289 lines
35 KiB
XML
289 lines
35 KiB
XML
<document id="3B1074A04E8DF4D581D55FE6433E80EF" ID-CLB-Dataset="30456" ID-DOI="10.11646/zootaxa.4399.2.1" ID-GBIF-Dataset="834234e8-ac96-4b19-a30e-3edc31fe4897" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206522" ID-ZooBank="43A975F2-05F6-4A29-A588-7787AF63589C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521899272936" checkinUser="plazi" docAuthor="Canseco-Márquez, Luis, Ramírez-González, Cynthia G. & Campbell, Jonathan A." docDate="2018" docId="B45787B9611CFF95FF6E711339B3FBA6" docLanguage="en" docName="zootaxa.4399.2.1.pdf" docOrigin="Zootaxa 4399 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chersodromus liebmanni Reinhardt 1860" docType="treatment" docVersion="7" lastPageNumber="156" masterDocId="486EFFC1611EFF90FFF97572381EFFFE" masterDocTitle="Taxonomic review of the rare Mexican snake genus Chersodromus (Serpentes: Dipsadidae), with the description of two new species" masterLastPageNumber="169" masterPageNumber="151" pageNumber="153" updateTime="1698532892634" updateUser="ExternalLinkService">
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<mods:title id="7AACC1E7E45AD644049FAAE646CC13DF">Taxonomic review of the rare Mexican snake genus Chersodromus (Serpentes: Dipsadidae), with the description of two new species</mods:title>
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<mods:namePart id="E2EA4945A811BB327E6BE87C92916A99">Canseco-Márquez, Luis</mods:namePart>
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<mods:namePart id="651A054709CE7DD85B364ED05F455105">Ramírez-González, Cynthia G.</mods:namePart>
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<mods:namePart id="06616D76F761B898C80598DEB260CD8E">Campbell, Jonathan A.</mods:namePart>
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<mods:title id="E72798F6CC6B164CBED43C5EB89144FE">Zootaxa</mods:title>
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<treatment id="B45787B9611CFF95FF6E711339B3FBA6" ID-DOI="http://doi.org/10.5281/zenodo.5965345" ID-GBIF-Taxon="142524238" ID-Zenodo-Dep="5965345" LSID="urn:lsid:plazi:treatment:B45787B9611CFF95FF6E711339B3FBA6" httpUri="http://treatment.plazi.org/id/B45787B9611CFF95FF6E711339B3FBA6" lastPageId="5" lastPageNumber="156" pageId="2" pageNumber="153">
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<subSubSection id="74E46524611CFF92FF6E71133977FB62" pageId="2" pageNumber="153" type="nomenclature">
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<paragraph id="3C4136AF611CFF92FF6E71133A85FB85" blockId="2.[151,667,1121,1181]" box="[151,667,1121,1147]" pageId="2" pageNumber="153">
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<heading id="670981C3611CFF92FF6E71133A85FB85" bold="true" box="[151,667,1121,1147]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
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<taxonomicName id="FBFE4D2C611CFF92FF6E71133A85FB85" authority="Reinhardt, 1860" authorityName="Reinhardt" authorityYear="1860" box="[151,667,1121,1147]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="liebmanni">
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<emphasis id="0E8AEABD611CFF92FF6E71133A85FB85" bold="true" box="[151,667,1121,1147]" pageId="2" pageNumber="153">
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<emphasis id="0E8AEABD611CFF92FF6E711339D4FB85" bold="true" box="[151,458,1121,1147]" italics="true" pageId="2" pageNumber="153">Chersodromus liebmanni</emphasis>
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Reinhardt, 1860
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="3C4136AF611CFF92FF6E71F63977FB62" blockId="2.[151,667,1121,1181]" box="[151,361,1156,1181]" pageId="2" pageNumber="153">
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<figureCitation id="A4C52A2A611CFF92FF6E71F638C4FB62" box="[151,218,1156,1181]" captionStart="FIGURE2" captionStartId="3.[151,250,1951,1973]" captionTargetBox="[345,1246,224,1919]" captionTargetId="figure@3.[340,1246,215,1919]" captionTargetPageId="3" captionText="FIGURE2. Dorsal(A), ventral (B) and lateral (C)views of the head (Scale = 3 mm); rightmaxilla (D, 2.61 mm); and from left to right: sulcate, lateral, and asulcate views of the hemipenis (E–G, Scale = 3.9 mm) of Chersodromus liebmanni (MZFC 28687)." httpUri="https://zenodo.org/record/1206526/files/figure.png" pageId="2" pageNumber="153">Fig. 2</figureCitation>
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,
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||
<figureCitation id="A4C52A2A611CFF92FF1F71F63914FB62" box="[230,266,1156,1180]" captionStart="FIGURE 3" captionStartId="4.[151,250,1591,1613]" captionTargetBox="[294,1299,179,1581]" captionTargetId="figure@4.[292,1299,179,1581]" captionTargetPageId="4" captionText="FIGURE 3. Adult of Chersodromus liebmanni in life (MZFC 28687) from Totontepec, Oaxaca (A), and its locality in Cuautlapan, Veracruz (B)." httpUri="https://zenodo.org/record/1206528/files/figure.png" pageId="2" pageNumber="153">3A</figureCitation>
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,
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<tableCitation id="717C0314611CFF92FEEF71F63977FB62" box="[278,361,1156,1181]" captionStart="TABLE 1" captionStartId="14.[151,239,620,643]" captionText="TABLE 1. Select quatitative and qualitative features increased by habitat/distribution data from the four currently recognized species of Chersodromus." pageId="2" pageNumber="153">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="74E46524611CFF92FF6E71BE3C70FA29" pageId="2" pageNumber="153" type="description">
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<paragraph id="3C4136AF611CFF92FF6E71BE3AC0FAE1" blockId="2.[151,1437,1228,1495]" pageId="2" pageNumber="153">
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<materialsCitation id="8C963CF2611CFF92FF6E71BE39A5FAFE" ID-GBIF-Occurrence="1836204294" collectionCode="ZMUC" country="Mexico" location="Mexico" pageId="2" pageNumber="153" specimenCode="ZMUC 60561" specimenCount="1" typeStatus="holotype">
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<taxonomicName id="FBFE4D2C611CFF92FF6E71BE3A57FB1C" authority="Reinhardt, 1861" authorityName="Reinhardt" authorityYear="1861" box="[151,585,1228,1250]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="liebmanni">
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<emphasis id="0E8AEABD611CFF92FF6E71BE3987FB1C" box="[151,409,1228,1250]" italics="true" pageId="2" pageNumber="153">Chersodromus liebmanni</emphasis>
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||
Reinhardt, 1861
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||
</taxonomicName>
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||
, Vidensk. Meddel. Naturhist. Foren. Kjöbenhavn 2:243.
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<typeStatus id="E345880D611CFF92FB6E71BE3CE7FB1C" box="[1175,1273,1228,1250]" pageId="2" pageNumber="153" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="6C589ED4611CFF92FAFD71BE3D86FB1C" box="[1284,1432,1228,1250]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="153">ZMUC 60561</specimenCode>
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. Type-locality: “
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<collectingCountry id="44E9763F611CFF92FE91719839A5FAFE" box="[360,443,1258,1280]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation id="8C963CF2611CFF92FE4771983AC0FAE1" ID-GBIF-Occurrence="1836204296" country="Mexico" location="Veracruz" pageId="2" pageNumber="153" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
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” Restricted to Cuautlapan,
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||
<collectingRegion id="FE3AF84D611CFF92FD1871993B20FAFE" box="[737,830,1259,1280]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
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,
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<collectingCountry id="44E9763F611CFF92FCB271983B85FAFE" box="[843,923,1258,1280]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
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, by Smith & Taylor (1950:347), and to Mirador,
|
||
<collectingRegion id="FE3AF84D611CFF92FF3E7078393AFAE1" box="[199,292,1290,1311]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
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,
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<collectingRegion id="FE3AF84D611CFF92FED4707B3984FAE1" box="[301,410,1289,1311]" country="Mexico" name="Distrito Federal" pageId="2" pageNumber="153">Mexico by</collectingRegion>
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Smith & Braestrup (1963:240).
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</materialsCitation>
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</paragraph>
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<paragraph id="3C4136AF611CFF92FF6E705A3ACAFA85" blockId="2.[151,1437,1228,1495]" pageId="2" pageNumber="153">
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<materialsCitation id="8C963CF2611CFF92FF6E705A39A2FAA2" ID-GBIF-Occurrence="1836204299" collectionCode="ZMUC" country="Mexico" location="Mexico" pageId="2" pageNumber="153" specimenCode="ZMUC 60562" specimenCount="2" typeStatus="holotype">
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<taxonomicName id="FBFE4D2C611CFF92FF6E705A3C5CFAC0" authority="Reinhardt, 1861 Vidensk. Meddel. Naturhist. Foren. Kjobenhavn" authorityName="Reinhardt" authorityYear="1861" box="[151,1090,1320,1342]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="nigricans">
|
||
<emphasis id="0E8AEABD611CFF92FF6E705A398BFAC0" box="[151,405,1320,1342]" italics="true" pageId="2" pageNumber="153">Chersodromus nigricans</emphasis>
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||
Reinhardt, 1861 Vidensk. Meddel. Naturhist. Foren. Kjöbenhavn
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||
</taxonomicName>
|
||
2:245.
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||
<typeStatus id="E345880D611CFF92FB6D705A3CE8FAC0" box="[1172,1270,1320,1342]" pageId="2" pageNumber="153" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="6C589ED4611CFF92FAFA705A3D87FAC0" box="[1283,1433,1320,1342]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="153">ZMUC 60562</specimenCode>
|
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.
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||
<typeStatus id="E345880D611CFF92FF3E703538E7FAA2" box="[199,249,1351,1372]" pageId="2" pageNumber="153">Type</typeStatus>
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locality: “
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<collectingCountry id="44E9763F611CFF92FE90703439A2FAA2" box="[361,444,1350,1372]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
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</materialsCitation>
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.
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||
<materialsCitation id="8C963CF2611CFF92FE4670343ACAFA85" ID-GBIF-Occurrence="1836204301" country="Mexico" location="Veracruz" pageId="2" pageNumber="153" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
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” Restricted to Cuautlapan,
|
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<collectingRegion id="FE3AF84D611CFF92FD1B70353B21FAA2" box="[738,831,1351,1372]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
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,
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||
<collectingCountry id="44E9763F611CFF92FCB270343B85FAA2" box="[843,923,1350,1372]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
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, by Smith & Taylor (1950:347), and to Mirador,
|
||
<collectingRegion id="FE3AF84D611CFF92FF3E7014393AFA85" box="[199,292,1382,1403]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
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,
|
||
<collectingRegion id="FE3AF84D611CFF92FED470173984FA85" box="[301,410,1381,1403]" country="Mexico" name="Distrito Federal" pageId="2" pageNumber="153">Mexico by</collectingRegion>
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Smith & Braetrup (1963:240).
|
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</materialsCitation>
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</paragraph>
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<paragraph id="3C4136AF611CFF92FF6E70F63A61FA46" blockId="2.[151,1437,1228,1495]" pageId="2" pageNumber="153">
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<materialsCitation id="8C963CF2611CFF92FF6E70F63A61FA46" ID-GBIF-Occurrence="1836204305" location="Huatusco" municipality="Huanusco" pageId="2" pageNumber="153" specimenCount="2" stateProvince="Veracruz" typeStatus="holotype">
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<taxonomicName id="FBFE4D2C611CFF92FF6E70F63A0FFA64" authority="Peters, 1861" authorityName="Peters" authorityYear="1861" box="[151,529,1412,1434]" class="Reptilia" family="Colubridae" genus="Opisthiodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="0E8AEABD611CFF92FF6E70F6399EFA64" box="[151,384,1412,1434]" italics="true" pageId="2" pageNumber="153">Opisthiodon torquatus</emphasis>
|
||
Peters, 1861
|
||
</taxonomicName>
|
||
, Monatsber. königl. Akad. Wiss. Berlin. 1861:461.
|
||
<typeStatus id="E345880D611CFF92FBBB70F63CBAFA64" box="[1090,1188,1412,1434]" pageId="2" pageNumber="153" type="holotype">Holotype</typeStatus>
|
||
: Berlin Museum.
|
||
<typeStatus id="E345880D611CFF92FA9070F73D82FA64" box="[1385,1436,1413,1434]" pageId="2" pageNumber="153">Type</typeStatus>
|
||
locality:
|
||
<collectingMunicipality id="DC25ACD5611CFF92FED970D1399BFA46" box="[288,389,1443,1464]" pageId="2" pageNumber="153">Huanusco</collectingMunicipality>
|
||
[=
|
||
<location id="39216074611CFF92FE5070D03A10FA46" LSID="urn:lsid:plazi:treatment:B45787B9611CFF95FF6E711339B3FBA6:39216074611CFF92FE5070D03A10FA46" box="[425,526,1442,1464]" municipality="Huanusco" name="Huatusco" pageId="2" pageNumber="153" stateProvince="Veracruz">Huatusco</location>
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],
|
||
<collectingRegion id="FE3AF84D611CFF92FDE470D13A64FA46" box="[541,634,1443,1464]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
|
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.
|
||
</materialsCitation>
|
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</paragraph>
|
||
<paragraph id="3C4136AF611CFF92FF6E70B03C70FA29" blockId="2.[151,1437,1228,1495]" box="[151,1134,1473,1495]" pageId="2" pageNumber="153">
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<materialsCitation id="8C963CF2611CFF92FF6E70B03C70FA29" ID-GBIF-Occurrence="1836204303" box="[151,1134,1473,1495]" country="Mexico" location="Not" pageId="2" pageNumber="153" specimenCount="1" typeStatus="holotype">
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<taxonomicName id="FBFE4D2C611CFF92FF6E70B039CCFA29" authority="Werner 1900" authorityName="Werner" authorityYear="1900" box="[151,466,1473,1495]" class="Reptilia" family="Colubridae" genus="Dirosema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="collare">
|
||
<emphasis id="0E8AEABD611CFF92FF6E70B03959FA29" box="[151,327,1473,1495]" italics="true" pageId="2" pageNumber="153">Dirosema collare</emphasis>
|
||
Werner 1900
|
||
</taxonomicName>
|
||
, Zool. Anz. 23:197.
|
||
<typeStatus id="E345880D611CFF92FD5A70B33B1BFA29" box="[675,773,1473,1495]" pageId="2" pageNumber="153" type="holotype">Holotype</typeStatus>
|
||
:
|
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<location id="39216074611CFF92FCF770B03B2AFA29" LSID="urn:lsid:plazi:treatment:B45787B9611CFF95FF6E711339B3FBA6:39216074611CFF92FCF770B03B2AFA29" box="[782,820,1474,1495]" country="Mexico" name="Not" pageId="2" pageNumber="153">Not</location>
|
||
traced. Type-locality:
|
||
<collectingCountry id="44E9763F611CFF92FBE070B33C77FA29" box="[1049,1129,1473,1495]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
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</paragraph>
|
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</subSubSection>
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<subSubSection id="74E46524611CFF92FF6E73763B83F89E" pageId="2" pageNumber="153" type="diagnosis">
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<paragraph id="3C4136AF611CFF92FF6E73763B83F89E" blockId="2.[151,1437,1540,1997]" pageId="2" pageNumber="153">
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<emphasis id="0E8AEABD611CFF92FF6E7376390FF9E3" bold="true" box="[151,273,1540,1565]" pageId="2" pageNumber="153">Diagnosis.</emphasis>
|
||
<taxonomicName id="FBFE4D2C611CFF92FED873763A5BF9E3" authorityName="Reinhardt" authorityYear="1861" box="[289,581,1540,1565]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="liebmanni">
|
||
<emphasis id="0E8AEABD611CFF92FED873763A5BF9E3" box="[289,581,1540,1565]" italics="true" pageId="2" pageNumber="153">Chersodromus liebmanni</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all Mexican species of snakes by having combination of prefrontals fused into single scale; postocular fused with supraocular; dorsal scales keeled in 17 rows at midbody, unreduced posteriorly; supralabials 7, third and fourth entering orbit; infralabials 7–8, usually 1– 5 contacting anterior chinshields (5 narrowly); mental contacting anterior chinshields; venter cream colored. This species may be distinguished from all species of
|
||
<taxonomicName id="FBFE4D2C611CFF92FCD073E63BD3F953" authorityName="Reinhardt" authorityYear="1861" box="[809,973,1684,1709]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="genus">
|
||
<emphasis id="0E8AEABD611CFF92FCD073E63BD3F953" box="[809,973,1684,1709]" italics="true" pageId="2" pageNumber="153">Chersodromus</emphasis>
|
||
</taxonomicName>
|
||
, except
|
||
<taxonomicName id="FBFE4D2C611CFF92FBCE73E73CFAF953" authorityName="Taylor" authorityYear="1949" box="[1079,1252,1684,1709]" class="Reptilia" family="Colubridae" genus="Schmidtophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="rubriventris">
|
||
<emphasis id="0E8AEABD611CFF92FBCE73E73CFAF953" box="[1079,1252,1684,1709]" italics="true" pageId="2" pageNumber="153">C. rubriventris</emphasis>
|
||
</taxonomicName>
|
||
, by having the postocular fused with supraocular; from
|
||
<taxonomicName id="FBFE4D2C611CFF92FD9A73CA3B15F92E" authorityName="Taylor" authorityYear="1949" box="[611,779,1719,1744]" class="Reptilia" family="Colubridae" genus="Schmidtophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="rubriventris">
|
||
<emphasis id="0E8AEABD611CFF92FD9A73CA3B15F92E" box="[611,779,1719,1744]" italics="true" pageId="2" pageNumber="153">C. rubriventris</emphasis>
|
||
</taxonomicName>
|
||
by having 17 scales around body (vs. 15), ventral surface uniformly cream (vs. bright orange) (
|
||
<tableCitation id="717C0314611CFF92FDCF73AE3A97F90A" box="[566,649,1756,1781]" captionStart="TABLE 1" captionStartId="14.[151,239,620,643]" captionText="TABLE 1. Select quatitative and qualitative features increased by habitat/distribution data from the four currently recognized species of Chersodromus." pageId="2" pageNumber="153">Table 1</tableCitation>
|
||
).
|
||
<taxonomicName id="FBFE4D2C611CFF92FD5873AF3C41F90B" authority="Baird & Girard (1853)" authorityName="Baird & Girard" authorityYear="1853" box="[673,1119,1756,1781]" class="Reptilia" family="Colubridae" genus="Ninia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="diademata">
|
||
<emphasis id="0E8AEABD611CFF92FD5873AF3B42F90B" box="[673,860,1756,1781]" italics="true" pageId="2" pageNumber="153">Ninia diademata</emphasis>
|
||
Baird & Girard (1853)
|
||
</taxonomicName>
|
||
might be confused with this species, and they are sympatric at some localities in
|
||
<collectingRegion id="FE3AF84D611CFF92FCD072723B63F8E6" box="[809,893,1792,1816]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="153">Oaxaca</collectingRegion>
|
||
and
|
||
<collectingRegion id="FE3AF84D611CFF92FC3A72723C34F8E9" box="[963,1066,1792,1815]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="153">Veracruz</collectingRegion>
|
||
.
|
||
<taxonomicName id="FBFE4D2C611CFF92FBC572733CE1F8E6" authorityName="Baird & Girard" authorityYear="1853" box="[1084,1279,1791,1816]" class="Reptilia" family="Colubridae" genus="Ninia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="species" species="diademata">
|
||
<emphasis id="0E8AEABD611CFF92FBC572733CE1F8E6" box="[1084,1279,1791,1816]" italics="true" pageId="2" pageNumber="153">Ninia diademata</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="FBFE4D2C611CFF92FF6E72563925F8C3" authorityName="Reinhardt" authorityYear="1861" box="[151,315,1828,1853]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="153" phylum="Chordata" rank="genus">
|
||
<emphasis id="0E8AEABD611CFF92FF6E72563925F8C3" box="[151,315,1828,1853]" italics="true" pageId="2" pageNumber="153">Chersodromus</emphasis>
|
||
</taxonomicName>
|
||
in having paired prefrontals shields, 19 strongly keeled dorsal scale rows, and two postoculars, larger body size (reaching at least
|
||
<quantity id="FB069B4A611CFF92FDEF723A3A77F8A1" box="[534,617,1864,1888]" metricMagnitude="0" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="153" unit="cm" value="330.0">330 cm</quantity>
|
||
TL), and more slender tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74E46524611CFF95FF3E721E3A73FC5B" lastPageId="5" lastPageNumber="156" pageId="2" pageNumber="153" type="description">
|
||
<paragraph id="3C4136AF611CFF92FF3E721E3A77F833" blockId="2.[151,1437,1540,1997]" pageId="2" pageNumber="153">
|
||
<emphasis id="0E8AEABD611CFF92FF3E721E3947F87B" bold="true" box="[199,345,1900,1925]" pageId="2" pageNumber="153">Description.</emphasis>
|
||
Small, slender snake; head moderately wider than the neck. Adults commonly reaching
|
||
<quantity id="FB069B4A611CFF92FAC3721E38DDF859" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="2" pageNumber="153" unit="mm" value="290.0" valueMax="330.0" valueMin="250.0">250–330 mm</quantity>
|
||
in total length with the largest known male
|
||
<quantity id="FB069B4A611CFF92FD5372E23B17F859" box="[682,777,1936,1960]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="153" unit="mm" value="310.0">310 mm</quantity>
|
||
, and the largest female
|
||
<quantity id="FB069B4A611CFF92FBF772E23C70F859" box="[1038,1134,1936,1960]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="153" unit="mm" value="330.0">330 mm</quantity>
|
||
. Tail comprises 16–22% of total length in males, 15–18% in females.
|
||
</paragraph>
|
||
<caption id="68816627611DFF93FF6E72ED38F8F80C" httpUri="https://zenodo.org/record/1206526/files/figure.png" pageId="3" pageNumber="154" startId="3.[151,250,1951,1973]" targetBox="[345,1246,224,1919]" targetPageId="3">
|
||
<paragraph id="3C4136AF611DFF93FF6E72ED38F8F80C" blockId="3.[151,1436,1951,2034]" pageId="3" pageNumber="154">
|
||
<emphasis id="0E8AEABD611DFF93FF6E72ED390CF84B" bold="true" box="[151,274,1951,1973]" pageId="3" pageNumber="154">FIGURE 2.</emphasis>
|
||
Dorsal (A), ventral (B) and lateral (C) views of the head (Scale = 3 mm); right maxilla (D, 2.61 mm); and from left to right: sulcate, lateral, and asulcate views of the hemipenis (E–G, Scale = 3.9 mm) of
|
||
<taxonomicName id="FBFE4D2C611DFF93FBBB72CF3D5AF82D" authorityName="Reinhardt" authorityYear="1861" box="[1090,1348,1981,2003]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="154" phylum="Chordata" rank="species" species="liebmanni">
|
||
<emphasis id="0E8AEABD611DFF93FBBB72CF3D5AF82D" box="[1090,1348,1981,2003]" italics="true" pageId="3" pageNumber="154">Chersodromus liebmanni</emphasis>
|
||
</taxonomicName>
|
||
(MZFC 28687).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="68816627611AFF94FF6E73453981F995" httpUri="https://zenodo.org/record/1206528/files/figure.png" pageId="4" pageNumber="155" startId="4.[151,250,1591,1613]" targetBox="[294,1299,179,1581]" targetPageId="4">
|
||
<paragraph id="3C4136AF611AFF94FF6E73453981F995" blockId="4.[151,1436,1591,1643]" pageId="4" pageNumber="155">
|
||
<emphasis id="0E8AEABD611AFF94FF6E73453906F9B3" bold="true" box="[151,280,1591,1613]" pageId="4" pageNumber="155">FIGURE 3.</emphasis>
|
||
Adult of
|
||
<taxonomicName id="FBFE4D2C611AFF94FE7073453A90F9B3" authorityName="Reinhardt" authorityYear="1861" box="[393,654,1591,1613]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="155" phylum="Chordata" rank="species" species="liebmanni">
|
||
<emphasis id="0E8AEABD611AFF94FE7073453A90F9B3" box="[393,654,1591,1613]" italics="true" pageId="4" pageNumber="155">Chersodromus liebmanni</emphasis>
|
||
</taxonomicName>
|
||
in life (MZFC 28687) from Totontepec, Oaxaca (A), and its locality in Cuautlapan, Veracruz (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3C4136AF611AFF94FF3E73E139FEF811" blockId="4.[151,1436,1683,2031]" pageId="4" pageNumber="155">
|
||
Prefrontals fused into a single, large scale. Supraoculars fused with postoculars. Frontal relatively broad and nasals divided. Small internasals 1/1, loreals 1/1, extending from the postnasal to the anterior edge of eye (no preoculars). No discrete postoculars are present (fused with supraocular). temporals 1+2 (rarely 1+1 owing to fusion of secondary series), supralabials 6–7 (usually 7) with 3–4 entering orbit, infralabials 6–8 (usually 7) with 1–4 contacting anterior chinshields (5 usually narrowly). Mental broader than long, first pair of infralabials not in contact along ventral midline. Anterior chinshields large with posterior pair of chinshields about one-third to onehalf the size and anterior pair. Dorsal scales keeled in 17-17-17 rows, rarely reduced by one either anteriorly or posteriorly. Preventrals 2 (rarely 1), ventrals
|
||
<quantity id="FB069B4A611AFF94FD7772FC3B13F859" box="[654,781,1934,1959]" metricMagnitude="0" metricUnit="m" metricValue="3.2638999999999996" metricValueMax="3.4543999999999997" metricValueMin="3.0734000000000004" pageId="4" pageNumber="155" unit="in" value="128.5" valueMax="136.0" valueMin="121.0">121–136 in</quantity>
|
||
males and
|
||
<quantity id="FB069B4A611AFF94FC7472FC3C12F859" box="[909,1036,1934,1959]" metricMagnitude="0" metricUnit="m" metricValue="3.3781999999999996" metricValueMax="3.556" metricValueMin="3.2004" pageId="4" pageNumber="155" unit="in" value="133.0" valueMax="140.0" valueMin="126.0">126–140 in</quantity>
|
||
females; divided subcaudals
|
||
<quantity id="FB069B4A611AFF94FAAF72FD38B3F832" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="10.668" metricValueMin="8.128" pageId="4" pageNumber="155" unit="in" value="37.0" valueMax="42.0" valueMin="32.0">32–42 in</quantity>
|
||
males and
|
||
<quantity id="FB069B4A611AFF94FEC672C139B6F832" box="[319,424,1971,1996]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.906" metricValueMin="7.874" pageId="4" pageNumber="155" unit="in" value="35.0" valueMax="39.0" valueMin="31.0">31–39 in</quantity>
|
||
females; dorsal scale rows at midlength of tail 6–8. Cloacal scute undivided. Pupil subcircular to vertically oval.
|
||
</paragraph>
|
||
<paragraph id="3C4136AF611BFF95FF3E75EA3BF7FDC3" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="156">A conspicuous yellow collar crosses the back of the head; the anterior border is irregular, usually having a middorsal forward extension to near the posterior tip of the frontal, extending on either side to involve about onehalf to two-thirds of the parietals, and laterally including the posterior portion of the anterior temporal and secondary temporals and portions of last several supralabials. Posterior border of the collar relatively straight, reaching the posterior edges of the parietals or nearly so, and extending posteriorly 0.5–1 scales. Dorsum dark gray to blackish and extends to the lateral portion of ventrals and subcaudals. A black head cap extends from the rostral to the anterior portion of parietals, covering internasals, a single fused prefrontal, supraoculars and fused postoculars, nasals, loreal, anterior portion of anterior temporal, and upper portion or edges of at least supralabials 3–5. The mental and infralabials 1–4 often have dark speckling or mottling. Belly immaculate cream or with varying amounts of dark mottling, especially along the midventer. Subcaudals usually have irregular dark mottling, especially along the midventer, and often become more heavily mottled posteriorly with distal portion of tail sometimes mostly dark. In a few individuals all subcaudals are mostly dark.</paragraph>
|
||
<paragraph id="3C4136AF611BFF95FF3E77353B44FD56" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="156">
|
||
<emphasis id="0E8AEABD611BFF95FF3E77353A30FD9E" bold="true" box="[199,558,583,608]" pageId="5" pageNumber="156">
|
||
Maxillary dentition (
|
||
<figureCitation id="A4C52A2A611BFF95FE4777353A01FD9E" box="[446,543,583,608]" captionStart="FIGURE2" captionStartId="3.[151,250,1951,1973]" captionTargetBox="[345,1246,224,1919]" captionTargetId="figure@3.[340,1246,215,1919]" captionTargetPageId="3" captionText="FIGURE2. Dorsal(A), ventral (B) and lateral (C)views of the head (Scale = 3 mm); rightmaxilla (D, 2.61 mm); and from left to right: sulcate, lateral, and asulcate views of the hemipenis (E–G, Scale = 3.9 mm) of Chersodromus liebmanni (MZFC 28687)." httpUri="https://zenodo.org/record/1206526/files/figure.png" pageId="5" pageNumber="156">Fig. 2 D</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Right maxilla extending anteriorly to level of suture between second and third supralabials, dorsoventrally compressed; in lateral view, anterior one-third curved dorsally; posterior end curved ventrally; maxillary teeth 7, slender, curved, longer posteriorly.
|
||
</paragraph>
|
||
<paragraph id="3C4136AF611BFF95FF3E77C63A73FC5B" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="156">
|
||
<emphasis id="0E8AEABD611BFF95FF3E77C63A9BFD35" bold="true" box="[199,645,692,717]" pageId="5" pageNumber="156">
|
||
Hemipenis morphology (
|
||
<figureCitation id="A4C52A2A611BFF95FE0977C63A68FD33" box="[496,630,692,717]" captionStart="FIGURE2" captionStartId="3.[151,250,1951,1973]" captionTargetBox="[345,1246,224,1919]" captionTargetId="figure@3.[340,1246,215,1919]" captionTargetPageId="3" captionText="FIGURE2. Dorsal(A), ventral (B) and lateral (C)views of the head (Scale = 3 mm); rightmaxilla (D, 2.61 mm); and from left to right: sulcate, lateral, and asulcate views of the hemipenis (E–G, Scale = 3.9 mm) of Chersodromus liebmanni (MZFC 28687)." httpUri="https://zenodo.org/record/1206526/files/figure.png" pageId="5" pageNumber="156">Fig. 2 E–G</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Retracted organ extends to level of tenth subcaudal, slightly bilobed, and semicapitate, sulcus spermaticus centrifugal with laterally expanded margins, ornamented with numerous spines of equal size, extending to tips of lobes; sulcus spermaticus bifurcates for about half length of hemipenial body; intrasulcar region and hemipenial body covered more-or-less homogeneously by spines, but on sulcate side of base, hook-shaped spine present on the right side of the hemipenis. In sulcate view, spines organized in inverted “V” shape before reaching capitation of hemipenis. On proximal region, two parallel folds protruding from body of hemipenis, ornamented by small spinules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74E46524611BFF95FF3776DD39B3FBA6" pageId="5" pageNumber="156" type="distribution">
|
||
<paragraph id="3C4136AF611BFF95FF3776DD39B3FBA6" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="156">
|
||
<emphasis id="0E8AEABD611BFF95FF3776DD3A1BFC36" bold="true" box="[206,517,943,968]" pageId="5" pageNumber="156">Distribution and habitat.</emphasis>
|
||
Inhabiting the Atlantic foothills and versant of west-central
|
||
<collectingRegion id="FE3AF84D611BFF95FB1C76C23D57FC39" box="[1253,1353,944,967]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="5" pageNumber="156">Veracruz</collectingRegion>
|
||
in the Orizaba region southward through the Sierra Zongolica to Sierra Negra in southeast
|
||
<collectingRegion id="FE3AF84D611BFF95FBB476A63C87FC13" box="[1101,1177,980,1005]" country="Mexico" name="Puebla" pageId="5" pageNumber="156">Puebla</collectingRegion>
|
||
and the Sierra Mixe in northern
|
||
<collectingRegion id="FE3AF84D611BFF95FF07768A394CFBEE" box="[254,338,1016,1040]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="156">Oaxaca</collectingRegion>
|
||
(
|
||
<figureCitation id="A4C52A2A611BFF95FE9D768539B9FBEE" box="[356,423,1015,1040]" captionStart="FIGURE 1" captionStartId="1.[151,250,1541,1563]" captionTargetBox="[151,1436,735,1519]" captionTargetId="figure@1.[151,1436,735,1519]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of the currently recognized species of the genus Chersodromus." httpUri="https://zenodo.org/record/1206524/files/figure.png" pageId="5" pageNumber="156">Fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="FBFE4D2C611BFF95FE3876853AC3FBEE" authorityName="Reinhardt" authorityYear="1861" box="[449,733,1015,1040]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="156" phylum="Chordata" rank="species" species="liebmanni">
|
||
<emphasis id="0E8AEABD611BFF95FE3876853AC3FBEE" box="[449,733,1015,1040]" italics="true" pageId="5" pageNumber="156">Chersodromus liebmanni</emphasis>
|
||
</taxonomicName>
|
||
inhabits upper rain forest and cloud forest (
|
||
<figureCitation id="A4C52A2A611BFF95FB2C76853D33FBEE" box="[1237,1325,1015,1040]" captionStart="FIGURE 3" captionStartId="4.[151,250,1591,1613]" captionTargetBox="[294,1299,179,1581]" captionTargetId="figure@4.[292,1299,179,1581]" captionTargetPageId="4" captionText="FIGURE 3. Adult of Chersodromus liebmanni in life (MZFC 28687) from Totontepec, Oaxaca (A), and its locality in Cuautlapan, Veracruz (B)." httpUri="https://zenodo.org/record/1206528/files/figure.png" pageId="5" pageNumber="156">Fig. 3B</figureCitation>
|
||
) between
|
||
<quantity id="FB069B4A611BFF95FF6E716E3928FBCA" box="[151,310,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="5" pageNumber="156" unit="m" value="1400.0" valueMax="1800.0" valueMin="1000.0">1000–1800 m</quantity>
|
||
above sea level (asl hereafter). Several specimens have been collected in agricultural fields and shade coffee plantations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |