172 lines
18 KiB
XML
172 lines
18 KiB
XML
<document id="3EE20E40DAE996225D0220870C257867" ID-DOI="10.11646/zootaxa.3985.4.3" ID-GBIF-Dataset="48cc8fed-5ec0-4ece-a82b-c2fa1ddd8893" ID-ISSN="1175-5326" ID-Zenodo-Dep="235350" ID-ZooBank="ECE53B6E-DBAF-4E98-BF8A-96FD9106EF35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461259944332" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Mock, Andrej & Enghoff, Henrik" docDate="2015" docId="682F87EBAD1A074484F4FE4AFB03581E" docLanguage="en" docName="zt03985p540.pdf" docOrigin="Zootaxa 3985 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thassoblaniulus simplarius Mauries 1985" docType="treatment" docVersion="9" lastPageNumber="536" masterDocId="9416FF93AD1707498463FFDEFF865D31" masterDocTitle="Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe" masterLastPageNumber="540" masterPageNumber="523" pageNumber="536" updateTime="1698609266564" updateUser="plazi">
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<mods:title id="0661EEA6A0C5932FF8312CAC63AA9A78">Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe</mods:title>
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<mods:name id="5286A8DA6B8D3745430057A607F0E4F9" type="personal">
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<mods:namePart id="0BA9532C53036ECA50A0653C07129637">Antić, Dragan Ž.</mods:namePart>
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<mods:namePart id="DE834B896C229863E641C2E52CA036B6">Mock, Andrej</mods:namePart>
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<mods:namePart id="AC67B609ED9EC0D0F2D1E13181324AF0">Enghoff, Henrik</mods:namePart>
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<mods:date id="81AC7FBA8583DCD71C03D65B733215BE">2015</mods:date>
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<treatment id="682F87EBAD1A074484F4FE4AFB03581E" ID-DOI="http://doi.org/10.5281/zenodo.5620900" ID-GBIF-Taxon="119636794" ID-Zenodo-Dep="5620900" LSID="urn:lsid:plazi:treatment:682F87EBAD1A074484F4FE4AFB03581E" httpUri="http://treatment.plazi.org/id/682F87EBAD1A074484F4FE4AFB03581E" lastPageNumber="536" pageId="13" pageNumber="536">
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<subSubSection id="A89C6576AD1A074484F4FE4AFE925CFE" pageId="13" pageNumber="536" type="nomenclature">
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<paragraph id="E03936FDAD1A074484F4FE4AFD195C9F" blockId="13.[151,671,404,463]" box="[151,671,404,430]" pageId="13" pageNumber="536">
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<heading id="BB718191AD1A074484F4FE4AFD195C9F" bold="true" box="[151,671,404,430]" fontSize="11" level="1" pageId="13" pageNumber="536" reason="1">
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<taxonomicName id="27864D7EAD1A074484F4FE4AFD195C9F" ID-CoL="564NL" authority="Mauries, 1985" authorityName="Mauries" authorityYear="1985" box="[151,671,404,430]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="simplarius">
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<emphasis id="D2F2EAEFAD1A074484F4FE4AFD195C9F" bold="true" box="[151,671,404,430]" pageId="13" pageNumber="536">
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<emphasis id="D2F2EAEFAD1A074484F4FE4AFE605C9F" bold="true" box="[151,486,404,430]" italics="true" pageId="13" pageNumber="536">Thassoblaniulus simplarius</emphasis>
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<bibRefCitation id="84174B0CAD1A0744858EFE4AFD195C9F" author="Mauries" box="[493,671,404,430]" pageId="13" pageNumber="536" refString="Mauries, J. - P. (1985) Les Diplopodes recoltes en 1982 par P. Beron et S. Andreev dans lile de Thasos (Grece). Thassoblaniulus, nouveau genre de Blaniulidae (Blaniulinae). Acta Zoologica Bulgarica, 28, 52 - 59." type="journal article" year="1985">Mauriès, 1985</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="E03936FDAD1A074484F4FE68FE925CFE" blockId="13.[151,671,404,463]" box="[151,276,438,463]" pageId="13" pageNumber="536">
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<figureCitation id="78BD2A78AD1A074484F4FE68FE925CFE" box="[151,276,438,463]" captionStart-0="FIGURES 34 – 37" captionStart-1="FIGURES 38 – 42" captionStartId-0="14.[151,264,1941,1963]" captionStartId-1="15.[151,264,1914,1936]" captionTargetBox-0="[151,1436,193,1919]" captionTargetBox-1="[190,1395,193,1896]" captionTargetId-0="figure@14.[151,1436,193,1920]" captionTargetId-1="figure@15.[190,1395,193,1897]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 34 – 37. Thassoblaniulus simplarius Mauriès, 1985, male. 34. anterior gonopods, anterior view; 35. anterior gonopods, posterior view; 36. anterior gonopods, lateral view; 37. first pair of legs, anterior view. f: postero-mesal flange, t: telopodite, cp: coxal processes. Scale lines: 0.1 mm." captionText-1="FIGURES 38 – 42. Thassoblaniulus simplarius Mauriès, 1985, male. 38. posterior gonopod, mesal view; 39. lamella of posterior gonopod, anterior view; 40. posterior gonopod, lateral view (basis of gonopod embedded in mounting glue); 41. apical part of lamella with solenomere, anterior view. 42. body ring 7, ventral view. ll: leaf-shaped lamella, so: solenomere. Scale lines: 0.1 mm for 38 – 40 and 42; 0.01 mm for 41." httpUri-0="https://zenodo.org/record/235360/files/figure.png" httpUri-1="https://zenodo.org/record/235361/files/figure.png" pageId="13" pageNumber="536">Figs 34–42</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A89C6576AD1A074484F4FE20FDB75F0A" pageId="13" pageNumber="536" type="materials_examined">
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<paragraph id="E03936FDAD1A074484F4FE20FDB75F0A" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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<emphasis id="D2F2EAEFAD1A074484F4FE20FEE65F26" bold="true" box="[151,352,510,535]" pageId="13" pageNumber="536">Material studied</emphasis>
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. Two males and five females (
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<collectionCode id="8697AE38AD1A074486A5FE21FC9D5F26" box="[710,795,511,535]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="13" pageNumber="536">ZMUC</collectionCode>
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) from Alistrati Cave, near Alistrati, Serres,
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<collectingCountry id="9891766DAD1A0744817AFE20FA1E5F26" box="[1305,1432,510,535]" name="Macedonia" pageId="13" pageNumber="536">Macedonia</collectingCountry>
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,
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<collectingCountry id="9891766DAD1A074484F4FDFDFF6F5F0A" box="[151,233,547,571]" name="Greece" pageId="13" pageNumber="536">Greece</collectingCountry>
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,
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<date id="9438103DAD1A07448497FDFCFEF35F0A" box="[244,373,546,571]" pageId="13" pageNumber="536" value="1987-06-28">28.06.1987</date>
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, leg. F. Gasparo.
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</paragraph>
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</subSubSection>
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<subSubSection id="A89C6576AD1A074484A4FD98FC6359D6" pageId="13" pageNumber="536" type="diagnosis">
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<paragraph id="E03936FDAD1A074484A4FD98FE385FFA" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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<emphasis id="D2F2EAEFAD1A074484A4FD98FEC75F6E" bold="true" box="[199,321,582,607]" pageId="13" pageNumber="536">Diagnosis.</emphasis>
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Differs clearly from
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<taxonomicName id="27864D7EAD1A07448621FD99FD205F6E" box="[578,678,582,607]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
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<emphasis id="D2F2EAEFAD1A07448621FD99FD205F6E" box="[578,678,582,607]" italics="true" pageId="13" pageNumber="536">T. radjai</emphasis>
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</taxonomicName>
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<emphasis id="D2F2EAEFAD1A074486D1FD98FD705F6E" bold="true" box="[690,758,582,607]" pageId="13" pageNumber="536">
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<taxonomicNameLabel id="C9C15794AD1A074486D1FD98FD705F6E" box="[690,758,582,607]" pageId="13" pageNumber="536" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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by numerous aspects of the anterior gonopods such as a different shape of distal part of coxal processes which are without shoulders, a bilobed mesal flange (
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<emphasis id="D2F2EAEFAD1A07448166FDB4FA885FB2" bold="true" box="[1285,1294,618,643]" pageId="13" pageNumber="536">f</emphasis>
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) and shorter telopodites (
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<emphasis id="D2F2EAEFAD1A07448541FD50FEAD5F96" bold="true" box="[290,299,654,679]" pageId="13" pageNumber="536">t</emphasis>
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), as well as by the presence of a higher number of ocelli and by the absence of a claw on the first pair of male legs (see
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<tableCitation id="AD040346AD1A07448538FD6CFE365FFA" box="[347,432,690,715]" captionStart="TABLE 2" captionStartId="6.[151,239,961,984]" captionTargetBox="[159,1427,1012,1804]" captionTargetPageId="6" captionText="TABLE 2. Differences between Thassoblaniulus simplarius Mauriès, 1985 and T. radjai sp. n." httpUri="http://table.plazi.org/id/B4F96675AD11074F84F4FC1FFB0C5EE9" pageId="13" pageNumber="536" tableUuid="B4F96675AD11074F84F4FC1FFB0C5EE9">Table 2</tableCitation>
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).
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</paragraph>
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<paragraph id="E03936FDAD1A074484A4FD08FDF05E06" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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<emphasis id="D2F2EAEFAD1A074484A4FD08FD935FDE" bold="true" box="[199,533,726,751]" pageId="13" pageNumber="536">Supplementary description.</emphasis>
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Specimens from the non-insular population from Alistrati Cave are somewhat different in view of the habitus than those from the population from the
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<typeStatus id="3F3D885FAD1A074487ADFD25FC785E22" box="[974,1022,763,787]" pageId="13" pageNumber="536">type</typeStatus>
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locality, Drakotrypa Cave, Island of Thasos, so additional details can be added:
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</paragraph>
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<paragraph id="E03936FDAD1A074484A4FC9CFEEB5EF7" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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Males: L
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<quantity id="277E9B18AD1A0744855AFC9DFE085E6B" box="[313,398,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="536" unit="mm" value="10.0">10 mm</quantity>
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, H
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<quantity id="277E9B18AD1A074485DFFC9CFD9E5E6B" box="[444,536,834,859]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="536" unit="mm" value="0.6">0.6 mm</quantity>
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. Females up to L
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<quantity id="277E9B18AD1A07448692FC9DFCC05E6B" box="[753,838,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="536" unit="mm" value="13.0">13 mm</quantity>
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, H
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<quantity id="277E9B18AD1A07448710FC9DFC565E6B" box="[883,976,835,859]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="536" unit="mm" value="0.7">0.7 mm</quantity>
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. Males with 37–41 body rings + telson (including 2 apodous rings), and with 5–6 ocelli on each side of the head. Females with 39–46 body rings + telson (including 2–3 apodous rings), and with 4–7 ocelli on each side of the head arranged mainly in one row, but also in two irregular rows.
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</paragraph>
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<paragraph id="E03936FDAD1A074484A4FC0CFBC5593E" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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Length of antennae 156 % of H in male, 137 % of H in female. Relative lengths of antennomeres 1–8 (8 = apical sensilla):
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<date id="9438103DAD1A0744852EFC28FE26593E" box="[333,416,1014,1039]" pageId="13" pageNumber="536" value="1917-08-18">8/18/17</date>
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/15/
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<date id="9438103DAD1A074485AEFC28FDA4593E" box="[461,546,1014,1039]" pageId="13" pageNumber="536" value="1917-06-15">17/15/6</date>
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/4 % in male,
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<date id="9438103DAD1A074486A3FC28FC95593E" box="[704,787,1014,1039]" pageId="13" pageNumber="536" value="1918-08-19">8/19/18</date>
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/15/
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<date id="9438103DAD1A07448723FC28FC13593E" box="[832,917,1014,1039]" pageId="13" pageNumber="536" value="1916-05-15">16/15/5</date>
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/4 % in female.
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</paragraph>
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<paragraph id="E03936FDAD1A074484A4FBC4FCDD5902" blockId="13.[151,1437,510,1327]" box="[199,859,1050,1075]" pageId="13" pageNumber="536">Length of setae ca. 10 % of H in male, 7 % of H in female.</paragraph>
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<paragraph id="E03936FDAD1A074484A4FBE0FAE7594A" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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Length of legs 102 % of H in male, 85 % of H in female. Relative lengths of podomeres from coxa to claw
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<date id="9438103DAD1A0744811BFBE1FF52594A" pageId="13" pageNumber="536" value="1921-11-13">21/ 13/11</date>
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/13/
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<date id="9438103DAD1A07448562FBBDFED1594A" box="[257,343,1123,1147]" pageId="13" pageNumber="536" value="2013-08-21">13/21/8</date>
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% in male, 23/
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<date id="9438103DAD1A07448661FBBDFDE3594A" box="[514,613,1123,1147]" pageId="13" pageNumber="536" value="2014-10-14">14/14/10</date>
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/
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<date id="9438103DAD1A0744860DFBBDFD44594A" box="[622,706,1123,1147]" pageId="13" pageNumber="536" value="2010-08-21">10/21/8</date>
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% in female; claw length/height
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<quantity id="277E9B18AD1A07448055FBBDFBE7594A" box="[1078,1121,1122,1147]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="13" pageNumber="536" unit="in" value="4.0">4 in</quantity>
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both male and female.
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</paragraph>
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<paragraph id="E03936FDAD1A074484A4FB58FC6359D6" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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<emphasis id="D2F2EAEFAD1A074484A4FB58FE7F59AE" bold="true" box="[199,505,1158,1183]" pageId="13" pageNumber="536">Note on cave adaptations.</emphasis>
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Like the new
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<taxonomicName id="27864D7EAD1A074486F9FB58FCD459AE" box="[666,850,1158,1183]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
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<emphasis id="D2F2EAEFAD1A074486F9FB58FCD459AE" box="[666,850,1158,1183]" italics="true" pageId="13" pageNumber="536">Thassoblaniulus</emphasis>
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</taxonomicName>
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species,
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<taxonomicName id="27864D7EAD1A074487D4FB59FBCF59AE" box="[951,1097,1158,1183]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="simplarius">
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<emphasis id="D2F2EAEFAD1A074487D4FB59FBCF59AE" box="[951,1097,1158,1183]" italics="true" pageId="13" pageNumber="536">T. simplarius</emphasis>
|
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</taxonomicName>
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not a real cave-dweller. Apart from the above-mentioned non-troglomorphic characters, this species also possesses a higher number of ocelli (4– 7 on each side of the head). Probably
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<taxonomicName id="27864D7EAD1A0744865EFB11FD4859D6" box="[573,718,1230,1255]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="simplarius">
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<emphasis id="D2F2EAEFAD1A0744865EFB11FD4859D6" box="[573,718,1230,1255]" italics="true" pageId="13" pageNumber="536">T. simplarius</emphasis>
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</taxonomicName>
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is a troglophilic species.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A89C6576AD1A074484A4FB2CFB03581E" pageId="13" pageNumber="536" type="distribution">
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||
<paragraph id="E03936FDAD1A074484A4FB2CFB03581E" blockId="13.[151,1437,510,1327]" pageId="13" pageNumber="536">
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||
<emphasis id="D2F2EAEFAD1A074484A4FB2CFEDF583A" bold="true" box="[199,345,1266,1291]" pageId="13" pageNumber="536">Distribution</emphasis>
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. Known only from two localities in eastern
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<collectingCountry id="9891766DAD1A0744870FFB2CFC6A583A" box="[876,1004,1266,1291]" name="Macedonia" pageId="13" pageNumber="536">Macedonia</collectingCountry>
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,
|
||
<collectingCountry id="9891766DAD1A07448798FB2DFBC9583A" box="[1019,1103,1267,1291]" name="Greece" pageId="13" pageNumber="536">Greece</collectingCountry>
|
||
: Drakotrypa Cave, Island of Thasos (
|
||
<typeStatus id="3F3D885FAD1A07448496FAC9FEA1581E" box="[245,295,1303,1327]" pageId="13" pageNumber="536">type</typeStatus>
|
||
locality) and Alistrati Cave, near Alistrati, Serres (
|
||
<figureCitation id="78BD2A78AD1A07448701FAC8FC33581E" box="[866,949,1302,1327]" captionStart="FIGURE 43" captionStartId="16.[151,250,1117,1139]" captionTargetBox="[151,1436,193,1095]" captionTargetId="figure@16.[151,1436,193,1095]" captionTargetPageId="16" captionText="FIGURE 43. Distribution of the genera Cibiniulus Verhoeff, 1897 and Thassoblaniulus Mauriès, 1985." httpUri="https://zenodo.org/record/235362/files/figure.png" pageId="13" pageNumber="536">Fig. 43</figureCitation>
|
||
, orange triangles).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |