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<document id="1EED87AA4FFD0BAE14A149A804F88FE1" ID-CLB-Dataset="30456" ID-DOI="10.11646/zootaxa.4399.2.1" ID-GBIF-Dataset="834234e8-ac96-4b19-a30e-3edc31fe4897" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206522" ID-ZooBank="43A975F2-05F6-4A29-A588-7787AF63589C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521899272936" checkinUser="plazi" docAuthor="Canseco-Márquez, Luis, Ramírez-González, Cynthia G. &amp; Campbell, Jonathan A." docDate="2018" docId="B45787B96116FF9BFF6E761539F1FF2A" docLanguage="en" docName="zootaxa.4399.2.1.pdf" docOrigin="Zootaxa 4399 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chersodromus australis Canseco-Márquez &amp; Ramírez-González &amp; Campbell 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="161" masterDocId="486EFFC1611EFF90FFF97572381EFFFE" masterDocTitle="Taxonomic review of the rare Mexican snake genus Chersodromus (Serpentes: Dipsadidae), with the description of two new species" masterLastPageNumber="169" masterPageNumber="151" pageNumber="159" updateTime="1698532892634" updateUser="ExternalLinkService">
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<mods:title id="A622C104CA20304091EE2143463FAACA">Taxonomic review of the rare Mexican snake genus Chersodromus (Serpentes: Dipsadidae), with the description of two new species</mods:title>
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<mods:namePart id="458D30A16ABAAA346E4D75403F884FDB">Canseco-Márquez, Luis</mods:namePart>
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<mods:namePart id="7CCD9EC140113AE2F907B0732A2476C8">Ramírez-González, Cynthia G.</mods:namePart>
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<mods:namePart id="DC09A2CE5E6B71B8BDF263A822B4F608">Campbell, Jonathan A.</mods:namePart>
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<mods:title id="1C82CF143BD40579ACED25E92C422391">Zootaxa</mods:title>
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<mods:date id="E1ECB2DCEED1B1524A93F29E1AB8EB65">2018</mods:date>
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<mods:number id="DDBDF7A1EA5BBAF33FAD2784941B34A7">2018-03-21</mods:number>
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<mods:number id="514C1587F2D38C4AC3B2D00F3D62E288">4399</mods:number>
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<treatment id="B45787B96116FF9BFF6E761539F1FF2A" ID-DOI="http://doi.org/10.5281/zenodo.5965351" ID-GBIF-Taxon="142524240" ID-Zenodo-Dep="5965351" LSID="urn:lsid:plazi:treatment:B45787B96116FF9BFF6E761539F1FF2A" httpUri="http://treatment.plazi.org/id/B45787B96116FF9BFF6E761539F1FF2A" lastPageId="11" lastPageNumber="161" pageId="8" pageNumber="159">
<subSubSection id="74E465246116FF98FF6E7615397CFC5D" pageId="8" pageNumber="159" type="nomenclature">
<paragraph id="3C4136AF6116FF98FF6E76153A03FC7C" blockId="8.[151,541,871,931]" box="[151,541,871,898]" pageId="8" pageNumber="159">
<heading id="670981C36116FF98FF6E76153A03FC7C" bold="true" box="[151,541,871,898]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
<emphasis id="0E8AEABD6116FF98FF6E76153A03FC7C" bold="true" box="[151,541,871,898]" pageId="8" pageNumber="159">
<taxonomicName id="FBFE4D2C6116FF98FF6E761539A6FC7F" authority="Canseco-Márquez &amp; Ramírez-González &amp; Campbell, 2018" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[151,440,871,897]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="australis" status="sp. nov.">
<emphasis id="0E8AEABD6116FF98FF6E761539A6FC7F" bold="true" box="[151,440,871,897]" italics="true" pageId="8" pageNumber="159">Chersodromus australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15B957C66116FF98FE46761A3A03FC7C" box="[447,541,872,898]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3C4136AF6116FF98FF6E76F8397CFC5D" blockId="8.[151,541,871,931]" box="[151,354,906,931]" pageId="8" pageNumber="159">
<figureCitation id="A4C52A2A6116FF98FF6E76F838C4FC5D" box="[151,218,906,931]" captionStart="FIGURE 6" captionStartId="9.[151,250,1971,1993]" captionTargetBox="[329,1262,511,1920]" captionTargetId="figure@9.[324,1262,499,1936]" captionTargetPageId="9" captionText="FIGURE 6. Dorsal (A), ventral (B), and lateral (C) views of the head (Scale = 3 mm); and its left maxilla (D, 2.43 mm) of the holotype of Chersodromus australis sp. nov. (MZFC 17618)." httpUri="https://zenodo.org/record/1206534/files/figure.png" pageId="8" pageNumber="159">Fig. 6</figureCitation>
,
<figureCitation id="A4C52A2A6116FF98FF1F76F93916FC5D" box="[230,264,907,931]" captionStart="FIGURE 7" captionStartId="10.[151,250,1374,1396]" captionTargetBox="[331,1262,185,1365]" captionTargetId="figure@10.[329,1271,181,1367]" captionTargetPageId="10" captionText="FIGURE 7. Holotype of Chersodromus australis sp. nov. (MZFC 17618) (A), and its type-locality in Chimalapas region of Oaxaca (B)." httpUri="https://zenodo.org/record/1206536/files/figure.png" pageId="8" pageNumber="159">7A</figureCitation>
<tableCitation id="717C03146116FF98FEF676F8397CFC5D" box="[271,354,906,931]" captionStart="TABLE 1" captionStartId="14.[151,239,620,643]" captionText="TABLE 1. Select quatitative and qualitative features increased by habitat/distribution data from the four currently recognized species of Chersodromus." pageId="8" pageNumber="159">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="74E465246116FF98FF6E76A03948FBC8" pageId="8" pageNumber="159" type="materials_examined">
<paragraph id="3C4136AF6116FF98FF6E76A03948FBC8" blockId="8.[151,1437,978,2014]" pageId="8" pageNumber="159">
<materialsCitation id="8C963CF26116FF98FF6E76A03948FBC8" ID-GBIF-Occurrence="1836204297" collectingDate="1995-06-27" collectionCode="MZFC" collectorName="An" country="Mexico" elevation="350" latitude="17.076517" location="San Isidro La Gringa" longLatPrecision="1" longitude="-94.064064" municipality="Municipality of Santa Maria Chimalapa" pageId="8" pageNumber="159" specimenCode="MZFC 17618" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis id="0E8AEABD6116FF98FF6E76A03915FC14" bold="true" box="[151,267,978,1003]" pageId="8" pageNumber="159">
<typeStatus id="E345880D6116FF98FF6E76A0391AFC15" box="[151,260,978,1003]" pageId="8" pageNumber="159" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="910B53796116FF98FEE876A1392DFC15" box="[273,307,979,1003]" pageId="8" pageNumber="159">An</collectorName>
adult female,
<specimenCode id="6C589ED46116FF98FE2B76A13A74FC15" box="[466,618,978,1003]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="8" pageNumber="159">MZFC 17618</specimenCode>
(field number EPR 388), collected by Edmundo Pérez Ramos on
<collectingDate id="5804E9876116FF98FABF76A138D1FBEC" pageId="8" pageNumber="159" value="1995-06-27">
<date id="4840106F6116FF98FABF76A138D1FBEC" pageId="8" pageNumber="159" value="1995-06-27">June 27 1995</date>
</collectingDate>
at
<location id="392160746116FF98FF0D768839FDFBEC" LSID="urn:lsid:plazi:treatment:B45787B96116FF9BFF6E761539F1FF2A:392160746116FF98FF0D768839FDFBEC" box="[244,483,1017,1042]" country="Mexico" latitude="17.076517" longLatPrecision="1" longitude="-94.064064" municipality="Municipality of Santa Maria Chimalapa" name="San Isidro La Gringa" pageId="8" pageNumber="159" stateProvince="Oaxaca">San Isidro La Gringa</location>
(
<geoCoordinate id="59CA50686116FF98FE0A768B3A8FFBEF" box="[499,657,1017,1042]" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="1" value="17.076517">17°04.591´N</geoCoordinate>
,
<geoCoordinate id="59CA50686116FF98FD6776883B23FBEF" box="[670,829,1018,1042]" direction="west" orientation="longitude" pageId="8" pageNumber="159" precision="1" value="-94.064064">94°03.844’ W</geoCoordinate>
;
<elevation id="B7D3D19C6116FF98FCB0768B3BA2FBEC" box="[841,956,1017,1042]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="159" unit="m" value="350.0">
<quantity id="FB069B4A6116FF98FCB0768B3B8CFBEF" box="[841,914,1017,1042]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="159" unit="m" value="350.0">350 m</quantity>
asl
</elevation>
),
<collectingMunicipality id="DC25ACD56116FF98FC28768B3D86FBEC" box="[977,1432,1017,1042]" pageId="8" pageNumber="159">Municipality of Santa María Chimalapa</collectingMunicipality>
,
<collectingRegion id="FE3AF84D6116FF98FF6E716C38F0FBC8" box="[151,238,1054,1078]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="159">Oaxaca</collectingRegion>
,
<collectingCountry id="44E9763F6116FF98FF00716F394CFBC8" box="[249,338,1053,1078]" name="Mexico" pageId="8" pageNumber="159">Mexico</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="74E465246116FF98FF3E71333C3CFA60" pageId="8" pageNumber="159" type="diagnosis">
<paragraph id="3C4136AF6116FF98FF3E71333ACBFB14" blockId="8.[151,1437,978,2014]" pageId="8" pageNumber="159">
<emphasis id="0E8AEABD6116FF98FF3E7133395FFBA4" bold="true" box="[199,321,1089,1114]" pageId="8" pageNumber="159">Diagnosis.</emphasis>
<taxonomicName id="FBFE4D2C6116FF98FEB771333A7CFBA4" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[334,610,1089,1114]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="australis">
<emphasis id="0E8AEABD6116FF98FEB771333A7CFBA4" box="[334,610,1089,1114]" italics="true" pageId="8" pageNumber="159">Chersodromus australis</emphasis>
</taxonomicName>
can be distinguished from all Mexican species of snakes by having combination of prefrontals fused into single scale; postocular discrete from supraocular; single anterior temporal present; dorsal scales keeled in 17 rows at midbody, unreduced posteriorly; supralabials usually 6, third and fourth entering orbit; infralabials 7–8, usually 1–4 contacting anterior chinshields); mental contacting anterior chinshields; ventral surface of body cream colored.
</paragraph>
<paragraph id="3C4136AF6116FF98FF3E71873C3CFA60" blockId="8.[151,1437,978,2014]" pageId="8" pageNumber="159">
<taxonomicName id="FBFE4D2C6116FF98FF3E718739C8FAF0" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[199,470,1269,1294]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="australis">
<emphasis id="0E8AEABD6116FF98FF3E718739C8FAF0" box="[199,470,1269,1294]" italics="true" pageId="8" pageNumber="159">Chersodromus australis</emphasis>
</taxonomicName>
differs of
<taxonomicName id="FBFE4D2C6116FF98FDA971843AE6FAF0" baseAuthorityName="Taylor" baseAuthorityYear="1949" box="[592,760,1269,1294]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="rubriventris">
<emphasis id="0E8AEABD6116FF98FDA971843AE6FAF0" box="[592,760,1269,1294]" italics="true" pageId="8" pageNumber="159">C. rubriventris</emphasis>
</taxonomicName>
by having 17 scales around body (vs. 15), by having eight infralabials (vs. six or seven), by having a cream colored belly (vs. orange), and by having the mental in contact with first pair of chinshields (vs. not in contact).
<taxonomicName id="FBFE4D2C6116FF98FD3D704F3BCFFAA8" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[708,977,1341,1366]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="australis">
<emphasis id="0E8AEABD6116FF98FD3D704F3BCFFAA8" box="[708,977,1341,1366]" italics="true" pageId="8" pageNumber="159">Chersodromus australis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="FBFE4D2C6116FF98FB90704C3CE5FAA8" authorityName="Reinhardt" authorityYear="1861" box="[1129,1275,1341,1366]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="liebmanni">
<emphasis id="0E8AEABD6116FF98FB90704C3CE5FAA8" box="[1129,1275,1341,1366]" italics="true" pageId="8" pageNumber="159">C. liebmanni</emphasis>
</taxonomicName>
by having six supralabials (vs. seven), and from the new species that follows by having a dark belly (vs. cream-colored) and in having four infralabials contacting anterior chinshields (vs. five or six) (
<tableCitation id="717C03146116FF98FC4470F73C0FFA60" box="[957,1041,1413,1438]" captionStart="TABLE 1" captionStartId="14.[151,239,620,643]" captionText="TABLE 1. Select quatitative and qualitative features increased by habitat/distribution data from the four currently recognized species of Chersodromus." pageId="8" pageNumber="159">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="74E465246116FF9AFF3E70DB3A8BF906" lastPageId="10" lastPageNumber="161" pageId="8" pageNumber="159" type="description">
<paragraph id="3C4136AF6116FF99FF3E70DB3BF9FE53" blockId="8.[151,1437,978,2014]" lastBlockId="9.[151,1437,151,429]" lastPageId="9" lastPageNumber="160" pageId="8" pageNumber="159">
<emphasis id="0E8AEABD6116FF98FF3E70DB3A13FA3C" bold="true" box="[199,525,1449,1474]" pageId="8" pageNumber="159">
Description of the
<typeStatus id="E345880D6116FF98FE5E70DB3A13FA3C" box="[423,525,1449,1474]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
</emphasis>
. Adult female; head length
<quantity id="FB069B4A6116FF98FCB570D83BB8FA3F" box="[844,934,1450,1474]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="8" pageNumber="159" unit="mm" value="7.3">7.3 mm</quantity>
; snout-vent length
<quantity id="FB069B4A6116FF98FB7970D83CFFFA3F" box="[1152,1249,1450,1474]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="159" unit="mm" value="180.0">180 mm</quantity>
; tail
<quantity id="FB069B4A6116FF98FAE370D83D73FA3F" box="[1306,1389,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="159" unit="mm" value="40.0">40 mm</quantity>
; the tail comprises 22% of total length; head slightly distinct from neck; snout long, 2.6 times head length, rounded in dorsal aspect, projecting anteriorly beyond lower jaw; rostral 0.6 times broader than high, 1.9 times longer than internasal suture, with posterior end approximately at level of anterior margin of nares; internasals broader than long (length vs width on L/R sides 0.59/0.56), rounded anteriorly, in contact laterally with anterior and posterior nasals; prefrontals fused together, posterior corner contacting eye; postnasal plus loreal 0.8 times length of snout, suture between them 0.2 times frontal length; frontal longer than wide (width/length 1.1); supraocular large, in broad contact with prefrontal and parietal, contacting postocular and frontal 1.0 times as long as horizontal diameter of eye; supraocular 1.0 times as long as loreal, forming about posterior half of dorsal margin of orbit, ventral margin not projecting anteriorly beyond margin of orbit, posteriorly exceeding 0.4 times length of posterior margin of orbit; parietals 1.4 times longer than broad, approximately 0.4 times head length, parietal suture 0.8 times as long as frontal; one postocular on right side, two on left side; nasal divided; postnasal 1.0 times as long as prenasal; combined length of prenasal and postnasal 1.8 times longer than loreal length; loreal 1.1 times wider that long, 2.6 times snout length, 1.0 times longer than horizontal diameter of eye, dorsal margin straight and ventral margin concave; eye small, 2.4 times snout length, vertical diameter 0.6 times its distance from lip; supralabials six, first and second in contact with postnasal, second and third in contact with loreal, third and fourth entering orbit, fifth largest, contacting anterior temporal; one anterior temporal in broad contact with fifth supralabial; two posterior temporals separating sixth supralabial from parietal; upper posterior temporal larger that lower; posterior temporals separated from each other by five nuchals contacting parietals; mental rounded anterior, 2.2 times broader than long, contacting anterior chinshields; infralabials eight, first four contacting anterior chinshields, fourth slightly and fifth in broad contact with posterior chinshields; anterior chinshields longer than broad (2.1/1.8), 1.5 times longer than posterior chinshields; posterior chinshields broadly contacting anterior chinshields; one preventral scale; infralabials and scales in chin region smooth; dorsals in 17-17-17 rows, keeled throughout body, without apical pits; ventrals 131; cloacal scute single; paired subcaudals 36.
</paragraph>
<caption id="688166276117FF99FF6E72C13B1AF816" httpUri="https://zenodo.org/record/1206534/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1971,1993]" targetBox="[329,1262,511,1920]" targetPageId="9">
<paragraph id="3C4136AF6117FF99FF6E72C13B1AF816" blockId="9.[151,1435,1971,2024]" pageId="9" pageNumber="160">
<emphasis id="0E8AEABD6117FF99FF6E72C1390DF837" bold="true" box="[151,275,1971,1993]" pageId="9" pageNumber="160">FIGURE 6.</emphasis>
Dorsal (A), ventral (B), and lateral (C) views of the head (Scale = 3 mm); and its left maxilla (D, 2.43 mm) of the holotype of
<taxonomicName id="FBFE4D2C6117FF99FEE872A03A1BF816" authority="Canseco-Márquez &amp; Ramírez-González &amp; Campbell, 2018" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[273,517,2002,2024]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="160" phylum="Chordata" rank="species" species="australis" status="sp. nov.">
<emphasis id="0E8AEABD6117FF99FEE872A03A1BF816" box="[273,517,2002,2024]" italics="true" pageId="9" pageNumber="160">Chersodromus australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15B957C66117FF99FDF272A03A45F816" box="[523,603,2002,2024]" pageId="9" pageNumber="160" rank="species">
<emphasis id="0E8AEABD6117FF99FDF272A03A45F816" bold="true" box="[523,603,2002,2024]" pageId="9" pageNumber="160">sp. nov.</emphasis>
</taxonomicNameLabel>
(MZFC 17618).
</paragraph>
</caption>
<caption id="688166276114FF9AFF6E702C390FFA6C" httpUri="https://zenodo.org/record/1206536/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1374,1396]" targetBox="[331,1262,185,1365]" targetPageId="10">
<paragraph id="3C4136AF6114FF9AFF6E702C390FFA6C" blockId="10.[151,1436,1374,1426]" pageId="10" pageNumber="161">
<emphasis id="0E8AEABD6114FF9AFF6E702C390AFA8A" bold="true" box="[151,276,1374,1396]" pageId="10" pageNumber="161">FIGURE 7.</emphasis>
Holotype of
<taxonomicName id="FBFE4D2C6114FF9AFE59702C3A8BFA8A" authority="Canseco-Márquez &amp; Ramírez-González &amp; Campbell, 2018" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[416,661,1374,1396]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="australis" status="sp. nov.">
<emphasis id="0E8AEABD6114FF9AFE59702C3A8BFA8A" box="[416,661,1374,1396]" italics="true" pageId="10" pageNumber="161">Chersodromus australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15B957C66114FF9AFD65702C3AF0FA8A" box="[668,750,1374,1396]" pageId="10" pageNumber="161" rank="species">
<emphasis id="0E8AEABD6114FF9AFD65702C3AF0FA8A" bold="true" box="[668,750,1374,1396]" pageId="10" pageNumber="161">sp. nov.</emphasis>
</taxonomicNameLabel>
(MZFC 17618) (A), and its type-locality in Chimalapas region of Oaxaca (B).
</paragraph>
</caption>
<paragraph id="3C4136AF6114FF9AFF3E70CD3D5EF999" blockId="10.[151,1436,1471,2036]" pageId="10" pageNumber="161">Color (alcohol after formalin). Dorsal ground color black, including lateral borders of ventral scales; dorsum of head with a white band crossing posterior region of parietals, including all of fifth and sixth supralabials, middle of fourth, and lower part of supralabials three and four; head, first three and portion of fourth supralabial black; mental, first three and portion of fourth infralabial black; lateral edges of ventrals colored similarly to dorsum; remaining ventral surface, including throat, cream; subcaudals cream with few scattered small black spots.</paragraph>
<paragraph id="3C4136AF6114FF9AFF3E73013A8BF906" blockId="10.[151,1436,1471,2036]" pageId="10" pageNumber="161">
<emphasis id="0E8AEABD6114FF9AFF3E73013A31F972" bold="true" box="[199,559,1651,1676]" pageId="10" pageNumber="161">
Maxillary dentition (
<figureCitation id="A4C52A2A6114FF9AFE3A73013A3FF972" box="[451,545,1651,1676]" captionStart="FIGURE 6" captionStartId="9.[151,250,1971,1993]" captionTargetBox="[329,1262,511,1920]" captionTargetId="figure@9.[324,1262,499,1936]" captionTargetPageId="9" captionText="FIGURE 6. Dorsal (A), ventral (B), and lateral (C) views of the head (Scale = 3 mm); and its left maxilla (D, 2.43 mm) of the holotype of Chersodromus australis sp. nov. (MZFC 17618)." httpUri="https://zenodo.org/record/1206534/files/figure.png" pageId="10" pageNumber="161">Fig. 6D</figureCitation>
).
</emphasis>
Left maxilla extending anteriorly to the level of suture between second and third supralabials; dorsoventrally compressed; in lateral view with dorsal flange at about midlevel, tapering at anterior and posterior ends; anterior portion of edentate, followed by seven long, slender, curved teeth; anterior five teeth subequal, posterior two slightly smaller.
</paragraph>
</subSubSection>
<subSubSection id="74E465246114FF9AFF3E72713AD3F8BE" pageId="10" pageNumber="161" type="etymology">
<paragraph id="3C4136AF6114FF9AFF3E72713AD3F8BE" blockId="10.[151,1436,1471,2036]" pageId="10" pageNumber="161">
<emphasis id="0E8AEABD6114FF9AFF3E72713958F8E2" bold="true" box="[199,326,1795,1820]" pageId="10" pageNumber="161">Etymology</emphasis>
. The specific name is derived from the Latin
<taxonomicName id="FBFE4D2C6114FF9AFC7472713BF1F8E2" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[909,1007,1795,1820]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="australis">
<emphasis id="0E8AEABD6114FF9AFC7472713BF1F8E2" box="[909,1007,1795,1820]" italics="true" pageId="10" pageNumber="161">australis</emphasis>
</taxonomicName>
, meaning southern, alluding to the southernmost distribution of species for the genus.
</paragraph>
</subSubSection>
<subSubSection id="74E465246114FF9BFF3E723939F1FF2A" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="distribution">
<paragraph id="3C4136AF6114FF9BFF3E723939F1FF2A" blockId="10.[151,1436,1471,2036]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<emphasis id="0E8AEABD6114FF9AFF3E723939E9F89A" bold="true" box="[199,503,1867,1892]" pageId="10" pageNumber="161">Distribution and habitat.</emphasis>
<taxonomicName id="FBFE4D2C6114FF9AFDF972393B11F89A" authorityName="Canseco-Márquez &amp; Ramírez-González &amp; Campbell" authorityYear="2018" box="[512,783,1867,1892]" class="Reptilia" family="Colubridae" genus="Chersodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="australis">
<emphasis id="0E8AEABD6114FF9AFDF972393B11F89A" box="[512,783,1867,1892]" italics="true" pageId="10" pageNumber="161">Chersodromus australis</emphasis>
</taxonomicName>
apparently has a relatively small distribution east of the Isthmus of Tehuantepec (
<figureCitation id="A4C52A2A6114FF9AFE48721D39ECF876" box="[433,498,1903,1928]" captionStart="FIGURE 1" captionStartId="1.[151,250,1541,1563]" captionTargetBox="[151,1436,735,1519]" captionTargetId="figure@1.[151,1436,735,1519]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of the currently recognized species of the genus Chersodromus." httpUri="https://zenodo.org/record/1206524/files/figure.png" pageId="10" pageNumber="161">Fig. 1</figureCitation>
) in the region known locally as Chimalapas; it occurs at low elevations (
<quantity id="FB069B4A6114FF9AFADD721D3D73F879" box="[1316,1389,1903,1928]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="161" unit="m" value="350.0">350 m</quantity>
asl) in tropical rain forest (
<figureCitation id="A4C52A2A6114FF9AFE6772E139E2F852" box="[414,508,1939,1964]" captionStart="FIGURE 7" captionStartId="10.[151,250,1374,1396]" captionTargetBox="[331,1262,185,1365]" captionTargetId="figure@10.[329,1271,181,1367]" captionTargetPageId="10" captionText="FIGURE 7. Holotype of Chersodromus australis sp. nov. (MZFC 17618) (A), and its type-locality in Chimalapas region of Oaxaca (B)." httpUri="https://zenodo.org/record/1206536/files/figure.png" pageId="10" pageNumber="161">Fig 7 B</figureCitation>
). A single specimen was obtained on
<date id="4840106F6114FF9AFC4072E63C31F852" box="[953,1071,1939,1964]" pageId="10" pageNumber="161" value="1995-06">June 1995</date>
under a rock at 16:00 hrs on a mountain near San Isidro la Gringa, municipality of Santa María Chimalapa. Other amphibians and reptiles recorded at the
<typeStatus id="E345880D6114FF9AFEBD72AE396AF80A" box="[324,372,2012,2036]" pageId="10" pageNumber="161">type</typeStatus>
locality included
<taxonomicName id="FBFE4D2C6114FF9AFDC772A93C42F80A" authority="(Firchein &amp; Smith 1957)" baseAuthorityName="Firchein &amp; Smith" baseAuthorityYear="1957" box="[574,1116,2011,2036]" class="Amphibia" family="Bufonidae" genus="Incilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="macrocristatus">
<emphasis id="0E8AEABD6114FF9AFDC772A93B5EF80A" box="[574,832,2011,2036]" italics="true" pageId="10" pageNumber="161">Incilius macrocristatus</emphasis>
(Firchein &amp; Smith 1957)
</taxonomicName>
,
<taxonomicName id="FBFE4D2C6114FF9AFB9272AF3D2DF80A" box="[1131,1331,2013,2036]" class="Reptilia" family="Dactyloidae" genus="Norops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="0E8AEABD6114FF9AFB9272AF3D2DF80A" box="[1131,1331,2013,2036]" italics="true" pageId="10" pageNumber="161">Norops pygmaeus</emphasis>
</taxonomicName>
(Álvarez del Toro &amp; Smith 1956),
<taxonomicName id="FBFE4D2C6115FF9BFE3075E53BFDFF4E" authority="Werler &amp; Shannon 1957" authorityName="Werler &amp; Shannon" authorityYear="1957" box="[457,995,151,176]" class="Reptilia" family="Xantusiidae" genus="Lepidophyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="tuxtlae">
<emphasis id="0E8AEABD6115FF9BFE3075E53AA7FF4E" box="[457,697,151,176]" italics="true" pageId="11" pageNumber="162">Lepidophyma tuxtlae</emphasis>
Werler &amp; Shannon 1957
</taxonomicName>
,
<taxonomicName id="FBFE4D2C6115FF9BFC0C75E53D7FFF4E" authority="(Garman 1884)" baseAuthorityName="Garman" baseAuthorityYear="1884" box="[1013,1377,151,176]" class="Reptilia" family="Viperidae" genus="Bothrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="asper">
<emphasis id="0E8AEABD6115FF9BFC0C75E53CBAFF4E" box="[1013,1188,151,176]" italics="true" pageId="11" pageNumber="162">Bothrops asper</emphasis>
(Garman 1884)
</taxonomicName>
, and
<taxonomicName id="FBFE4D2C6115FF9BFF6E75CE39F5FF2A" authority="(Smith 1940)" baseAuthorityName="Smith" baseAuthorityYear="1940" box="[151,491,188,213]" class="Reptilia" family="Colubridae" genus="Tantillita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="lintoni">
<emphasis id="0E8AEABD6115FF9BFF6E75CE3953FF2B" box="[151,333,188,213]" italics="true" pageId="11" pageNumber="162">Tantillita lintoni</emphasis>
(Smith 1940)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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