373 lines
44 KiB
XML
373 lines
44 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F530FFDC0986A38B6355FAA5" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus kudrnai Serrano 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="4" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Serrano, Artur R. M.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
|
||
<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
|
||
<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-03-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5256</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>35</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
|
||
<mods:identifier type="ZooBank">CE0AC7CD-E55D-484C-9346-86F16C985DBE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7745350" ID-Zenodo-Dep="7745350" LSID="urn:lsid:plazi:treatment:AE0487C8F530FFDC0986A38B6355FAA5" httpUri="http://treatment.plazi.org/id/AE0487C8F530FFDC0986A38B6355FAA5" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<subSubSection box="[151,495,151,178]" pageId="3" pageNumber="4" type="nomenclature">
|
||
<paragraph blockId="3.[151,495,151,213]" box="[151,495,151,178]" pageId="3" pageNumber="4">
|
||
<heading bold="true" box="[151,495,151,178]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
|
||
<emphasis bold="true" box="[151,495,151,178]" pageId="3" pageNumber="4">
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[151,421,151,178]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis bold="true" box="[151,421,151,178]" italics="true" pageId="3" pageNumber="4">Macrocheilus kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[428,495,151,177]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,323,187,213]" pageId="3" pageNumber="4" type="description">
|
||
<paragraph blockId="3.[151,495,151,213]" box="[151,323,187,213]" pageId="3" pageNumber="4">
|
||
<figureCitation box="[151,243,187,213]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="3" pageNumber="4">Figs. 1b</figureCitation>
|
||
,
|
||
<figureCitation box="[255,286,187,213]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="3" pageNumber="4">2b</figureCitation>
|
||
,
|
||
<figureCitation box="[297,323,187,213]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="3" pageNumber="4">3a</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="4" type="materials_examined">
|
||
<paragraph blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[151,325,259,285]" pageId="3" pageNumber="4">Type material.</emphasis>
|
||
<materialsCitation collectingDate="2001-06-25" collectorName="A. Kudrna Jr. Lgt. & Th. Assmann" country="Benin" determinerName="Th. Assmann" location="Bensekou" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus box="[332,436,259,285]" pageId="3" pageNumber="4">Holotype</typeStatus>
|
||
male: “NE
|
||
<collectingCountry box="[569,635,259,285]" name="Benin" pageId="3" pageNumber="4">Benin</collectingCountry>
|
||
\
|
||
<locationDeviation box="[650,773,259,286]" location="Kandi" pageId="3" pageNumber="4">E of Kandi</locationDeviation>
|
||
\
|
||
<quantity box="[788,845,259,285]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="4" unit="km" value="4.0">4 km</quantity>
|
||
W of
|
||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F530FFDC0986A38B6355FAA5:23726005F530FFDB0A83A21F652BFEB5" box="[914,1024,259,285]" country="Benin" name="Bensekou" pageId="3" pageNumber="4">Bensekou</location>
|
||
\ 24–
|
||
<date box="[1081,1207,259,286]" pageId="3" pageNumber="4" value="2001-06-25">
|
||
<collectingDate box="[1081,1207,259,286]" pageId="3" pageNumber="4" value="2001-06-25">25.06.2001</collectingDate>
|
||
</date>
|
||
\
|
||
<collectorName box="[1220,1426,259,286]" pageId="3" pageNumber="4">A. Kudrna Jr. Lgt.</collectorName>
|
||
” \\ “Collection\
|
||
<collectorName box="[323,473,295,321]" pageId="3" pageNumber="4">Th. Assmann</collectorName>
|
||
\ Bleckede, CAB” (yellow label)\\ “
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[888,1041,295,321]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="spec. nov.">
|
||
<emphasis box="[888,1041,295,321]" italics="true" pageId="3" pageNumber="4">Macrocheilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1051,1168,295,321]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel box="[1051,1168,295,321]" pageId="3" pageNumber="4" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
[h]\ det.
|
||
<determinerName box="[1279,1432,295,321]" pageId="3" pageNumber="4">Th. Assmann</determinerName>
|
||
, 2008” [t] \\ “
|
||
</materialsCitation>
|
||
<typeStatus box="[292,399,331,357]" pageId="3" pageNumber="4">Holotype</typeStatus>
|
||
male\
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[475,722,331,357]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis box="[475,722,331,357]" italics="true" pageId="3" pageNumber="4">Macrocheilus kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[729,791,331,357]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel box="[729,791,331,357]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
\ A. Serrano det. 2022 [t] [h] [red card].
|
||
<materialsCitation collectingDate="2005-08-24" collectingDateMax="2005-08-28" collectingDateMin="2005-08-24" collectorName="Phillipe Moretto & Collection P. Sch" country="Burkina Faso" determinerName="Th. Assmann" elevation="232" location="Pama" longitude="0.7163889" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus box="[1240,1342,332,358]" pageId="3" pageNumber="4">Paratype</typeStatus>
|
||
:
|
||
<specimenCount box="[1353,1433,331,357]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
|
||
: “
|
||
<collectingCountry box="[161,316,367,394]" name="Burkina Faso" pageId="3" pageNumber="4">Burkina Faso</collectingCountry>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F530FFDC0986A38B6355FAA5:23726005F530FFDB0854A27360A8FE21" box="[325,387,367,393]" country="Burkina Faso" longitude="0.7163889" name="Pama" pageId="3" pageNumber="4">Pama</location>
|
||
\
|
||
<date box="[401,591,367,394]" pageId="3" pageNumber="4" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">
|
||
<collectingDate box="[401,591,367,394]" pageId="3" pageNumber="4" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">24–28.VIII.2005</collectingDate>
|
||
</date>
|
||
,\ 11º 17′00′N,
|
||
<geoCoordinate box="[752,868,367,396]" degrees="0" direction="east" minutes="42" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="59" value="0.7163889">0º 42′ 59E</geoCoordinate>
|
||
,
|
||
<quantity box="[879,947,367,393]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="3" pageNumber="4" unit="m" value="232.0">
|
||
<elevation box="[879,947,367,393]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="3" pageNumber="4" unit="m" value="232.0">232 m</elevation>
|
||
</quantity>
|
||
\
|
||
<collectorName box="[962,1145,367,394]" pageId="3" pageNumber="4">Phillipe Moretto</collectorName>
|
||
leg” \\ “Macr-07” (yellow label)\\ “
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[251,404,403,429]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="spec. nov.">
|
||
<emphasis box="[251,404,403,429]" italics="true" pageId="3" pageNumber="4">Macrocheilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[413,528,403,429]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel box="[413,528,403,429]" pageId="3" pageNumber="4" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
” [h]\ det.
|
||
<determinerName box="[647,799,403,429]" pageId="3" pageNumber="4">Th. Assmann</determinerName>
|
||
, 2008” \\ (
|
||
<collectorName box="[919,1123,403,430]" pageId="3" pageNumber="4">Collection P. Sch</collectorName>
|
||
̧le, Herrenberg)\\ “
|
||
</materialsCitation>
|
||
<typeStatus box="[1336,1437,403,429]" pageId="3" pageNumber="4">Paratype</typeStatus>
|
||
male\
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[222,471,439,465]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis box="[222,471,439,465]" italics="true" pageId="3" pageNumber="4">Macrocheilus kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[480,545,439,465]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel box="[480,545,439,465]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
\ A. Serrano det. 2022 [t] [h] [red card].
|
||
<materialsCitation box="[1012,1326,439,465]" collectionCode="PSC" country="Burkina Faso" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus box="[1012,1116,439,465]" pageId="3" pageNumber="4">Holotype</typeStatus>
|
||
deposited in
|
||
<collectionCode box="[1273,1326,439,465]" pageId="3" pageNumber="4">PSC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectionCode="TAC" country="Burkina Faso" pageId="3" pageNumber="4" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus box="[1338,1436,440,466]" pageId="3" pageNumber="4">Paratype</typeStatus>
|
||
deposited in
|
||
<collectionCode box="[294,351,475,501]" country="USA" lsid="urn:lsid:biocol.org:col:14176" name="Tarleton State University, Biological Sciences Department" pageId="3" pageNumber="4" type="Herbarium">TAC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[199,466,511,537]" pageId="3" pageNumber="4">Differential diagnosis.</emphasis>
|
||
Head and pronotum dark brown to blackish, lateral margins lighter; antennae and mouthparts brown or reddish-brown; legs brown; a short tooth just before the posterior angles of pronotum; elytra blackish, with one pair of median ovoid shaped orange spots; labrum long, anterior half ogive-shaped advanced; shiny and pubescent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
|
||
<paragraph blockId="3.[151,1437,259,2014]" box="[199,1075,655,682]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[199,345,655,681]" pageId="3" pageNumber="4">Description.</emphasis>
|
||
Length of
|
||
<typeStatus box="[468,570,655,681]" pageId="3" pageNumber="4">holotype</typeStatus>
|
||
:
|
||
<quantity box="[581,670,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="3" pageNumber="4" unit="mm" value="9.2">9.2 mm</quantity>
|
||
. Length of
|
||
<typeStatus box="[797,898,656,682]" pageId="3" pageNumber="4">paratype</typeStatus>
|
||
:
|
||
<quantity box="[908,987,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="4" unit="mm" value="9.5">9.5mm</quantity>
|
||
(male).
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="4">
|
||
<emphasis box="[199,260,691,717]" italics="true" pageId="3" pageNumber="4">Head</emphasis>
|
||
(
|
||
<figureCitation box="[277,360,691,717]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="3" pageNumber="4">Fig. 1b</figureCitation>
|
||
). Slightly wider than long (males: length
|
||
<quantity box="[837,1004,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" metricValueMax="1.84" metricValueMin="1.82" pageId="3" pageNumber="4" unit="mm" value="1.83" valueMax="1.84" valueMin="1.82">1.82–1.84 mm</quantity>
|
||
, width:
|
||
<quantity box="[1097,1266,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.175" metricValueMax="2.27" metricValueMin="2.08" pageId="3" pageNumber="4" unit="mm" value="2.175" valueMax="2.27" valueMin="2.08">2.08–2.27 mm</quantity>
|
||
). Surface shining, coarsely and deeply punctate, less dense in middle of the frons, each puncture with a long, decumbent and fine brown-yellowish seta, without microsculpture; frontal lateral impressions well marked and wrinkled, frons not excavated behind clypeus; last maxillary palpomere securiform, pubescent; eyes strongly prominent, temples long with rows of large setae along hind margin of eyes. Clypeus slightly punctured each puncture with a long, decumbent and fine brown-yellowish seta, anterior half with a row of 8–10 large submarginal setiferous punctures. Labrum smooth, brown or dark brown and shiny, 1.1times wider than long (males: length
|
||
<quantity box="[1074,1240,907,934]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" metricValueMax="9.6" metricValueMin="8.6" pageId="3" pageNumber="4" unit="mm" value="0.9099999999999999" valueMax="0.96" valueMin="0.86">0.86–0.96 mm</quantity>
|
||
, width
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="9.7" metricValueMax="10.1" metricValueMin="9.3" pageId="3" pageNumber="4" unit="mm" value="0.97" valueMax="1.01" valueMin="0.93">0.93–1.01 mm</quantity>
|
||
), posterior half subparallel-sided, anterior half slightly rounded along margins, surface protruded and ogiveshaped advanced, strongly lateral grooves of posterior half curved inward; a large seta at the widest point near the rim and 4 setae at the apex, two exactly on the margin from the top, and two beneath the margin just on the top (
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="3" pageNumber="4">Fig. 2b</figureCitation>
|
||
); posterior lateral half up to widest point with a fringe of short setae underneath. A pair of supraorbital setae on each side (
|
||
<figureCitation box="[274,357,1087,1113]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="3" pageNumber="4">Fig. 1b</figureCitation>
|
||
). Mentum with a central tooth, strongly thin and sharp like a needle, bent upwards at the end. Antennae with the first three antennomeres less densely pubescent with rather long setae, the fourth and following antennomeres are densely pubescent with shorter setae, first and third antennomeres almost cylindrical, second and fourth sub-pyriform, the remaining depressed, slightly longer than wide, with a conspicuous median longitudinal blackish smooth area; last antennomere more or less acuminate.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="4">
|
||
<emphasis box="[199,280,1267,1293]" italics="true" pageId="3" pageNumber="4">Thorax</emphasis>
|
||
. Pronotum cordiform (
|
||
<figureCitation box="[544,628,1267,1293]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="3" pageNumber="4">Fig. 1b</figureCitation>
|
||
), transverse, about 1.5–1.6 times wider than long (males: length
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.6" metricValueMin="1.47" pageId="3" pageNumber="4" unit="mm" value="1.535" valueMax="1.6" valueMin="1.47">1.47– 1.60 mm</quantity>
|
||
, width:
|
||
<quantity box="[342,508,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" metricValueMax="2.43" metricValueMin="2.29" pageId="3" pageNumber="4" unit="mm" value="2.36" valueMax="2.43" valueMin="2.29">2.29–2.43 mm</quantity>
|
||
), maximum width before the middle length; surface covered with coarse punctures, being more dense throughout the discal surface; each puncture bear a long and fine brown-yellowish seta, intervals smooth; anterior angles rounded, anterior margin arcuate; posterior margin straight in the middle, strongly rightangled sinuate towards the posterior angles which bear a tooth and a long seta; posterior margin not margined, anterior margin margined in the middle, lateral margins with complete and distinct margination since the anterior angles, reaching the posterior angles, slightly sinuate just anterior to posterior angles, forming a short tooth just before the angles, a long seta behind the widest part of pronotum, anterior angles with some large bristles; anterior transverse sulcus not distinct, pronotal base with a pair of longitudinal deep foveae separated by a median sulcus in a flat and depressed area that reaches the posterior margin. Scutellum triangular, strongly setiferous punctured. Prosternum coarsely punctured, without microsculpture.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,259,2014]" lastBlockId="4.[151,1437,150,1293]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<emphasis box="[199,269,1663,1689]" italics="true" pageId="3" pageNumber="4">Elytra</emphasis>
|
||
(
|
||
<figureCitation box="[286,367,1663,1689]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="3" pageNumber="4">Fig. 1b</figureCitation>
|
||
) 1.6 times longer than wide (males: length
|
||
<quantity box="[865,1031,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="5.97" metricValueMax="6.07" metricValueMin="5.87" pageId="3" pageNumber="4" unit="mm" value="5.97" valueMax="6.07" valueMin="5.87">5.87‒6.07 mm</quantity>
|
||
, width
|
||
<quantity box="[1116,1284,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="3.73" metricValueMax="3.78" metricValueMin="3.68" pageId="3" pageNumber="4" unit="mm" value="3.73" valueMax="3.78" valueMin="3.68">3.68‒3.78 mm</quantity>
|
||
), subparallelsided, widening slightly towards the posterior third, not covering the last two tergites; only seven striae visible, well-marked, all punctured and not reaching the base and the apex; intervals slightly convex, almost plane, coarse setiferous punctures in three rows on 2
|
||
<superScript attach="left" box="[579,595,1771,1785]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
‒7
|
||
<superScript attach="left" box="[629,642,1771,1785]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
intervals and two rows on 1
|
||
<superScript attach="left" box="[953,964,1771,1785]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
||
interval, decumbent seta brown-yellowish sub-erected and directed posteriad; lateral margin rounded in the shoulders, ending in deep furrow densely punctuated towards the scutellum. Scutellar striae visible through the deep punctures in the anterior region of first interval. Basal pore present. Umbilicate series with about 25 (
|
||
<typeStatus box="[732,833,1879,1905]" pageId="3" pageNumber="4">holotype</typeStatus>
|
||
) and 30 (
|
||
<typeStatus box="[935,1035,1879,1905]" pageId="3" pageNumber="4">paratype</typeStatus>
|
||
) pores, not interrupted in the middle although pores are more separated from each other in this area; 1 subapical pore shifted inward in the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[327,340,1951,1965]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
striae; apex with 4 extra pores,
|
||
<quantity box="[697,739,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="4" unit="in" value="2.0">2 in</quantity>
|
||
the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[1132,1145,1951,1965]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
striae and of 7
|
||
<superScript attach="left" box="[1311,1324,1951,1965]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
striae, respectively, and
|
||
<quantity box="[324,367,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="4" unit="in" value="2.0">2 in</quantity>
|
||
the lengthening of the 6
|
||
<superScript attach="left" box="[639,652,1987,2001]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
and 3
|
||
<superScript attach="left" box="[720,734,1987,2001]" fontSize="6" pageId="3" pageNumber="4">rd</superScript>
|
||
intervals; apical transparent membrane extending from sutural angle to the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[629,642,150,164]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||
interval. A median, more or less circular orange patch extends from the 2
|
||
<superScript attach="left" box="[165,181,186,200]" fontSize="6" pageId="4" pageNumber="5">nd</superScript>
|
||
to the 6
|
||
<superScript attach="left" box="[272,285,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||
striae (
|
||
<figureCitation box="[368,450,187,213]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="4" pageNumber="5">Fig. 1b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis box="[199,260,223,248]" italics="true" pageId="4" pageNumber="5">Legs.</emphasis>
|
||
brown or reddish-brown coloured; all tarsomeres pubescent, first three fore tarsomeres not dilated, fourth tarsomere slightly triangulate; fore tibiae on the outer edge with two bristles.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis box="[199,372,295,321]" italics="true" pageId="4" pageNumber="5">Ventral surface</emphasis>
|
||
. Almost all underside dark brown; whole underside, including abdominal sternites coarsely punctured and pubescent (yellowish colour), except gula, ventral and lateral sides of neck and propleura. Pubescence of prosternum and proepisternum suberected and slightly directed anteriad; pubescence of the remaining ventral sclerites dressed and directed posteriad. Elytral epipleura moderately punctured, pubescence dressed and directed posteriad.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis box="[199,366,475,501]" italics="true" pageId="4" pageNumber="5">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[383,466,475,501]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">Fig. 3a</figureCitation>
|
||
). Median lobe short, not knee shaped at the base, slightly dilated behind the middle, apical lamella short, acuminated at apex (dorsal view), downward (lateral view), ostium with one pair of scaled and sclerotized stripes on latero-dorsal region, bending downward near apex, becoming membranous and ending in a very large curled inward sclerotized plate; left paramere slightly longer than wide, rounded at apex, with a tooth in postero-dorsal edge.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" box="[199,415,655,681]" pageId="4" pageNumber="5">
|
||
<emphasis box="[199,283,655,681]" italics="true" pageId="4" pageNumber="5">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[199,473,690,716]" pageId="4" pageNumber="5">Intraspecific variation.</emphasis>
|
||
There is variability between the
|
||
<typeStatus box="[842,940,691,717]" pageId="4" pageNumber="5">holotype</typeStatus>
|
||
and
|
||
<typeStatus box="[995,1091,691,717]" pageId="4" pageNumber="5">paratype</typeStatus>
|
||
in the number of pores of umbilicate series (25 vs. 30).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[199,334,763,789]" pageId="4" pageNumber="5">Etymology.</emphasis>
|
||
This species is named in honour of Arnošt Kudrna (Ceské Budejovice,
|
||
<collectingCountry box="[1154,1338,763,789]" name="Czech Republic" pageId="4" pageNumber="5">Czech Republic</collectingCountry>
|
||
), a wellknown specialist in
|
||
<taxonomicName box="[369,513,799,825]" class="Insecta" family="Cicindelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Cicindelidae</taxonomicName>
|
||
, who collected the
|
||
<collectingCountry box="[723,790,799,825]" name="Benin" pageId="4" pageNumber="5">Benin</collectingCountry>
|
||
specimen and other
|
||
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[1015,1168,799,825]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1015,1168,799,825]" italics="true" pageId="4" pageNumber="5">Macrocheilus</emphasis>
|
||
</taxonomicName>
|
||
specimens here studied.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[199,315,835,861]" pageId="4" pageNumber="5">Remarks.</emphasis>
|
||
Taking into account the general shape of median lobe of aedeagus (
|
||
<figureCitation box="[1062,1142,835,861]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">Fig. 3a</figureCitation>
|
||
),
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[1162,1408,835,861]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis box="[1162,1408,835,861]" italics="true" pageId="4" pageNumber="5">Macrocheilus kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
seems to be close to
|
||
<taxonomicName authority="Basilewsky, 1967" authorityName="Basilewsky" authorityYear="1967" box="[422,782,871,897]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="clasispilus">
|
||
<emphasis box="[422,578,871,897]" italics="true" pageId="4" pageNumber="5">M. clasispilus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[585,782,871,897]" pageId="4" pageNumber="5" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
(see Basilewsky 1963,
|
||
<figureCitation box="[1047,1140,871,897]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="4" pageNumber="5">Figs. 2b</figureCitation>
|
||
,
|
||
<figureCitation box="[1151,1179,871,897]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">3b</figureCitation>
|
||
<emphasis box="[1186,1209,871,896]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
|
||
.
|
||
<figureCitation box="[1223,1304,871,897]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">Fig. 3a</figureCitation>
|
||
, this work). However, some external morphological characters allow separating both species. The pronotum of the new species is slightly less transverse than that of the latter species (1.52–1.56
|
||
<emphasis box="[889,912,943,968]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
|
||
. 1.65–1.67). On the other hand, just anterior to the posterior angles of pronotum, a short tooth occurs in the former species, but it is not present in the latter species (cf.
|
||
<figureCitation box="[198,290,1015,1041]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="4" pageNumber="5">Figs. 1b</figureCitation>
|
||
and
|
||
<figureCitation box="[349,378,1014,1041]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="4" pageNumber="5">5c</figureCitation>
|
||
). In addition, the elytral intervals of
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[802,1051,1015,1041]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis box="[802,1051,1015,1041]" italics="true" pageId="4" pageNumber="5">Macrocheilus kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1060,1125,1015,1041]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel box="[1060,1125,1015,1041]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are almost plane while the intervals of
|
||
<taxonomicName authorityName="Basilewsky" authorityYear="1967" box="[284,438,1051,1077]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="clasispilus">
|
||
<emphasis box="[284,438,1051,1077]" italics="true" pageId="4" pageNumber="5">M. clasispilus</emphasis>
|
||
</taxonomicName>
|
||
are subconvex. The new species also exhibits a more or less similar general shape pattern of the median lobe (in lateral and dorsal views) as in
|
||
<taxonomicName authority="(Reiche, 1832)" baseAuthorityName="Reiche" baseAuthorityYear="1832" box="[772,1224,1087,1113]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="biguttatus">
|
||
<emphasis box="[772,1046,1087,1113]" italics="true" pageId="4" pageNumber="5">Macrocheilus biguttatus</emphasis>
|
||
(Reiche, 1832)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Klug, 1834" authorityName="Klug" authorityYear="1834" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">Macrocheilus dorsalis</emphasis>
|
||
Klug, 1834
|
||
</taxonomicName>
|
||
, but is slightly different (cf.
|
||
<figureCitation box="[693,821,1123,1149]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">Figs. 3a, 3e</figureCitation>
|
||
and
|
||
<figureCitation box="[875,905,1123,1149]" captionStart="FIGURE 7" captionStartId="19.[152,255,1531,1555]" captionTargetBox="[163,1424,429,1485]" captionTargetId="figure-63@19.[151,1436,421,1492]" captionTargetPageId="19" captionText="FIGURE 7.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. dorsalis, ZAM (TAC); b, M. elegantulus, DOC? (MRAC); c, M. labrosus, RCA (PSC); d, M. longicollis, MOZ (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745297" httpUri="https://zenodo.org/record/7745297/files/figure.png" pageId="4" pageNumber="5">7a</figureCitation>
|
||
). Moreover, the general body colour of the new species and these two species is quite different.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="5">
|
||
Concerning
|
||
<taxonomicName authorityName="Klug" authorityYear="1834" box="[337,465,1195,1221]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis box="[337,465,1195,1221]" italics="true" pageId="4" pageNumber="5">M. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
, its left paramere does not present a tooth at the postero-dorsal edge. The new species differs from the remaining African species by the median lobe shape, among other external morphological characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,638,1266,1293]" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph blockId="4.[151,1437,150,1293]" box="[199,638,1266,1293]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[199,345,1266,1292]" pageId="4" pageNumber="5">Distribution</emphasis>
|
||
.
|
||
<collectingCountry box="[359,426,1267,1293]" name="Benin" pageId="4" pageNumber="5">Benin</collectingCountry>
|
||
and
|
||
<collectingCountry box="[481,634,1267,1293]" name="Burkina Faso" pageId="4" pageNumber="5">Burkina Faso</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |