treatments-xml/data/CD/4D/87/CD4D87D9FFAF6529FF33BEB46135F8C9.xml

<document id="CA3F3E065C3945B244B7B24A26AFF1C1" ID-CLB-Dataset="27710" ID-DOI="10.11646/zootaxa.4543.1.5" ID-GBIF-Dataset="498bf72a-fb02-4c71-adab-0450f4b0432b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617729" ID-ZooBank="92B59468-BDEE-43E7-83B0-24BC0DBE556D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928171120" checkinUser="plazi" docAuthor="Molineri, Carlos &amp; Granados-Martinez, Cristian E." docDate="2019" docId="CD4D87D9FFAF6529FF33BEB46135F8C9" docLanguage="en" docName="zootaxa.4543.1.5.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus cristales Molineri &amp; Granados-Martinez 2019, sp. nov." docType="treatment" docVersion="8" masterDocId="3174FFA1FFAA652CFFA4BE236010FF87" masterDocTitle="Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia" updateTime="1698715347377" updateUser="ExternalLinkService">
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<mods:title id="1060923648411906E5FFB35F4EC2867A">Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia</mods:title>
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<mods:name id="88C83148C64EAD1D36AC0F4D6E650462" type="personal">
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<mods:namePart id="DDF6C1C37EA7779C1141BAC873CA5F2A">Molineri, Carlos</mods:namePart>
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<mods:namePart id="C3BE5155B85E67887824B440A3E34E6D">Granados-Martinez, Cristian E.</mods:namePart>
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<mods:date id="07A1803DB00D7337D69D1A74B458234F">2019</mods:date>
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<subSubSection id="0DFE6544FFAF6529FF33BEB461E0FF35" box="[151,496,151,178]" pageId="5" type="nomenclature">
<paragraph id="455B36CFFFAF6529FF33BEB461E0FF35" blockId="5.[151,1114,151,247]" box="[151,496,151,178]" pageId="5">
<heading id="1E1381A3FFAF6529FF33BEB461E0FF35" bold="true" box="[151,496,151,178]" fontSize="11" level="1" pageId="5" reason="1">
<emphasis id="7790EADDFFAF6529FF33BEB461E0FF35" bold="true" box="[151,496,151,178]" pageId="5">
<taxonomicName id="82E44D4CFFAF6529FF33BEB4619CFF36" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[151,396,151,177]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="cristales" status="sp. nov.">
<emphasis id="7790EADDFFAF6529FF33BEB4619CFF36" bold="true" box="[151,396,151,177]" italics="true" pageId="5">Campsurus cristales</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CA357A6FFAF6529FE37BEBB61E0FF35" box="[403,496,152,178]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAF6529FF33BE99644AFF70" pageId="5" type="description">
<paragraph id="455B36CFFFAF6529FF33BE996168FF54" blockId="5.[151,1114,151,247]" box="[151,376,186,211]" pageId="5">
(
<figureCitation id="DDDF2A4AFFAF6529FF3BBE9960F7FF54" box="[159,231,186,211]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1</figureCitation>
D–G, 4A–E)
</paragraph>
<paragraph id="455B36CFFFAF6529FF33BEFD644AFF70" blockId="5.[151,1114,151,247]" box="[151,1114,222,247]" pageId="5">
http://zoobank.org/NomenclaturalActs/ 
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</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAF6529FF33BF05640AFD90" pageId="5" type="diagnosis">
<paragraph id="455B36CFFFAF6529FF33BF05640AFD90" blockId="5.[151,1437,294,1870]" pageId="5">
<emphasis id="7790EADDFFAF6529FF33BF056101FEB8" bold="true" box="[151,273,294,319]" pageId="5">Diagnosis.</emphasis>
<taxonomicName id="82E44D4CFFAF6529FEBEBF046212FEB8" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[282,514,294,319]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="cristales" status="sp. nov.">
<emphasis id="7790EADDFFAF6529FEBEBF046212FEB8" box="[282,514,294,319]" italics="true" pageId="5">Campsurus cristales</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFAF6529FDAFBF056276FEB8" bold="true" box="[523,614,294,319]" pageId="5">
<taxonomicNameLabel id="6CA357A6FFAF6529FDAFBF056276FEB8" box="[523,614,294,319]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is described from male and female adults. It can be distinguished from other species of 
<taxonomicName id="82E44D4CFFAF6529FEE8BF6861DCFEE5" authorityName="Eaton" authorityYear="1871" box="[332,460,331,354]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="7790EADDFFAF6529FEE8BF6861DCFEE5" box="[332,460,331,354]" italics="true" pageId="5">Campsurus</emphasis>
</taxonomicName>
by: 1) small to medium size (length of male FW 7.0–8.0 mm, female FW 
<quantity id="821C9B2AFFAF6529FAAEBF686562FEE4" box="[1290,1394,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="5" unit="mm" value="10.7">10.7 mm</quantity>
); 2) posterior margin of male abdominal sternum IX tri–lobed (
<figureCitation id="DDDF2A4AFFAF6529FCF5BF4D63A9FE00" box="[849,953,366,391]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1D</figureCitation>
); 3) pedestal almost touching each other medially at base, strongly elongated, parastylus broadly rounded apically, relatively wide and somewhat projected distally (
<figureCitation id="DDDF2A4AFFAF6529FF5BBF95614DFE48" box="[255,349,438,463]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Fig. 1D</figureCitation>
, 
<figureCitation id="DDDF2A4AFFAF6529FED4BF946185FE48" box="[368,405,439,463]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">4D</figureCitation>
), inner corner rounded and slightly projected (
<figureCitation id="DDDF2A4AFFAF6529FC6EBF95641EFE48" box="[970,1038,438,463]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Fig. 1</figureCitation>
D–4D); 4) penes with a strongly expanded dorsal area (
<emphasis id="7790EADDFFAF6529FE0EBFFF61D5FE74" box="[426,453,476,499]" italics="true" pageId="5">ea</emphasis>
in 
<figureCitation id="DDDF2A4AFFAF6529FE5CBFF96245FE74" box="[504,597,474,499]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4E</figureCitation>
), main lobe sclerotized, long and relatively acute apically, secondary membranous lobe shorter, rounded apically and cylindrical (
<figureCitation id="DDDF2A4AFFAF6529FC9DBFDD6392FD90" box="[825,898,510,535]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1</figureCitation>
D–E, 4D–E).
</paragraph>
</subSubSection>
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<paragraph id="455B36CFFFAF6529FF63BC0161F1FD20" blockId="5.[151,1437,294,1870]" pageId="5">
<emphasis id="7790EADDFFAF6529FF63BC016165FDBC" bold="true" box="[199,373,546,571]" pageId="5">Type material.</emphasis>
<materialsCitation id="F58C3C92FFAF6529FED8BC016496FDD8" ID-GBIF-Occurrence="2147524670" collectingDate="2016-10-26" collectorName="La Macarena &amp; C Granados" country="Colombia" county="La Macarena" elevation="257" latitude="2.2677777" location="Cano Cristales" longLatPrecision="21" longitude="-73.78306" municipality="Pailones" pageId="5" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="9A5F886DFFAF6529FED8BC0161F4FDBC" box="[380,484,546,571]" pageId="5" type="holotype">Holotype</typeStatus>
<specimenCount id="53E2FD46FFAF6529FE48BC006211FDBD" box="[492,513,547,570]" pageId="5" type="male">♂</specimenCount>
imago from 
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, 
<collectingRegion id="8720F82DFFAF6529FC93BC006363FDBD" box="[823,883,547,570]" country="Colombia" name="Meta" pageId="5">Meta</collectingRegion>
, 
<collectingCounty id="AC3A4E43FFAF6529FC24BC006405FDBD" box="[896,1045,547,570]" pageId="5">
<collectorName id="E8115319FFAF6529FC24BC006405FDBD" box="[896,1045,547,570]" pageId="5">La Macarena</collectorName>
</collectingCounty>
, 
<collectingMunicipality id="A53FACB5FFAF6529FB85BC016493FDBC" box="[1057,1155,546,571]" pageId="5">Pailones</collectingMunicipality>
, 
<location id="403B6014FFAF6529FB2ABC006526FDBC" LSID="urn:lsid:plazi:treatment:CD4D87D9FFAF6529FF33BEB46135F8C9:403B6014FFAF6529FB2ABC006526FDBC" box="[1166,1334,546,571]" country="Colombia" county="La Macarena" latitude="2.2677777" longLatPrecision="21" longitude="-73.78306" municipality="Pailones" name="Cano Cristales" pageId="5" stateProvince="Meta">Caño Cristales</location>
, 
<geoCoordinate id="20D05008FFAF6529FAE6BC0060A0FDD8" direction="north" orientation="latitude" pageId="5" precision="15" value="2.2677777">N 2° 16' 4&quot;</geoCoordinate>
/ 
<geoCoordinate id="20D05008FFAF6529FF62BC646176FDD8" box="[198,358,582,607]" direction="west" orientation="longitude" pageId="5" precision="15" value="-73.78306">W 73° 46' 59&quot;</geoCoordinate>
, 
<quantity id="821C9B2AFFAF6529FED6BC6561ABFDD9" box="[370,443,582,607]" metricMagnitude="2" metricUnit="m" metricValue="2.57" pageId="5" unit="m" value="257.0">
<elevation id="CEC9D1FCFFAF6529FED6BC6561ABFDD9" box="[370,443,582,607]" metricMagnitude="2" metricUnit="m" metricValue="2.57" pageId="5" unit="m" value="257.0">257 m</elevation>
</quantity>
, 
<date id="315A100FFFAF6529FE63BC65622AFDD8" box="[455,570,582,607]" pageId="5" value="2016-10-26">
<collectingDate id="211EE9E7FFAF6529FE63BC65622AFDD8" box="[455,570,582,607]" pageId="5" value="2016-10-26">26.x.2016</collectingDate>
</date>
, 
<collectingMethod id="9CA54ED8FFAF6529FDE2BC6562BCFDD9" box="[582,684,582,607]" pageId="5">light trap</collectingMethod>
18–20 hs, 
<collectorName id="E8115319FFAF6529FC83BC6463BCFDD8" box="[807,940,582,607]" pageId="5">C Granados</collectorName>
col. (MUSENUV)
</materialsCitation>
. 
<materialsCitation id="F58C3C92FFAF6529FB34BC6461BBFD04" ID-GBIF-Occurrence="2147524576" collectionCode="CM" country="Colombia" pageId="5" specimenCode="IBN803" specimenCount="16" specimenCount-male="16" stateProvince="Meta" typeStatus="paratype">
<typeStatus id="9A5F886DFFAF6529FB34BC646511FDD8" box="[1168,1281,583,607]" pageId="5" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="53E2FD46FFAF6529FAA8BC656554FDD9" box="[1292,1348,582,607]" pageId="5" type="male">16 ♂</specimenCount>
imagos (1 on slide 
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<collectionCode id="23F5AE0AFFAF6529FED2BC4861B3FD04" box="[374,419,619,643]" country="China" name="Chongqing Museum" pageId="5">CM</collectionCode>
)
</materialsCitation>
and 
<materialsCitation id="F58C3C92FFAF6529FE42BC486356FD04" ID-GBIF-Occurrence="2147524704" box="[486,838,618,643]" collectionCode="CM" country="Colombia" pageId="5" specimenCode="IBN804" specimenCount="1" specimenCount-female="1" stateProvince="Meta" typeStatus="paratype">
<specimenCount id="53E2FD46FFAF6529FE42BC486203FD05" box="[486,531,619,643]" pageId="5" type="female">1 ♀</specimenCount>
imago (slide 
<specimenCode id="15429EB4FFAF6529FD10BC486301FD04" box="[692,785,619,643]" collectionCode="IBN" pageId="5">IBN804</specimenCode>
<collectionCode id="23F5AE0AFFAF6529FCB5BC48632EFD04" box="[785,830,619,643]" country="China" name="Chongqing Museum" pageId="5">CM</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="F58C3C92FFAF6529FCF0BC4861CDFD20" ID-GBIF-Occurrence="2147524705" collectingDate="2016-10-26" collectionCode="IBN, CEBUC" collectorName="La Macarena &amp; C Granados" country="Colombia" county="La Macarena" elevation="257" latitude="2.2677777" location="Cano Cristales" longLatPrecision="21" longitude="-73.78306" municipality="Pailones" pageId="5" specimenCount="17" specimenCount-female="1" specimenCount-male="16" stateProvince="Meta" typeStatus="paratype">
same data as holotype (
<specimenCount id="53E2FD46FFAF6529FBC1BC486481FD05" box="[1125,1169,619,643]" pageId="5" type="male">8 ♂</specimenCount>
and 
<specimenCount id="53E2FD46FFAF6529FB69BC4864E0FD04" box="[1229,1264,619,643]" pageId="5" type="female">1♀</specimenCount>
in 
<collectionCode id="23F5AE0AFFAF6529FABDBC48655CFD05" box="[1305,1356,619,642]" pageId="5">IBN</collectionCode>
, 
<specimenCount id="53E2FD46FFAF6529FAFEBC48656DFD04" box="[1370,1405,619,643]" pageId="5" type="male">4♂</specimenCount>
in MUSENUV, 
<specimenCount id="53E2FD46FFAF6529FE88BCAC615FFD20" box="[300,335,655,679]" pageId="5" type="male">4♂</specimenCount>
in 
<collectionCode id="23F5AE0AFFAF6529FED6BCAC61C5FD20" box="[370,469,655,679]" pageId="5">CEBUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="455B36CFFFAF6529FF63BC916569FA89" blockId="5.[151,1437,294,1870]" pageId="5">
Male imago. Length (mm): body, 7.2–7.7; fore wing, 7.0–8.0; hind wing, 3.5–4.0; fore leg, 3.5–4.2; cerci, 20.0–21.0. General coloration yellowish white, widely shaded with gray (
<figureCitation id="DDDF2A4AFFAF6529FC7FBCF56427FD68" box="[987,1079,726,751]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4A</figureCitation>
). Head completely shaded with black dorsally, and on antenna (except for hyaline flagellum), ventrally with gray marks along carinae. Thorax (
<figureCitation id="DDDF2A4AFFAF6529FF3BBD3D613DFCB0" box="[159,301,798,823]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4A, 4C</figureCitation>
). Pronotum translucent, extensively shaded with black, darker on anterior portion, medial line, and margins; prosternum yellowish white. Mesonotum yellowish, shaded with gray medially on parapsidal sutures, and on a triangle between PSP; pleura and sterna yellowish white, shaded with slightly gray. Metanotum yellowish white, shaded with gray medially. Meso furcasternal plates as in 
<figureCitation id="DDDF2A4AFFAF6529FCD8BDA963C5FC24" box="[892,981,906,931]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4C</figureCitation>
. Legs. Fore leg completely shaded with gray; middle and hind legs yellowish translucent, shaded completely with gray. Wings. Membrane hyaline, except C and Sc areas shaded with brownish gray, veins hyaline, except C, Sc, and R 
<subScript id="D960348AFFAF6529FBA8BDC26404FC68" attach="left" box="[1036,1044,993,1007]" fontSize="6" pageId="5">1</subScript>
brownish on basal 2/3. Hind wing hyaline, except base of veins yellowish. Abdomen (
<figureCitation id="DDDF2A4AFFAF6529FD45BDDA632AFB95" box="[737,826,1017,1042]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4A</figureCitation>
) whitish translucent, extensively shaded with gray on terga, except on small paler dots. Abdominal sterna translucent white, shaded with gray on medial line of sterna VII–IX. Genitalia (
<figureCitation id="DDDF2A4AFFAF6529FED0BA6261AFFBDD" box="[372,447,1089,1114]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1</figureCitation>
D–E, 4D–E)): yellowish white; sternum IX with three slightly marked lobes on hind margin (arrows in 
<figureCitation id="DDDF2A4AFFAF6529FED7BA4661C9FBF9" box="[371,473,1125,1150]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1D</figureCitation>
), medial line and hind margin black; pedestal of each side touching basally, with apically rounded parastylus (
<emphasis id="7790EADDFFAF6529FE7BBAA96211FB26" box="[479,513,1162,1185]" italics="true" pageId="5">mo</emphasis>
in 
<figureCitation id="DDDF2A4AFFAF6529FD8CBAAA6292FB25" box="[552,642,1161,1186]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4D</figureCitation>
), round inner corner small, slightly projected (
<emphasis id="7790EADDFFAF6529FB32BAA964A2FB26" box="[1174,1202,1162,1185]" italics="true" pageId="5">mi</emphasis>
in 
<figureCitation id="DDDF2A4AFFAF6529FB7DBAAA6523FB25" box="[1241,1331,1161,1186]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4D</figureCitation>
); forceps short and translucent, shaded with gray basally; base of main lobe of penes sclerotized, very expanded dorsally (
<emphasis id="7790EADDFFAF6529FA25BA8D658BFB42" box="[1409,1435,1198,1221]" italics="true" pageId="5">ea</emphasis>
in 
<figureCitation id="DDDF2A4AFFAF6529FF1EBAF2611CFB6D" box="[186,268,1233,1258]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Figs. 4</figureCitation>
D–E), apical portion narrowing abruptly and finger like, main lobe sclerotized and acute apically, secondary lobe membranous and rounded apically, slender and cylindrical. Caudal filament hyaline translucent.
</paragraph>
<paragraph id="455B36CFFFAF6529FF63BB3A629FF98D" blockId="5.[151,1437,294,1870]" pageId="5">
Female adult. Length (mm): body, 8.0; fore wing, 10.7; hind wing, 4.5; fore leg, 1.2; cerci broken. Similar to male imago, except pigmentation more heavily marked (
<figureCitation id="DDDF2A4AFFAF6529FCB5BB1E6379FAD1" box="[785,873,1341,1366]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4B</figureCitation>
). Thorax. Pronotum shaded with gray, except pale transverse line between anterior and posterior portions; thin black medial line surrounded by a pale area along anterior portion of pronotum. Mesonotum (
<figureCitation id="DDDF2A4AFFAF6529FD28BBA662F5FA19" box="[652,741,1413,1438]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4B</figureCitation>
) additionally shaded gray on anterior triangular mark (after mesonotal protuberance). Legs shaded completely with gray. Wing membranes yellowish translucent. Abdomen shaded extensively with black on dorsum (
<figureCitation id="DDDF2A4AFFAF6529FDDEBBEE62C3FA61" box="[634,723,1485,1510]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Fig. 4B</figureCitation>
); sternum VIII with small anteromedian subovate socket (
<figureCitation id="DDDF2A4AFFAF6529FAC0BBEE60B0F98D" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="5">Figs. 1</figureCitation>
F–G). Caudal filaments translucent whitish.
</paragraph>
<paragraph id="455B36CFFFAF6529FF63B8356521F9AA" blockId="5.[151,1437,294,1870]" box="[199,1329,1557,1582]" pageId="5">Egg. Length, 275–300 µm; width, 225–250 µm. Oval, cup–shaped, yellowish with whitish polar cap.</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAF6529FF63B81A63A7F9D5" box="[199,951,1593,1618]" pageId="5" type="etymology">
<paragraph id="455B36CFFFAF6529FF63B81A63A7F9D5" blockId="5.[151,1437,294,1870]" box="[199,951,1593,1618]" pageId="5">
<emphasis id="7790EADDFFAF6529FF63B81A615EF9D5" bold="true" box="[199,334,1593,1618]" pageId="5">Etymology.</emphasis>
The name alludes to the 
<typeStatus id="9A5F886DFFAF6529FDC2B8196286F9D5" box="[614,662,1594,1618]" pageId="5">type</typeStatus>
locality (Caño Cristales).
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAF6529FF63B87E629BF9F1" box="[199,651,1629,1654]" pageId="5" type="distribution">
<paragraph id="455B36CFFFAF6529FF63B87E629BF9F1" blockId="5.[151,1437,294,1870]" box="[199,651,1629,1654]" pageId="5">
<emphasis id="7790EADDFFAF6529FF63B87E6149F9F1" bold="true" box="[199,345,1629,1654]" pageId="5">Distribution</emphasis>
(
<figureCitation id="DDDF2A4AFFAF6529FECCB87E61BDF9F1" box="[360,429,1629,1654]" captionStart="FIGURE 5" captionStartId="7.[151,250,1026,1048]" captionTargetBox="[152,1435,196,1002]" captionTargetId="figure@7.[151,1436,193,1005]" captionTargetPageId="7" captionText="FIGURE 5. Map indicating the distributional records of the new species." httpUri="https://zenodo.org/record/2617739/files/figure.png" pageId="5">Fig. 5</figureCitation>
). 
<collectingCountry id="3DF3765FFFAF6529FE66B87E6220F9F1" box="[450,560,1629,1654]" name="Colombia" pageId="5">Colombia</collectingCountry>
(
<collectingRegion id="8720F82DFFAF6529FD9BB87D626FF9F1" box="[575,639,1630,1654]" country="Colombia" name="Meta" pageId="5">Meta</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAF6529FF63B8A26135F8C9" pageId="5" type="discussion">
<paragraph id="455B36CFFFAF6529FF63B8A26135F8C9" blockId="5.[151,1437,294,1870]" pageId="5">
<emphasis id="7790EADDFFAF6529FF63B8A2615CF91D" bold="true" box="[199,332,1665,1690]" pageId="5">Discussion.</emphasis>
<taxonomicName id="82E44D4CFFAF6529FEF1B8A1622CF91D" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[341,572,1665,1690]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="cristales" status="sp. nov.">
<emphasis id="7790EADDFFAF6529FEF1B8A1622CF91D" box="[341,572,1665,1690]" italics="true" pageId="5">Campsurus cristales</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFAF6529FDE1B8A262B1F91D" bold="true" box="[581,673,1665,1690]" pageId="5">
<taxonomicNameLabel id="6CA357A6FFAF6529FDE1B8A262B1F91D" box="[581,673,1665,1690]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to 
<taxonomicName id="82E44D4CFFAF6529FCD1B8A164BFF91D" authority="Molineri &amp; Salles" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[885,1199,1665,1690]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="janae">
<emphasis id="7790EADDFFAF6529FCD1B8A163C6F91E" box="[885,982,1666,1689]" italics="true" pageId="5">C. janae</emphasis>
Molineri &amp; Salles
</taxonomicName>
, and will key to that species in 
<bibRefCitation id="21754B3EFFAF6529FEA8B886623AF939" author="Molineri, C. &amp; Salles, F. F." box="[268,554,1701,1726]" pageId="5" pagination="301 - 354" refId="ref4106" refString="Molineri, C. &amp; Salles, F. F. (2017) Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus. Zootaxa, 4300 (3), 301 - 354. https: // doi. org / 10.11646 / zootaxa. 4300.3.1" type="journal article" year="2017">Molineri &amp; Salles (2017)</bibRefCitation>
. Nevertheless, the pedestals are broader and parastylus more projected distally in the new species. Also, the penes are even more expanded in a dorsal ovate flat structure (ea in 
<figureCitation id="DDDF2A4AFFAF6529FAAEB8EA6549F965" box="[1290,1369,1737,1762]" captionStart="FIGURES 4" captionStartId="6.[151,264,1938,1961]" captionTargetBox="[209,1379,193,1918]" captionTargetId="figure@6.[208,1379,193,1918]" captionTargetPageId="6" captionText="FIGURES 4A–E. Campsurus cristales sp. nov.: A, male imago, d.v.; B, female imago, d.v. (PSP= Postero Scutal Protuberances); C, male imago, v.v. (mfp= mesofurcasternal plates); D, male genitalia, v.v.; E, same, l.v. (ea ear–like projection; im= inner projected margin of pedestal; ml= main lobe; om= outer projected margin of pedestal; sl= secondary lobe." httpUri="https://zenodo.org/record/2617737/files/figure.png" pageId="5">Figs. 4</figureCitation>
D–E), which is more circular in 
<taxonomicName id="82E44D4CFFAF6529FE18B8CD620BF882" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[444,539,1774,1797]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="janae">
<emphasis id="7790EADDFFAF6529FE18B8CD620BF882" box="[444,539,1774,1797]" italics="true" pageId="5">C. janae</emphasis>
</taxonomicName>
. Additionally, the main lobe of penes in the new species is longer and apically acute (it is shorter and rounded in 
<taxonomicName id="82E44D4CFFAF6529FDBEB931626AF8AE" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[538,634,1810,1833]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="janae">
<emphasis id="7790EADDFFAF6529FDBEB931626AF8AE" box="[538,634,1810,1833]" italics="true" pageId="5">C. janae</emphasis>
</taxonomicName>
). Finally, the tri–lobed hind margin of the sternum IX is unique for the new species.
</paragraph>
</subSubSection>
</treatment>
</document>