="Pocock" box="[350,491,222,244]" pageId="18" pageNumber="406" refString="Pocock, R. I. (1890) Report on a small collection of scorpions and centipedes sent from Madras by Mr Edgar Thurston. Annals and Magazine of Natural History, 5 (6), 236 - 250. [+ plate 4]" type="journal article" year="1890">Pocock, 1890</bibRefCitation>
</taxonomicName>
, 247, Plate 12,
<figureCitation id="D9C12A7A4066F14EBAC60AF7C5C7FF4B" box="[646,755,222,244]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1673,1696]" captionTargetBox="[175,1413,204,1644]" captionTargetId="figure@3.[164,1423,193,1652]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Otostigmus ateles Holotype. 1. Antennal articles 1 – 3 dorsal. 2. Forcipular coxosternal tooth-plates. 3. Sternite 17. 4. Ultimate segments ventral. FIGURES 5 – 11. Otostigmus australianus 5. Sternite 12 Lectotype. 6. Sternite of ultimate leg-bearing segment spm. 2 Belenden Ker (after Lewis, 2003). 7. Sternite of ultimate leg-bearing segment spm. 3 Belenden Ker (after Lewis, 2003). 8. Right forcipular trochanteroprefemoral process and tooth-plates spm. 7 Malanda. 9. Sternite 6 spm. 7 Malanda. 10. Sternite 12 spm. 7 Malanda after (Lewis, 2003). 11. Ultimate segments ventral spm. 7 Malanda. Scale bars = 1.0 mm except Figs 1 &amp; 2 = 0.5 mm." httpUri="https://zenodo.org/record/224920/files/figure.png" pageId="18" pageNumber="406">Figs 2 &amp; 2</figureCitation>
b. Madras,
<collectingCountry id="39ED766F4066F14EBB240AF7C4AFFF4B" box="[868,923,222,244]" name="India" pageId="18" pageNumber="406">India</collectingCountry>
.
<taxonomicName id="86FA4D7C4066F14EB8D70AD7C6FBFEAC" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[151,463,253,275]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps" subGenus="Otostigmus">
<emphasis id="738EEAED4066F14EB8D70AD7C784FEAC" box="[151,176,254,275]" italics="true" pageId="18" pageNumber="406">O.</emphasis>
(
<emphasis id="738EEAED4066F14EB8FE0AD7C7E4FEAC" box="[190,208,254,275]" italics="true" pageId="18" pageNumber="406">O</emphasis>
.)
<emphasis id="738EEAED4066F14EB8A50AD4C603FEAC" box="[229,311,253,275]" italics="true" pageId="18" pageNumber="406">ruficeps</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9040AD4C6FBFEAC" author="Attems" box="[324,463,253,275]" pageId="18" pageNumber="406" refString="Attems, C. (1930) Scolopendromorpha, Das Tierreich, 54, Berlin: Walter de Gruyter, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
</taxonomicName>
, 141.
</paragraph>
<paragraph id="414536FF4066F14EB8D70B35C6E8FE8D" blockId="18.[151,1296,222,367]" box="[151,476,284,306]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D70B35C691FE8D" authority="Khanna, 1994" authorityName="Khanna" authorityYear="1994" box="[151,421,284,306]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="738EEAED4066F14EB8D70B35C63CFE8D" box="[151,264,284,306]" italics="true" pageId="18" pageNumber="406">O. ruficeps</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9540B35C691FE8D" author="Khanna" box="[276,421,284,306]" pageId="18" pageNumber="406" refString="Khanna, V. (1994) Centipede fauna of NE States, India (Chilopoda Scolopendromorpha), Record of the Zoological Survey of India, 94, 459 - 472." type="journal article" year="1994">Khanna, 1994</bibRefCitation>
</taxonomicName>
, 466.
</paragraph>
<paragraph id="414536FF4066F14EB8D70B12C224FED0" blockId="18.[151,1296,222,367]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D70B12C6F0FEEF" authority="Lewis, 1996" authorityName="Lewis" authorityYear="1996" box="[151,452,314,336]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps" subGenus="Otostigmus">
<emphasis id="738EEAED4066F14EB8D70B12C784FEEF" box="[151,176,315,336]" italics="true" pageId="18" pageNumber="406">O.</emphasis>
(
<emphasis id="738EEAED4066F14EB8FE0B12C7E4FEEF" box="[190,208,315,336]" italics="true" pageId="18" pageNumber="406">O</emphasis>
.)
<emphasis id="738EEAED4066F14EB8A50B13C603FEEF" box="[229,311,314,336]" italics="true" pageId="18" pageNumber="406">ruficeps</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9040B13C6F0FEEF" author="Lewis" box="[324,452,314,336]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (1996) The type specimens and identity of the species described in the genus Otostigmus (subgenus Otostigmus) by R. I. Pocock in the collection of the Natural History Museum, London (Scolopendromorpha: Scolopendridae). Journal of Natural History, 30, 823 - 834. http: // dx. doi. org / 10.1080 / 00222939600770451" type="journal article" year="1996">Lewis, 1996</bibRefCitation>
</taxonomicName>
, 828,
<figureCitation id="D9C12A7A4066F14EBA410B13C57AFEEF" box="[513,590,314,336]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="18" pageNumber="406">Figs 20</figureCitation>
–23. (Incorrectly synonymised under
<taxonomicName id="86FA4D7C4066F14EBB880B12C37DFEEF" ID-CoL="4B7VZ" box="[968,1097,314,336]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="738EEAED4066F14EBB880B12C37DFEEF" box="[968,1097,314,336]" italics="true" pageId="18" pageNumber="406">O. multidens</emphasis>
</taxonomicName>
).
<taxonomicName id="86FA4D7C4066F14EB8D70B73C528FED0" authority="Joshi &amp; Edgecombe 2013" authorityName="Joshi &amp; Edgecombe" authorityYear="2013" box="[151,540,345,367]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="738EEAED4066F14EB8D70B73C63CFED0" box="[151,264,345,367]" italics="true" pageId="18" pageNumber="406">O. ruficeps</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9540B70C528FED0" author="Joshi" box="[276,540,345,367]" pageId="18" pageNumber="406" refString="Joshi, J. &amp; Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. http: // dx. doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi &amp; Edgecombe 2013</bibRefCitation>
</taxonomicName>
, 119, Figs 68–77 (revised species diagnosis). Karnataka and Kerala,
<collectingCountry id="39ED766F4066F14EBC940B70C23FFED0" box="[1236,1291,345,367]" name="India" pageId="18" pageNumber="406">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14EB8D70BB5C4C7FD32" pageId="18" pageNumber="406" type="diagnosis">
<paragraph id="414536FF4066F14EB8D70BB5C4C7FD32" blockId="18.[151,1437,412,797]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8D70BB5C625FE0A" bold="true" box="[151,273,412,437]" pageId="18" pageNumber="406">Diagnosis.</emphasis>
(From
<bibRefCitation id="256B4B0E4066F14EB92F0BB5C591FE0B" author="Joshi" box="[367,677,412,437]" pageId="18" pageNumber="406" refString="Joshi, J. &amp; Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. http: // dx. doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi &amp; Edgecombe 2013</bibRefCitation>
).
<taxonomicName id="86FA4D7C4066F14EBAFC0BB5C408FE0C" box="[700,828,412,435]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="738EEAED4066F14EBAFC0BB5C408FE0C" box="[700,828,412,435]" italics="true" pageId="18" pageNumber="406">Otostigmus</emphasis>
</taxonomicName>
with 21 elongate antennal articles, the basal 2–2.5 glabrous dorsally; forcipular tooth-plates with 4+4 large teeth, small (fifth) inner tooth variably present; four teeth along margin of forcipular trochanteroprefemoral process; longitudinal median depression on posterior half of tergite of ultimate leg-bearing segment; two apical spines on coxopleural process, dorsal spine rarely present; lateral spine mostly lacking; two tarsal spurs on legs
<date id="3544103F4066F14EBAB90805C4B7FDFB" box="[761,899,556,580]" pageId="18" pageNumber="406" value="1919-01-04" valueMax="1919-01-01" valueMin="1919-01-04">1–4 to 1–19</date>
(usually
<date id="3544103F4066F14EBBAF0805C3BCFDFB" box="[1007,1160,556,580]" pageId="18" pageNumber="406" value="1919-01-17" valueMax="1919-01-01" valueMin="1919-01-17">1–17 to 1–19</date>
), one on leg 20; a tibial spur typically on legs 1–6 to 1–8; ultimate leg prefemur with 2 VM spines and 2–3 VL spines. (The leg shown in Joshi &amp; Edgecombe’s Fig. 74 has three rows each of two prefemoral spines).
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14EB88708BEC422FCA2" pageId="18" pageNumber="406" type="discussion">
<paragraph id="414536FF4066F14EB88708BEC422FCA2" blockId="18.[151,1437,412,797]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB88708BEC600FD0F" bold="true" box="[199,308,663,688]" pageId="18" pageNumber="406">Remarks</emphasis>
.
<bibRefCitation id="256B4B0E4066F14EB90B08BEC6DEFD0F" author="Lewis" box="[331,490,663,688]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (1996) The type specimens and identity of the species described in the genus Otostigmus (subgenus Otostigmus) by R. I. Pocock in the collection of the Natural History Museum, London (Scolopendromorpha: Scolopendridae). Journal of Natural History, 30, 823 - 834. http: // dx. doi. org / 10.1080 / 00222939600770451" type="journal article" year="1996">Lewis (1996)</bibRefCitation>
synonymised Pocock’s
<taxonomicName id="86FA4D7C4066F14EBB5308B1C4ACFD0F" box="[787,920,664,688]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="738EEAED4066F14EBB5308B1C4ACFD0F" box="[787,920,664,688]" italics="true" pageId="18" pageNumber="406">O. ruficeps</emphasis>
</taxonomicName>
from
<collectingCountry id="39ED766F4066F14EBBAF08BEC31DFD0F" box="[1007,1065,663,688]" name="India" pageId="18" pageNumber="406">India</collectingCountry>
under
<taxonomicName id="86FA4D7C4066F14EBCC908B1C3C2FD0F" box="[1161,1270,664,688]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="738EEAED4066F14EBCC908B1C3C2FD0F" box="[1161,1270,664,688]" italics="true" pageId="18" pageNumber="406">multidens</emphasis>
</taxonomicName>
based on an examination of the
<typeStatus id="9E41885D4066F14EB93D0895C6EBFD6A" box="[381,479,700,725]" pageId="18" pageNumber="406" type="holotype">holotype</typeStatus>
which has only 5+5 coxosternal teeth and the last four pairs of legs missing. Its distribution when compared to
<taxonomicName id="86FA4D7C4066F14EB9BB08C9C5B8FD47" box="[507,652,736,760]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="738EEAED4066F14EB9BB08C9C5B8FD47" box="[507,652,736,760]" italics="true" pageId="18" pageNumber="406">O. multidens</emphasis>
</taxonomicName>
, is disjunct. The redescription of the species by Joshi &amp; Edgecombe (1913) is based on new material from the Western Ghats.
</paragraph>
</subSubSection>
</treatment>
</document>
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