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<document id="18EC2DE641EEA8B5CC1440B14DD27EA0" ID-DOI="10.11646/zootaxa.3889.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4952403" ID-ZooBank="1F3F076D-A4F1-4B4A-977D-4B99581C6779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730508465" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2014" docId="A7173150FC5EFF9FFECC006482BC9AAE" docLanguage="en" docName="zt03889p484.pdf" docOrigin="Zootaxa 3889 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ogyges menchuae Cano 2014, new species" docType="treatment" docVersion="3" lastPageNumber="467" masterDocId="5B2E4928FC50FF8FFE5B063880699A56" masterDocTitle="Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly" masterLastPageNumber="484" masterPageNumber="451" pageNumber="465" updateTime="1698967030688" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="3002A91DC8AB13F6DE5CA81D554033BD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1CA7D2BD7BA815D159D8650ADE65C08">
<mods:title id="A77904C1A1643962F626146029228CA0">Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly</mods:title>
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<mods:name id="491BF4E58298816FAEB435BA8996F88B" type="personal">
<mods:role id="FA2A9F6A574ECF16F9E1F1D944FECD71">
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<mods:namePart id="A52190A36BE7CB0B91D686B8B11B6082">Cano, Enio B.</mods:namePart>
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<mods:typeOfResource id="B4DE3F370BA18B0195B6B67FC48B7BCF">text</mods:typeOfResource>
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<mods:title id="0D78DB46D7F3382AAE31A2C3EAD216FB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="405D1E3069E88707EB0A7AA70A2774A6">
<mods:date id="79F99FCB5406109ADEE6187AB88B76EF">2014</mods:date>
<mods:detail id="FA5D7A5AB03542479608656166F36950" type="pubDate">
<mods:number id="1AEDACE332B27DD22C463B1CAE69F4A3">2014-12-03</mods:number>
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<mods:detail id="09EA950023D29B3F476800B716E027D9" type="volume">
<mods:number id="C6218B848115C309F3CFFE5225322B91">3889</mods:number>
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<mods:number id="FB37DEFEEBC2FE78178A25C4A128F6B9">4</mods:number>
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<mods:classification id="0AE3B22364D0428CCFFEEBFD24B0CAD9">journal article</mods:classification>
<mods:identifier id="E3050B494FF207B911AE4ED7BE520DCB" type="DOI">10.11646/zootaxa.3889.4.1</mods:identifier>
<mods:identifier id="2C5C6C040C9FD6E4AB55CAD6292BE7D5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7F79EAD287BBD6838111F6890234344D" type="Zenodo-Dep">4952403</mods:identifier>
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<treatment id="A7173150FC5EFF9FFECC006482BC9AAE" LSID="urn:lsid:plazi:treatment:A7173150FC5EFF9FFECC006482BC9AAE" httpUri="http://treatment.plazi.org/id/A7173150FC5EFF9FFECC006482BC9AAE" lastPageId="16" lastPageNumber="467" pageId="14" pageNumber="465">
<subSubSection id="67A4D3CDFC5EFF81FECC006482359C20" box="[151,604,1628,1654]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="2F018046FC5EFF81FECC006482359C20" blockId="14.[151,604,1628,1687]" box="[151,604,1628,1654]" pageId="14" pageNumber="465">
<heading id="7449372AFC5EFF81FECC006482359C20" bold="true" box="[151,604,1628,1654]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<emphasis id="1DCA5C54FC5EFF81FECC006482359C20" bold="true" box="[151,604,1628,1654]" pageId="14" pageNumber="465">
<taxonomicName id="E8BEFBC5FC5EFF81FECC006481A99C20" authority="Cano" authorityName="Cano" authorityYear="2014" box="[151,448,1628,1654]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="menchuae" status="sp. nov.">
<emphasis id="1DCA5C54FC5EFF81FECC0064811A9C20" bold="true" box="[151,371,1628,1654]" italics="true" pageId="14" pageNumber="465">Ogyges menchuae</emphasis>
Cano
</taxonomicName>
,
<taxonomicNameLabel id="06F9E12FFC5EFF81FF97006482359C20" box="[460,604,1628,1654]" pageId="14" pageNumber="465" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF81FECC0046813B9CC1" box="[151,338,1662,1687]" pageId="14" pageNumber="465" type="description">
<paragraph id="2F018046FC5EFF81FECC0046813B9CC1" blockId="14.[151,604,1628,1687]" box="[151,338,1662,1687]" pageId="14" pageNumber="465">
<figureCitation id="B7859CC3FC5EFF81FECC004681749CC1" box="[151,285,1662,1687]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="16.[151,250,1926,1948]" captionStartId-1="17.[151,250,1945,1967]" captionTargetBox-0="[159,1436,310,1896]" captionTargetBox-1="[162,1427,206,1916]" captionTargetId-0="figure-89@16.[151,1436,301,1905]" captionTargetId-1="figure-45@17.[151,1436,193,1924]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 11. Paratype of Ogyges menchuae: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 12. Ogyges menchuae: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952433" figureDoi-1="http://doi.org/10.5281/zenodo.4952435" httpUri-0="https://zenodo.org/record/4952433/files/figure.png" httpUri-1="https://zenodo.org/record/4952435/files/figure.png" pageId="14" pageNumber="465">Figs. 11–12</figureCitation>
,
<figureCitation id="B7859CC3FC5EFF81FF730046813B9CC1" box="[296,338,1662,1687]" captionStart="FIGURE 21" captionStartId="28.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure-318@28.[151,1436,193,1354]" captionTargetPageId="28" captionText="FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili." figureDoi="http://doi.org/10.5281/zenodo.4952455" httpUri="https://zenodo.org/record/4952455/files/figure.png" pageId="14" pageNumber="465">21p</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF81FECC00FE84ED9D39" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="2F018046FC5EFF81FECC00FE84ED9D39" blockId="14.[151,1437,1734,2011]" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FECC00FE81789C89" bold="true" box="[151,273,1734,1759]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">Diagnosis.</emphasis>
<taxonomicName id="E8BEFBC5FC5EFF81FF4200FF818E9C88" authorityName="Cano" authorityYear="2014" box="[281,487,1734,1758]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="menchuae">
<emphasis id="1DCA5C54FC5EFF81FF4200FF818E9C88" box="[281,487,1734,1758]" italics="true" pageId="14" pageNumber="465">Ogyges menchuae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="E8BEFBC5FC5EFF81FCE700FF84FF9C89" authority="Schuster &amp; Reyes-Castillo" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" box="[700,1174,1734,1759]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cakchiqueli">
<emphasis id="1DCA5C54FC5EFF81FCE700FF83099C88" box="[700,864,1734,1758]" italics="true" pageId="14" pageNumber="465">O. cakchiqueli</emphasis>
Schuster &amp; Reyes-Castillo
</taxonomicName>
,
<taxonomicName id="E8BEFBC5FC5EFF81FAF900FF85FE9C89" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[1186,1431,1734,1759]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="1DCA5C54FC5EFF81FAF900FF85529C88" box="[1186,1339,1734,1758]" italics="true" pageId="14" pageNumber="465">O. championi</emphasis>
(Bates)
</taxonomicName>
, and
<taxonomicName id="E8BEFBC5FC5EFF81FE9000D382BD9D55" authority="Schuster &amp; Reyes-Castillo. These" authorityName="Schuster &amp; Reyes-Castillo. These" authorityYear="1990" box="[203,724,1770,1795]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="kekchii">
<emphasis id="1DCA5C54FC5EFF81FE9000D381299D55" box="[203,320,1771,1795]" italics="true" pageId="14" pageNumber="465">O. kekchii</emphasis>
Schuster &amp; Reyes-Castillo. These
</taxonomicName>
species are in the “
<taxonomicName id="E8BEFBC5FC5EFF81FD9F00D384539D55" box="[964,1082,1771,1795]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="laevior">
<emphasis id="1DCA5C54FC5EFF81FD9F00D383B19D54" box="[964,984,1771,1794]" italics="true" pageId="14" pageNumber="465">O</emphasis>
.
<emphasis id="1DCA5C54FC5EFF81FDB000D384539D55" box="[1003,1082,1771,1795]" italics="true" pageId="14" pageNumber="465">laevior</emphasis>
</taxonomicName>
” [=
<taxonomicName id="E8BEFBC5FC5EFF81FA2D00D3857A9D55" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[1142,1299,1771,1795]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="1DCA5C54FC5EFF81FA2D00D384E39D54" box="[1142,1162,1771,1794]" italics="true" pageId="14" pageNumber="465">O</emphasis>
.
<emphasis id="1DCA5C54FC5EFF81FAC600D3857A9D55" box="[1181,1299,1771,1795]" italics="true" pageId="14" pageNumber="465">championi</emphasis>
</taxonomicName>
] lineage of Schuster &amp; Reyes Castillo (1990). The large internal tubercles separate it from similar species, except from
<taxonomicName id="E8BEFBC5FC5EFF81FBDA013782409D1D" authority="Schuster &amp; Reyes-Castillo" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="tzutuhili">
<emphasis id="1DCA5C54FC5EFF81FBDA0137809A9D1D" italics="true" pageId="14" pageNumber="465">O. tzutuhili</emphasis>
Schuster &amp; Reyes-Castillo
</taxonomicName>
and
<taxonomicName id="E8BEFBC5FC5EFF81FC38010B842B9D1D" authority="Reyes-Castillo &amp; Castillo" authorityName="Reyes-Castillo &amp; Castillo" authorityYear="1986" box="[611,1090,1842,1867]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="marilucasae">
<emphasis id="1DCA5C54FC5EFF81FC38010B837B9D1D" box="[611,786,1843,1867]" italics="true" pageId="14" pageNumber="465">O. marilucasae</emphasis>
Reyes-Castillo &amp; Castillo
</taxonomicName>
, but is easily separated by the narrow antennal club (very wide in
<taxonomicName id="E8BEFBC5FC5EFF81FC7D016F82CC9D38" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" box="[550,677,1878,1902]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="tzutuhili">
<emphasis id="1DCA5C54FC5EFF81FC7D016F82539D38" box="[550,570,1879,1902]" italics="true" pageId="14" pageNumber="465">O</emphasis>
.
<emphasis id="1DCA5C54FC5EFF81FC12016E82CC9D38" box="[585,677,1878,1902]" italics="true" pageId="14" pageNumber="465">tzutuhili</emphasis>
</taxonomicName>
and
<taxonomicName id="E8BEFBC5FC5EFF81FC80016F83EE9D38" authorityName="Reyes-Castillo &amp; Castillo" authorityYear="1986" box="[731,903,1878,1902]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="marilucasae">
<emphasis id="1DCA5C54FC5EFF81FC80016F83EE9D38" box="[731,903,1878,1902]" italics="true" pageId="14" pageNumber="465">O. marilucasae</emphasis>
</taxonomicName>
), and the smaller size.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF80FE9C014285FA9C23" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465" type="description">
<paragraph id="2F018046FC5EFF80FE9C0142851199D6" blockId="14.[151,1437,1734,2011]" lastBlockId="15.[151,1437,151,2018]" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FE9C014281309DC5" bold="true" box="[199,345,1914,1939]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">Description.</emphasis>
<typeStatus id="F0053EE4FC5EFF81FF3E014281A49DC5" box="[357,461,1914,1939]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
male, black adult. Total length 29.24, elytral length 16.20, pronotal length 8.38, pronotal width 10.61, humeral width 10.03. Head: anterior border of labrum straight, dorsal surface granular. Clypeus vertical, anterior border straight, posteriorly microgranulate, separated from the mediofrontal area by an inflection; mediofrontal tubercles conical, with apex rounded, moderately enlarged and markedly visible in lateral view. Mediofrontal area slightly microtuberculate, moderately roughened; internal tubercles present and well developed, directed forward and slightly upward, connected to the curved and strongly marked posterofrontal ridges; angle of posterofrontal ridges concave and united to the central horn. Laterofrontal areas roughened, with sparse granulations. Lateropostfrontal areas glabrous, smooth, somewhat roughened and with sparse microtubercles or granulations. Central horn short with apex conical, not free and with median longitudinal (mostly deep) groove dorsally in basal half; lateroposterior tubercles elongate, rounded, directed forward, separated from the central horn by a small groove with sparse or moderate microgranulations. Postfrontal groove deeper at sides and separated from lateropostfrontal areas. Supraorbital ridge with mostly unequal or almost equal anterior tubercles; posterior 1/2 bifurcate, with the supraorbital fossae very deep; external ridge well marked. Ocular canthus with apex swollen, apex forming a right angle. Eyes reduced, width =
<quantity id="E8462DA3FC5FFF80FA7107C784F89841" box="[1066,1169,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="466" unit="mm" value="0.54">0.54 mm</quantity>
(each eye). Interocular width =
<quantity id="E8462DA3FC5FFF80FF5B041C8104986D" box="[256,365,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" pageId="15" pageNumber="466" unit="mm" value="6.15">6.15 mm</quantity>
. Head width (measured between tips of canthi) =
<quantity id="E8462DA3FC5FFF80FD94041C8455986D" box="[975,1084,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.92" pageId="15" pageNumber="466" unit="mm" value="6.92">6.92 mm</quantity>
; ratio both eyes/head = 0.16. Postorbital pits situated immediately behind supraorbital ridges, almost rectangular, shallow and with a few punctures; external borders with setose punctures. Ligula slightly protuberant basally; with a central apical tooth small and with a transversal, well marked anterior ventral ridge; setose punctures present medially. Lateral lobes of mentum with abundant setose punctures; lateral basal scars oval, brilliant and punctate setose, internal border longitudinally striate. Medial basal mentum bare and with sparse micropunctures visible at great magnification. Hypostomal process elongate, wide medially and narrow in apical third, without lateral depression. Infraocular ridge short, external and basal area punctate-pubescent. Dorsal tooth of mandible occupies 1/2 length of the mandible; internal face of mandible in dorsal view, granular. Antennal club (
<figureCitation id="B7859CC3FC5FFF80FDAD057C8415990B" box="[1014,1148,836,861]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="16.[151,250,1926,1948]" captionStartId-1="17.[151,250,1945,1967]" captionTargetBox-0="[159,1436,310,1896]" captionTargetBox-1="[162,1427,206,1916]" captionTargetId-0="figure-89@16.[151,1436,301,1905]" captionTargetId-1="figure-45@17.[151,1436,193,1924]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 11. Paratype of Ogyges menchuae: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 12. Ogyges menchuae: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952433" figureDoi-1="http://doi.org/10.5281/zenodo.4952435" httpUri-0="https://zenodo.org/record/4952433/files/figure.png" httpUri-1="https://zenodo.org/record/4952435/files/figure.png" pageId="15" pageNumber="466">Figs. 11–12</figureCitation>
)
<quantity id="E8462DA3FC5FFF80FAD7057C8498990D" box="[1164,1265,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="15" pageNumber="466" unit="mm" value="1.88">1.88 mm</quantity>
long,
<quantity id="E8462DA3FC5FFF80FB63057C85F4990D" box="[1336,1437,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="15" pageNumber="466" unit="mm" value="1.31">1.31 mm</quantity>
wide; with last two antennomeres wider than the antepenultimate, this slightly more than twice as wide as long.
</paragraph>
<paragraph id="2F018046FC5FFF80FE9C05B4836F9E95" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">Thorax: Lateral fossae of pronotum with micropunctures of rugose aspect, visible with moderate magnification. Pronotum with marginal groove narrow, widened at anterior margin; anterior angles rounded, slightly projected anteriorly immediately behind eyes. Pronotum with dense and shallow micropunctures of rugose aspect, visible at moderate magnification, running from the lateral fossae to the anterior border, internal to marginal groove; disc with sparse micropunctures visible at great magnification. Hypomeron with abundant small shallow punctures. Prosternellum completely shagreened. Mesosternum bare, lateral depressions elongate, rugose (shagreened). Mesepisternum with large and oval rugose (shagreened) area extended to posterior margin. Metasternum with anterior angles bare; disc not delimited by punctures; marginal groove bare and narrow, posterior part 1.5–2.0 times wider than the median part.</paragraph>
<paragraph id="2F018046FC5FFF80FE9C02F782159F79" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">Elytra: Striations marked and with small but defined punctures, wider and deeper from 5–10; interstriae 1–7 with sparse micropunctures, more abundant and visible at moderate magnification on interstriae 8–10; junctions of striations 1 and 10 without extra punctures.</paragraph>
<paragraph id="2F018046FC5FFF80FE9C0304808D9F2E" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">Legs: Profemur with anterioventral groove marked, metafemur widened; mesotibia with two (rarely three) spines.</paragraph>
<paragraph id="2F018046FC5FFF80FE9C03BC842B9FCB" blockId="15.[151,1437,151,2018]" box="[199,1090,1412,1437]" pageId="15" pageNumber="466">Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.</paragraph>
<paragraph id="2F018046FC5FFF80FE9C039F82899FB3" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">
Aedeagus: In ventral view phallus globose; parameres and phallobase separated; in dorsal view, ventrodorsal basal sclerotizations of the phallus present (
<figureCitation id="B7859CC3FC5FFF80FCDB03F482BA9FB3" box="[640,723,1484,1509]" captionStart="FIGURE 12" captionStartId="17.[151,250,1945,1967]" captionTargetBox="[162,1427,206,1916]" captionTargetId="figure-45@17.[151,1436,193,1924]" captionTargetPageId="17" captionText="FIGURE 12. Ogyges menchuae: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4952435" httpUri="https://zenodo.org/record/4952435/files/figure.png" pageId="15" pageNumber="466">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="2F018046FC5FFF80FE9C03D785FA9C23" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">
Variation (
<emphasis id="1DCA5C54FC5FFF80FF1A03C981269C5E" box="[321,335,1521,1544]" italics="true" pageId="15" pageNumber="466">n</emphasis>
= 20): Total length 29.36–33.15 ( = 31.43), elytral length 16.11–18.66 ( = 17.29), pronotal length 7.40–9.13 ( = 8.56), pronotal width 10.12–11.72 ( = 10.93), humeral width 9.46–10.71 ( = 10.17). Mediofrontal area more or less smooth to somewhat or moderately roughened, with scarce to barely visible microgranulations. Some specimens present an anterior keel in the concavity of the angle of posterofrontal ridges.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5FFF80FE9C004785EA9DB7" pageId="15" pageNumber="466" type="materials_examined">
<paragraph id="2F018046FC5FFF80FE9C0047823E9CCE" blockId="15.[151,1437,151,2018]" box="[199,599,1663,1688]" pageId="15" pageNumber="466">
<emphasis id="1DCA5C54FC5FFF80FE9C004781D99CC1" box="[199,432,1663,1688]" inLineHeading="true" pageId="15" pageNumber="466" reason="1">
<emphasis id="1DCA5C54FC5FFF80FE9C004781C09CCE" bold="true" box="[199,425,1663,1688]" pageId="15" pageNumber="466">Material examined</emphasis>
.
</emphasis>
<specimenCount id="39B84BCFFC5FFF80FFEC0047823A9CCE" box="[439,595,1663,1688]" pageId="15" pageNumber="466" type="generic">35 specimens</specimenCount>
.
</paragraph>
<paragraph id="2F018046FC5FFF80FE9C009C83649CB6" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">
<emphasis id="1DCA5C54FC5FFF80FE9C009C811C9CED" box="[199,373,1700,1725]" inLineHeading="true" pageId="15" pageNumber="466" reason="1">
<emphasis id="1DCA5C54FC5FFF80FE9C009C81079CEB" bold="true" box="[199,366,1700,1725]" pageId="15" pageNumber="466">Type material</emphasis>
.
</emphasis>
<materialsCitation id="9FD68A1BFC5FFF80FF27009C83649CB6" collectingDate="1998-07" collectorName="E. B. Cano" country="Guatemala" elevation="2100" location="Laj Chimel" municipality="Uspantan" pageId="15" pageNumber="466" specimenCount="1" stateProvince="Quiche" typeStatus="holotype">
<typeStatus id="F0053EE4FC5FFF80FF27009C81809CEB" box="[380,489,1700,1725]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="57A9C0D6FC5FFF80FFAF009C82CE9CEB" box="[500,679,1700,1725]" name="Guatemala" pageId="15" pageNumber="466">GUATEMALA</collectingCountry>
:
<collectingRegion id="ED7A4EA4FC5FFF80FCEA009C836C9CEB" box="[689,773,1700,1725]" country="Guatemala" name="Quiche" pageId="15" pageNumber="466">Quiché</collectingRegion>
,
<collectingMunicipality id="CF651A3CFC5FFF80FD4B009C83149CEB" box="[784,893,1700,1725]" pageId="15" pageNumber="466">Uspantán</collectingMunicipality>
, aldea
<location id="2A61D69DFC5FFF80FD90009C84249CEB" LSID="urn:lsid:plazi:treatment:A7173150FC5EFF9FFECC006482BC9AAE:2A61D69DFC5FFF80FD90009C84249CEB" box="[971,1101,1700,1725]" country="Guatemala" municipality="Uspantan" name="Laj Chimel" pageId="15" pageNumber="466" stateProvince="Quiche">Laj Chimel</location>
, montaña al norte de la aldea,
<quantity id="E8462DA3FC5FFF80FECC00FF80989C89" box="[151,241,1735,1760]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="466" unit="m" value="2100.0">
<elevation id="A4936775FC5FFF80FECC00FF80989C89" box="[151,241,1735,1760]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="466" unit="m" value="2100.0">2100 m</elevation>
</quantity>
.,
<date id="5B00A686FC5FFF80FF5A00F081009CB6" box="[257,361,1735,1760]" pageId="15" pageNumber="466" value="1998-07">
<collectingDate id="4B445F6EFC5FFF80FF5A00F081009CB6" box="[257,361,1735,1760]" pageId="15" pageNumber="466" value="1998-07">VII 1998</collectingDate>
</date>
, bosque nuboso.
<collectorName id="824BE590FC5FFF80FC7100F082F49CB6" box="[554,669,1735,1760]" pageId="15" pageNumber="466">E.B. Cano</collectorName>
collector.
</materialsCitation>
</paragraph>
<paragraph id="2F018046FC5FFF80FE9C00D4810C9DE8" blockId="15.[151,1437,151,2018]" pageId="15" pageNumber="466">
<materialsCitation id="9FD68A1BFC5FFF80FE9C00D4839F9D53" box="[199,1014,1772,1797]" collectingDate="1998-07" collectorName="E. B. Cano" country="Guatemala" elevation="2100" location="Laj Chimel" municipality="Uspantan" pageId="15" pageNumber="466" specimenCount="1" specimenCount-male="1" stateProvince="Quiche" typeStatus="paratype">
<typeStatus id="F0053EE4FC5FFF80FE9C00D481509D52" box="[199,313,1772,1796]" pageId="15" pageNumber="466" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="57A9C0D6FC5FFF80FF1F00D4819E9D53" box="[324,503,1772,1797]" name="Guatemala" pageId="15" pageNumber="466">GUATEMALA</collectingCountry>
: same data as holotype (
<specimenCount id="39B84BCFFC5FFF80FD4B00D483379D53" box="[784,862,1772,1797]" pageId="15" pageNumber="466" type="male">1 male</specimenCount>
, 7 unknown)
</materialsCitation>
;
<materialsCitation id="9FD68A1BFC5FFF80FA5900D4823E9D7E" collectingDate="1998-07" collectorName="E. B. Cano" country="Guatemala" elevation="2100" location="Norte Laj Chimel" municipality="San Pedro" pageId="15" pageNumber="466" specimenCount="1" specimenCount-male="1" stateProvince="Quiche" typeStatus="paratype">
same data except
<location id="2A61D69DFC5FFF80FA9600D485F19D53" LSID="urn:lsid:plazi:treatment:A7173150FC5EFF9FFECC006482BC9AAE:2A61D69DFC5FFF80FA9600D485F19D53" box="[1229,1432,1772,1797]" country="Guatemala" municipality="San Pedro" name="Norte Laj Chimel" pageId="15" pageNumber="466">Norte Laj Chimel</location>
,
<collectingMunicipality id="CF651A3CFC5FFF80FECC013781629D7E" box="[151,267,1807,1832]" pageId="15" pageNumber="466">San Pedro</collectingMunicipality>
,
<quantity id="E8462DA3FC5FFF80FF4D0137810D9D7E" box="[278,356,1807,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="466" unit="m" value="2100.0">
<elevation id="A4936775FC5FFF80FF4D0137810D9D7E" box="[278,356,1807,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="466" unit="m" value="2100.0">2100m</elevation>
</quantity>
(
<specimenCount id="39B84BCFFC5FFF80FF2D013781AB9D7E" box="[374,450,1807,1832]" pageId="15" pageNumber="466" type="male">1 male</specimenCount>
, 1 unknown)
</materialsCitation>
;
<materialsCitation id="9FD68A1BFC5FFF80FC3A012885FE9D7E" box="[609,1431,1807,1832]" collectingDate="2011-05-30" collectionCode="V" collectorName="Aldea Laj Chimel &amp; Gonzalez-Madrid" country="Guatemala" elevation="2100" location="Norte Laj Chimel" municipality="San Pedro" pageId="15" pageNumber="466" specimenCount="1" stateProvince="Quiche" typeStatus="paratype">
same data except
<collectorName id="824BE590FC5FFF80FD7C0137839B9D7E" box="[807,1010,1807,1832]" pageId="15" pageNumber="466">Aldea Laj Chimel</collectorName>
, A.
<collectorName id="824BE590FC5FFF80FA4F013784899D7E" box="[1044,1248,1807,1832]" pageId="15" pageNumber="466">González-Madrid</collectorName>
,
<date id="5B00A686FC5FFF80FAB0013785369D7E" box="[1259,1375,1807,1832]" pageId="15" pageNumber="466" value="2011-05-30">
<collectingDate id="4B445F6EFC5FFF80FAB0013785369D7E" box="[1259,1375,1807,1832]" pageId="15" pageNumber="466" value="2011-05-30">
30
<collectionCode id="49AF1883FC5FFF80FB550128854B9D71" box="[1294,1314,1808,1831]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="466" type="Museum">V</collectionCode>
2011
</collectingDate>
</date>
(13)
</materialsCitation>
;
<materialsCitation id="9FD68A1BFC5FFF80FECC010F82B89D25" collectingDate="2011-04-30" collectingDateMax="2011-05-30" collectingDateMin="2011-04-30" collectionCode="V" collectorName="Aldea Laj Chimel &amp; San Pablo &amp; Gonzalez-Madrid" country="Guatemala" elevation="2035" latitude="15.458525" location="Norte Laj Chimel" longLatPrecision="1" longitude="-90.773926" municipality="San Pedro" pageId="15" pageNumber="466" specimenCount="3" specimenCount-male="3" stateProvince="Quiche" typeStatus="paratype">
same data except,
<collectorName id="824BE590FC5FFF80FF31010E82539D19" box="[362,570,1846,1871]" pageId="15" pageNumber="466">Aldea Laj Chimel</collectorName>
, road to
<collectorName id="824BE590FC5FFF80FCC4010E837C9D19" box="[671,789,1846,1871]" pageId="15" pageNumber="466">San Pablo</collectorName>
,
<date id="5B00A686FC5FFF80FD79010E83CE9D19" box="[802,935,1846,1871]" pageId="15" pageNumber="466" value="2011-04-30">
<collectingDate id="4B445F6EFC5FFF80FD79010E83CE9D19" box="[802,935,1846,1871]" pageId="15" pageNumber="466" value="2011-04-30">30 IV 2011</collectingDate>
</date>
,
<geoCoordinate id="4A8AE681FC5FFF80FDEF010E84309D19" box="[948,1113,1843,1871]" degrees="15" direction="north" minutes="27" orientation="latitude" pageId="15" pageNumber="466" precision="1" seconds="30.69" value="15.458525">
15
<superScript id="D8CB2D0EFC5FFF80FD8B010B83BE9D17" attach="none" box="[976,983,1843,1857]" fontSize="6" pageId="15" pageNumber="466">o</superScript>
27'30.69''N
</geoCoordinate>
,
<geoCoordinate id="4A8AE681FC5FFF80FA3D010E85649D19" box="[1126,1293,1843,1871]" degrees="90" direction="west" minutes="46" orientation="longitude" pageId="15" pageNumber="466" precision="1" seconds="26.11" value="-90.773926">
90
<superScript id="D8CB2D0EFC5FFF80FAD9010B84E09D17" attach="none" box="[1154,1161,1843,1857]" fontSize="6" pageId="15" pageNumber="466">o</superScript>
46'26.11''W
</geoCoordinate>
,
<quantity id="E8462DA3FC5FFF80FB41010E851A9D18" box="[1306,1395,1846,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.035" pageId="15" pageNumber="466" unit="m" value="2035.0">
<elevation id="A4936775FC5FFF80FB41010E851A9D18" box="[1306,1395,1846,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.035" pageId="15" pageNumber="466" unit="m" value="2035.0">2035 m</elevation>
</quantity>
, A.
<collectorName id="824BE590FC5FFF80FECC016281359D25" box="[151,348,1882,1907]" pageId="15" pageNumber="466">Gónzález-Madrid</collectorName>
<date id="5B00A686FC5FFF80FF3F016281B39D25" box="[356,474,1882,1907]" pageId="15" pageNumber="466" value="2011-05-30">
<collectingDate id="4B445F6EFC5FFF80FF3F016281B39D25" box="[356,474,1882,1907]" pageId="15" pageNumber="466" value="2011-05-30">
30
<collectionCode id="49AF1883FC5FFF80FFDC016381F29D24" box="[391,411,1883,1906]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="466" type="Museum">V</collectionCode>
2011
</collectingDate>
</date>
(
<specimenCount id="39B84BCFFC5FFF80FFB0016282539D25" box="[491,570,1882,1907]" pageId="15" pageNumber="466" type="male">3 male</specimenCount>
, 1 unknown)
</materialsCitation>
;
<materialsCitation id="9FD68A1BFC5FFF80FC86016382A59DCC" collectingDate="2011-04-30" collectingDateMax="2011-05-30" collectingDateMin="2011-04-30" collectionCode="V" collectorName="Aldea Laj Chimel &amp; San Pablo" country="Guatemala" elevation="2035" latitude="15.458525" location="Norte Laj Chimel" longLatPrecision="1" longitude="-90.773926" municipality="San Pedro" pageId="15" pageNumber="466" specimenCount="1" stateProvince="Quiche" typeStatus="paratype">
same data except,
<collectorName id="824BE590FC5FFF80FDF7016284119D25" box="[940,1144,1882,1907]" pageId="15" pageNumber="466">Aldea Laj Chimel</collectorName>
, road to
<collectorName id="824BE590FC5FFF80FA80016285399D25" box="[1243,1360,1882,1907]" pageId="15" pageNumber="466">San Pablo</collectorName>
,
<date id="5B00A686FC5FFF80FB00016280B89DCC" pageId="15" pageNumber="466" value="2011-04-30">
<collectingDate id="4B445F6EFC5FFF80FB00016280B89DCC" pageId="15" pageNumber="466" value="2011-04-30">30 IV 2011</collectingDate>
</date>
,
<geoCoordinate id="4A8AE681FC5FFF80FE8501B981ED9DCC" box="[222,388,1918,1946]" degrees="15" direction="north" minutes="27" orientation="latitude" pageId="15" pageNumber="466" precision="1" seconds="30.69" value="15.458525">
15
<superScript id="D8CB2D0EFC5FFF80FEA00146816B9DDA" attach="right" box="[251,258,1918,1932]" fontSize="6" pageId="15" pageNumber="466">o</superScript>
27'30.69''N
</geoCoordinate>
,
<geoCoordinate id="4A8AE681FC5FFF80FFCA01B982509DCC" box="[401,569,1918,1946]" degrees="90" direction="west" minutes="46" orientation="longitude" pageId="15" pageNumber="466" precision="1" seconds="26.11" value="-90.773926">
90
<superScript id="D8CB2D0EFC5FFF80FFF6014681DD9DDA" attach="none" box="[429,436,1918,1932]" fontSize="6" pageId="15" pageNumber="466">o</superScript>
46'26.11''W
</geoCoordinate>
,
<quantity id="E8462DA3FC5FFF80FC1C01B982F79DCF" box="[583,670,1921,1946]" metricMagnitude="3" metricUnit="m" metricValue="2.035" pageId="15" pageNumber="466" unit="m" value="2035.0">
<elevation id="A4936775FC5FFF80FC1C01B982F79DCF" box="[583,670,1921,1946]" metricMagnitude="3" metricUnit="m" metricValue="2.035" pageId="15" pageNumber="466" unit="m" value="2035.0">2035 m</elevation>
</quantity>
(6)
</materialsCitation>
;
<materialsCitation id="9FD68A1BFC5FFF80FC8101BA81089DE8" collectingDate="2012-06-15" collectingDateMax="2011-05-30" collectingDateMin="2011-04-30" collectionCode="VI" collectorName="Montana El Amay &amp; A. Zamora" country="Guatemala" elevation="2035" latitude="15.458525" location="Norte Laj Chimel" longLatPrecision="1" longitude="-90.773926" municipality="San Pedro" pageId="15" pageNumber="466" specimenCount="2" specimenCount-male="2" stateProvince="Quiche" typeStatus="paratype">
same data except,
<collectorName id="824BE590FC5FFF80FDEB01BA84EF9DCC" box="[944,1158,1921,1946]" pageId="15" pageNumber="466">Montaña El Amay</collectorName>
, in logs
<date id="5B00A686FC5FFF80FAB001B9851A9DCC" box="[1259,1395,1921,1946]" pageId="15" pageNumber="466" value="2012-06-15">
<collectingDate id="4B445F6EFC5FFF80FAB001B9851A9DCC" box="[1259,1395,1921,1946]" pageId="15" pageNumber="466" value="2012-06-15">
15
<collectionCode id="49AF1883FC5FFF80FB4A01BA85479DCF" box="[1297,1326,1922,1945]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="466">VI</collectionCode>
2012
</collectingDate>
</date>
,
<collectorName id="824BE590FC5FFF80FBDA01B980879DEB" pageId="15" pageNumber="466">A. Zamora</collectorName>
(
<specimenCount id="39B84BCFFC5FFF80FEA6019D81309DE8" box="[253,345,1957,1982]" pageId="15" pageNumber="466" type="male">2 males</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2F018046FC5FFF80FE9C01F185EA9DB7" blockId="15.[151,1437,151,2018]" box="[199,1411,1993,2018]" pageId="15" pageNumber="466">
<materialsCitation id="9FD68A1BFC5FFF80FE9C01F182719DB4" box="[199,536,1993,2018]" collectionCode="UVGC" pageId="15" pageNumber="466" specimenCount="1" typeStatus="holotype">
<typeStatus id="F0053EE4FC5FFF80FE9C01F181469DB4" box="[199,303,1993,2018]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="49AF1883FC5FFF80FF9E01F182719DB4" box="[453,536,1993,2018]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="15" pageNumber="466">UVGC</collectionCode>
</materialsCitation>
.
<materialsCitation id="9FD68A1BFC5FFF80FC7901F284859DB4" box="[546,1260,1993,2018]" collectionCode="UVGC, IEXA, USAC, IBUNAM, SNM" pageId="15" pageNumber="466" specimenCount="1" typeStatus="paratype">
<typeStatus id="F0053EE4FC5FFF80FC7901F282E69DB4" box="[546,655,1994,2018]" pageId="15" pageNumber="466" type="paratype">Paratypes</typeStatus>
deposited in
<collectionCode id="49AF1883FC5FFF80FD7D01F183119DB4" box="[806,888,1993,2018]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="15" pageNumber="466">UVGC</collectionCode>
,
<collectionCode id="49AF1883FC5FFF80FDD801F183A09DB4" box="[899,969,1993,2018]" pageId="15" pageNumber="466">IEXA</collectionCode>
,
<collectionCode id="49AF1883FC5FFF80FD8F01F184489DB4" box="[980,1057,1993,2018]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="15" pageNumber="466">USAC</collectionCode>
,
<collectionCode id="49AF1883FC5FFF80FA7701F184C89DB4" box="[1068,1185,1993,2018]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="15" pageNumber="466" type="University or college">IBUNAM</collectionCode>
,
<collectionCode id="49AF1883FC5FFF80FAF701F184859DB4" box="[1196,1260,1993,2018]" country="Slovakia" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="15" pageNumber="466">SNM</collectionCode>
</materialsCitation>
, and
<collectionCode id="49AF1883FC5FFF80FB7D01F285169DB7" box="[1318,1407,1994,2017]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="466">MNHN</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC40FF9FFE9C06AF82BC9AAE" pageId="16" pageNumber="467" type="etymology">
<paragraph id="2F018046FC40FF9FFE9C06AF85F59A83" blockId="16.[151,1436,151,248]" pageId="16" pageNumber="467">
<emphasis id="1DCA5C54FC40FF9FFE9C06AF81279AE6" bold="true" box="[199,334,151,176]" inLineHeading="true" pageId="16" pageNumber="467" reason="1">Etymology.</emphasis>
Named in honor of the Nobel Prize winner, Dr. Rigoberta Menchú, who was born in Laj Chimel.
<emphasis id="1DCA5C54FC40FF9FFE9C0684819E9A83" bold="true" box="[199,503,188,213]" inLineHeading="true" pageId="16" pageNumber="467" reason="1">Distribution and ecology.</emphasis>
Known only from middle altitude cloud forest at Cerro El Amay, of Laj Chimel,
</paragraph>
<paragraph id="2F018046FC40FF9FFECC06E782BC9AAE" blockId="16.[151,1436,151,248]" box="[151,725,223,248]" pageId="16" pageNumber="467">
<materialsCitation id="9FD68A1BFC40FF9FFECC06E782B89AAE" box="[151,721,223,248]" country="Guatemala" elevation="1950" location="Quiche" pageId="16" pageNumber="467" specimenCount="1" stateProvince="Quiche">
<collectingRegion id="ED7A4EA4FC40FF9FFECC06E780829AAE" box="[151,235,223,248]" country="Guatemala" name="Quiche" pageId="16" pageNumber="467">Quiché</collectingRegion>
,
<collectingCountry id="57A9C0D6FC40FF9FFEAD06E7811B9AAE" box="[246,370,223,248]" name="Guatemala" pageId="16" pageNumber="467">Guatemala</collectingCountry>
,
<quantity id="E8462DA3FC40FF9FFF2606E782719AA1" box="[381,536,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="16" pageNumber="467" unit="m" value="1950.0" valueMax="2100.0" valueMin="1800.0">
<elevation id="A4936775FC40FF9FFF2606E782719AA1" box="[381,536,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="16" pageNumber="467" unit="m" value="1950.0" valueMax="2100.0" valueMin="1800.0">1800–2100 m</elevation>
</quantity>
altitude (
<figureCitation id="B7859CC3FC40FF9FFCD906E782A19AAE" box="[642,712,223,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1904,1926]" captionTargetBox="[171,1398,927,1870]" captionTargetId="figure-358@6.[149,1434,899,1893]" captionTargetPageId="6" captionText="FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi." figureDoi="http://doi.org/10.5281/zenodo.4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="16" pageNumber="467">Fig. 4</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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