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<document id="10155E336342B1683BF757E026EF0180" ID-DOI="10.11646/zootaxa.3889.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4952403" ID-ZooBank="1F3F076D-A4F1-4B4A-977D-4B99581C6779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730508465" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2014" docId="A7173150FC5BFF81FECC0114809B9C5E" docLanguage="en" docName="zt03889p484.pdf" docOrigin="Zootaxa 3889 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ogyges llama Cano 2014, new species" docType="treatment" docVersion="4" lastPageNumber="465" masterDocId="5B2E4928FC50FF8FFE5B063880699A56" masterDocTitle="Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly" masterLastPageNumber="484" masterPageNumber="451" pageNumber="462" updateTime="1698967030688" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7E5A873A708A19C5923086158D9F5CF3">Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly</mods:title>
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<mods:namePart id="C6B0BE9777FA6337FD24B539F34C2083">Cano, Enio B.</mods:namePart>
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<mods:date id="84D0D6633F3EE956FC3DBD81D6E4EDB0">2014</mods:date>
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<mods:number id="6C0F1095D7D12CB47CB53993C4349857">3889</mods:number>
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<mods:classification id="567EA1CC68823D81652DD2861E5090D4">journal article</mods:classification>
<mods:identifier id="ECC4ED50F6D20E6E62A2188611972B7B" type="DOI">10.11646/zootaxa.3889.4.1</mods:identifier>
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<treatment id="A7173150FC5BFF81FECC0114809B9C5E" ID-DOI="http://doi.org/10.5281/zenodo.4953257" ID-Zenodo-Dep="4953257" LSID="urn:lsid:plazi:treatment:A7173150FC5BFF81FECC0114809B9C5E" httpUri="http://treatment.plazi.org/id/A7173150FC5BFF81FECC0114809B9C5E" lastPageId="14" lastPageNumber="465" pageId="11" pageNumber="462">
<subSubSection id="67A4D3CDFC5BFF84FECC0114824A9D10" box="[151,547,1836,1862]" pageId="11" pageNumber="462" type="nomenclature">
<paragraph id="2F018046FC5BFF84FECC0114824A9D10" blockId="11.[151,547,1836,1895]" box="[151,547,1836,1862]" pageId="11" pageNumber="462">
<heading id="7449372AFC5BFF84FECC0114824A9D10" bold="true" box="[151,547,1836,1862]" fontSize="11" level="1" pageId="11" pageNumber="462" reason="1">
<emphasis id="1DCA5C54FC5BFF84FECC0114824A9D10" bold="true" box="[151,547,1836,1862]" pageId="11" pageNumber="462">
<taxonomicName id="E8BEFBC5FC5BFF84FECC011481EE9D10" authority="Cano" authorityName="Cano" authorityYear="2014" box="[151,391,1836,1862]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="llama" status="sp. nov.">
<emphasis id="1DCA5C54FC5BFF84FECC011481529D10" bold="true" box="[151,315,1836,1862]" italics="true" pageId="11" pageNumber="462">Ogyges llama</emphasis>
Cano
</taxonomicName>
,
<taxonomicNameLabel id="06F9E12FFC5BFF84FFC80114824A9D10" box="[403,547,1836,1862]" pageId="11" pageNumber="462" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5BFF84FECC0176812C9D31" box="[151,325,1870,1895]" pageId="11" pageNumber="462" type="description">
<paragraph id="2F018046FC5BFF84FECC0176812C9D31" blockId="11.[151,547,1836,1895]" box="[151,325,1870,1895]" pageId="11" pageNumber="462">
<figureCitation id="B7859CC3FC5BFF84FECC017681669D31" box="[151,271,1870,1895]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="12.[151,250,1946,1968]" captionStartId-1="13.[151,250,1733,1755]" captionTargetBox-0="[170,1432,582,1914]" captionTargetBox-1="[164,1421,207,1695]" captionTargetId-0="figure-210@12.[151,1436,556,1925]" captionTargetId-1="figure-135@13.[151,1436,193,1712]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 9. Paratype of Ogyges llama: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 10. Ogyges llama: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952425" figureDoi-1="http://doi.org/10.5281/zenodo.4952427" httpUri-0="https://zenodo.org/record/4952425/files/figure.png" httpUri-1="https://zenodo.org/record/4952427/files/figure.png" pageId="11" pageNumber="462">
Figs. 9–
<geoCoordinate id="4A8AE681FC5BFF84FEA8017681669D31" box="[243,271,1870,1895]" degrees="10, 21" direction="north" orientation="latitude" pageId="11" pageNumber="462" precision="555" value="10.21">10</geoCoordinate>
</figureCitation>
,
<figureCitation id="B7859CC3FC5BFF84FF400176812C9D31" box="[283,325,1870,1895]" captionStart="FIGURE 21" captionStartId="28.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure-318@28.[151,1436,193,1354]" captionTargetPageId="28" captionText="FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili." figureDoi="http://doi.org/10.5281/zenodo.4952455" httpUri="https://zenodo.org/record/4952455/files/figure.png" pageId="11" pageNumber="462">21n</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5BFF83FECC01AE83879B4B" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="2F018046FC5BFF83FECC01AE83879B4B" blockId="11.[151,1436,1942,2004]" lastBlockId="12.[151,1437,151,503]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<emphasis id="1DCA5C54FC5BFF84FECC01AE81789DF9" bold="true" box="[151,273,1942,1967]" inLineHeading="true" pageId="11" pageNumber="462" reason="1">Diagnosis.</emphasis>
By the elytra with wide, deep dorsal punctures,
<taxonomicName id="E8BEFBC5FC5BFF84FD1F01AF83C09DF9" authorityName="Cano" authorityYear="2014" box="[836,937,1943,1967]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="llama">
<emphasis id="1DCA5C54FC5BFF84FD1F01AF83C09DF9" box="[836,937,1943,1967]" italics="true" pageId="11" pageNumber="462">O. llama</emphasis>
</taxonomicName>
is similar to the “
<taxonomicName id="E8BEFBC5FC5BFF84FA2601AF85629DF9" baseAuthorityName="Casey" baseAuthorityYear="1897" box="[1149,1291,1943,1967]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="crassulus">
<emphasis id="1DCA5C54FC5BFF84FA2601AF85629DF9" box="[1149,1291,1943,1967]" italics="true" pageId="11" pageNumber="462">O. crassulus</emphasis>
</taxonomicName>
” lineage (as suggested by
<bibRefCitation id="4B2FFDB7FC5BFF84FF680183825F9D82" author="Schuster, J. C. &amp; Cano, E. B. &amp; Boucher, S." box="[307,566,1979,2004]" pageId="11" pageNumber="462" pagination="115 - 132" refId="ref19637" refString="Schuster, J. C., Cano, E. B. &amp; Boucher, S. (2005) Ogyges and Veturius (Coleoptera: Passalidae) in Central America: synonymies, range extensions and new species. Acta Zoologica Mexicana (n. s.), 21, 115 - 132." type="journal article" year="2005">
Schuster
<emphasis id="1DCA5C54FC5BFF84FFC7018481A79D85" box="[412,462,1979,2003]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. (2005))
</bibRefCitation>
. It is easily separated from
<taxonomicName id="E8BEFBC5FC5BFF84FD290183839C9D85" authorityName="Schuster, Cano, &amp; Boucher" authorityYear="2005" box="[882,1013,1979,2003]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="monzoni">
<emphasis id="1DCA5C54FC5BFF84FD290183839C9D85" box="[882,1013,1979,2003]" italics="true" pageId="11" pageNumber="462">O. monzoni</emphasis>
</taxonomicName>
and
<taxonomicName id="E8BEFBC5FC5BFF84FA74018384E29D85" authorityName="Schuster, Cano, &amp; Boucher" authorityYear="2005" box="[1071,1163,1979,2004]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="aluxi">
O.
<emphasis id="1DCA5C54FC5BFF84FA08018384E29D85" box="[1107,1163,1979,2003]" italics="true" pageId="11" pageNumber="462">aluxi</emphasis>
</taxonomicName>
by the smaller size, the central horn not free or scarcely free and by the medial central mentum weakly rugopunctate. It can be separated from
<taxonomicName id="E8BEFBC5FC5CFF83FE8F068481299A82" authorityName="Cano" authorityYear="2014" box="[212,320,188,212]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="laurae">
<emphasis id="1DCA5C54FC5CFF83FE8F068481299A82" box="[212,320,188,212]" italics="true" pageId="12" pageNumber="463">O. laurae</emphasis>
</taxonomicName>
by the mesotibia with two spines (instead of one) and the tumose posterior portion of central horn. From the sympatric or almost sympatric
<taxonomicName id="E8BEFBC5FC5CFF83FC3606D882959AAE" baseAuthorityName="Casey" baseAuthorityYear="1897" box="[621,764,224,248]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="crassulus">
<emphasis id="1DCA5C54FC5CFF83FC3606D882959AAE" box="[621,764,224,248]" italics="true" pageId="12" pageNumber="463">O. crassulus</emphasis>
</taxonomicName>
it is separated by the form of aedeagus, with lateral lobes separated from basal piece, and the widest punctures on posterior striae 6–9.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5CFF81FE9C071F85679E2B" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463" type="description">
<paragraph id="2F018046FC5CFF81FE9C071F8429986B" blockId="12.[151,1437,151,503]" lastBlockId="14.[151,1437,151,1544]" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463">
<emphasis id="1DCA5C54FC5CFF83FE9C071F81309B16" bold="true" box="[199,345,295,320]" inLineHeading="true" pageId="12" pageNumber="463" reason="1">Description.</emphasis>
<typeStatus id="F0053EE4FC5CFF83FF05071F81AF9B16" box="[350,454,295,320]" pageId="12" pageNumber="463" type="holotype">Holotype</typeStatus>
male, black (dorsum) to dark brown (venter) teneral, with blue iridescence. Total length 22.61, elytral length 11.85, pronotal length 6.23, pronotal width 8.16, humeral width 7.56. Head: anterior border of labrum slightly concave. Clypeus inclined (&gt; 90
<superScript id="D8CB2D0EFC5CFF83FCE7075782AA9B2B" attach="none" box="[700,707,367,381]" fontSize="6" pageId="12" pageNumber="463">o</superScript>
); anterior border straight, lacking a central invagination, with a scabrose (to almost granular) sulcus separating it from the smooth (almost shagreened at great magnification) mediofrontal area. Mediofrontal tubercles rounded and defined; internal tubercles and posterofrontal ridges absent. Laterofrontal areas smooth. Lateropostfrontal areas wide, glabrous, and smooth; a marked or weakly marked keel runs laterally from the mediopostfrontal structure, forming the anterior border of lateropostfrontal areas. A wide, bare, smooth fossa present in front of the mediopostfrontal structure. Central horn short, narrow, with apex weakly free (or almost not free), directed upward, without median longitudinal groove posteriorly; lateroposterior tubercles not elongate, forming an angle directed forward and sideways, separated from the central horn by a groove. Postfrontal groove shallow, medially very narrow (constrained by the extended mediopostfrontal structure), gradually enlarged at sides and almost continuous with the lateropostfrontal areas. Supraorbital ridge with equal anterior tubercles; posterior 1/2 bifurcate with the supraorbital fossae deep; external ridge well marked. Ocular canthus with apex straight or slightly swollen, ventrally covering at least the half of eye. Eyes reduced. Eye width =
<quantity id="E8462DA3FC5EFF81FECC06E780969AA1" box="[151,255,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="465" unit="mm" value="0.42">0.42 mm</quantity>
. Interocular width =
<quantity id="E8462DA3FC5EFF81FFAB06E782319AA1" box="[496,600,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.61" pageId="14" pageNumber="465" unit="mm" value="4.61">4.61 mm</quantity>
. Head width (measured between tips of ocular canthus) =
<quantity id="E8462DA3FC5EFF81FAA806E785359AA1" box="[1267,1372,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" pageId="14" pageNumber="465" unit="mm" value="5.85">5.85 mm</quantity>
; ratio both eye widths/head width = 0.14. Postorbital pits oval, elongate and covered with sparse setae on external border. Ligula slightly protuberant basally, with small central tooth and without a ventral anterior transversal carina (a tumosity is present); medially with setose punctures. Lateral lobes of mentum with moderate setose punctures, lateral basal scars oval and opaque (shagreened); medial basal mentum shiny, with a few scattered small and scabrous setose punctures, setae long and slender. Hypostomal process elongate, without lateral depression, wide medially and narrow in apical third. Infraocular ridge present, short and smooth, surrounded by striate, setose punctures. Mandible with dorsal tooth occupying 1/2 of his length; internal face of mandible in dorsal view smooth, not granular. Antepenultimate antennomere of antennal club subequal in length but less wide than the penultimate (
<figureCitation id="B7859CC3FC5EFF81FF71041C81CF986B" box="[298,422,548,573]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="12.[151,250,1946,1968]" captionStartId-1="13.[151,250,1733,1755]" captionTargetBox-0="[170,1432,582,1914]" captionTargetBox-1="[164,1421,207,1695]" captionTargetId-0="figure-210@12.[151,1436,556,1925]" captionTargetId-1="figure-135@13.[151,1436,193,1712]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 9. Paratype of Ogyges llama: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 10. Ogyges llama: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952425" figureDoi-1="http://doi.org/10.5281/zenodo.4952427" httpUri-0="https://zenodo.org/record/4952425/files/figure.png" httpUri-1="https://zenodo.org/record/4952427/files/figure.png" pageId="14" pageNumber="465">Figs. 9–10</figureCitation>
); antennal club measures
<quantity id="E8462DA3FC5EFF81FC9D041C8340986D" box="[710,809,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="14" pageNumber="465" unit="mm" value="1.11">1.11 mm</quantity>
wide and
<quantity id="E8462DA3FC5EFF81FDC7041C8469986D" box="[924,1024,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="14" pageNumber="465" unit="mm" value="1.23">1.23 mm</quantity>
long.
</paragraph>
<caption id="7BC1D0CEFC5CFF83FECC01A284809DE6" ID-DOI="http://doi.org/10.5281/zenodo.4952425" ID-Zenodo-Dep="4952425" box="[151,1257,1946,1968]" httpUri="https://zenodo.org/record/4952425/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,1946,1968]" targetBox="[170,1432,582,1914]" targetPageId="12">
<paragraph id="2F018046FC5CFF83FECC01A284809DE6" blockId="12.[151,1257,1946,1968]" box="[151,1257,1946,1968]" pageId="12" pageNumber="463">
<emphasis id="1DCA5C54FC5CFF83FECC01A2817B9DF9" bold="true" box="[151,274,1946,1968]" pageId="12" pageNumber="463">FIGURE 9.</emphasis>
Paratype of
<taxonomicName id="E8BEFBC5FC5CFF83FFC801A282769DE6" authorityName="Cano" authorityYear="2014" box="[403,543,1946,1968]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="llama">
<emphasis id="1DCA5C54FC5CFF83FFC801A282769DE6" box="[403,543,1946,1968]" italics="true" pageId="12" pageNumber="463">Ogyges llama</emphasis>
</taxonomicName>
: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view.
</paragraph>
</caption>
<caption id="7BC1D0CEFC5DFF82FECC00FD80929CAC" ID-DOI="http://doi.org/10.5281/zenodo.4952427" ID-Zenodo-Dep="4952427" httpUri="https://zenodo.org/record/4952427/files/figure.png" pageId="13" pageNumber="464" startId="13.[151,250,1733,1755]" targetBox="[164,1421,207,1695]" targetPageId="13">
<paragraph id="2F018046FC5DFF82FECC00FD80929CAC" blockId="13.[151,1435,1733,1786]" pageId="13" pageNumber="464">
<emphasis id="1DCA5C54FC5DFF82FECC00FD81769C8D" bold="true" box="[151,287,1733,1755]" pageId="13" pageNumber="464">FIGURE 10.</emphasis>
<taxonomicName id="E8BEFBC5FC5DFF82FF7E00FD81D89C8D" authorityName="Cano" authorityYear="2014" box="[293,433,1733,1755]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="llama">
<emphasis id="1DCA5C54FC5DFF82FF7E00FD81D89C8D" box="[293,433,1733,1755]" italics="true" pageId="13" pageNumber="464">Ogyges llama</emphasis>
</taxonomicName>
: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus.
</paragraph>
</caption>
<paragraph id="2F018046FC5EFF81FE9C047F84959943" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">Thorax: Lateral fossae and marginal groove of pronotum with abundant striate micropunctures, visible at great magnification; anterior angles rounded; disc smooth except for scattered shallow micropunctures visible at high magnification. Prosternellum opaque, completely shagreened. Mesosternum glabrous; lateral depressions elongate and rugose (shagreened). Mesepisternum with oval, very elongate shagreened area, reaching the posterior border. Metasternum anterior angles glabrous; disc delimited by 12–16 well marked punctures (some partially fused) on each side; marginal groove glabrous, narrow and smooth, posteriorly two times wider than medially.</paragraph>
<paragraph id="2F018046FC5EFF81FE9C052782F39929" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">Elytra: Opaque, with blue iridescence; striations well marked and with defined punctures, slightly deeper and wider between striae 5–10; anterior 1/3 of sixth interstriae of the width of a puncture of striae 6 or 7; junctions of striations 1 and 10 with some extra punctures.</paragraph>
<paragraph id="2F018046FC5EFF81FE9C05B4808D999E" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">Legs: Profemur with anterioventral groove marked; metafemur moderately widened; mesotibia with two spines.</paragraph>
<paragraph id="2F018046FC5EFF81FE9C05EC842B99BB" blockId="14.[151,1437,151,1544]" box="[199,1090,980,1005]" pageId="14" pageNumber="465">Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.</paragraph>
<paragraph id="2F018046FC5EFF81FE9C05CF84DD9E46" blockId="14.[151,1437,151,1544]" box="[199,1204,1015,1040]" pageId="14" pageNumber="465">
Aedeagus: In ventral view phallus globose; parameres and phallobase separated (
<figureCitation id="B7859CC3FC5EFF81FA0F05CF84CE9E46" box="[1108,1191,1015,1040]" captionStart="FIGURE 10" captionStartId="13.[151,250,1733,1755]" captionTargetBox="[164,1421,207,1695]" captionTargetId="figure-135@13.[151,1436,193,1712]" captionTargetPageId="13" captionText="FIGURE 10. Ogyges llama: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4952427" httpUri="https://zenodo.org/record/4952427/files/figure.png" pageId="14" pageNumber="465">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="2F018046FC5EFF81FE9C022485679E2B" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">
Variation (
<emphasis id="1DCA5C54FC5EFF81FF66022581229E62" box="[317,331,1053,1076]" italics="true" pageId="14" pageNumber="465">n</emphasis>
= 30): Total length 19.30–24.49 ( = 22.95), elytral length 11.7–13.24 ( = 12.59), pronotal length 5.60–7.18 ( = 6.38); pronotal width 7.55–8.69 ( = 8.19); humeral width 6.8–8.26 ( = 7.71). Disc of metasternum delimited by 12–24 well marked punctures (some striate or partially fused) on each side.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF81FE9C02BF853B9F03" pageId="14" pageNumber="465" type="materials_examined">
<paragraph id="2F018046FC5EFF81FE9C02BF823E9EF6" blockId="14.[151,1437,151,1544]" box="[199,599,1159,1184]" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FE9C02BF81D99EC9" box="[199,432,1159,1184]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">
<emphasis id="1DCA5C54FC5EFF81FE9C02BF81C09EF6" bold="true" box="[199,425,1159,1184]" pageId="14" pageNumber="465">Material examined</emphasis>
.
</emphasis>
<specimenCount id="39B84BCFFC5EFF81FFEC02BF823A9EF6" box="[439,595,1159,1184]" pageId="14" pageNumber="465" type="generic">44 specimens</specimenCount>
.
</paragraph>
<paragraph id="2F018046FC5EFF81FE9C029482919F66" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FE9C0294811D9E95" box="[199,372,1196,1221]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">
<emphasis id="1DCA5C54FC5EFF81FE9C029481049E93" bold="true" box="[199,365,1196,1221]" pageId="14" pageNumber="465">Type material</emphasis>
.
</emphasis>
<materialsCitation id="9FD68A1BFC5EFF81FF27029482BC9EBE" collectingDate="2011-10-02" collectorName="Bosque &amp; Coll. F. Camposeco." country="Honduras" elevation="1580" location="Sula" pageId="14" pageNumber="465" specimenCount="1" stateProvince="Cortes" typeStatus="holotype">
<typeStatus id="F0053EE4FC5EFF81FF270294818D9E93" box="[380,484,1196,1221]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="57A9C0D6FC5EFF81FC74029482C99E93" box="[559,672,1196,1221]" name="Honduras" pageId="14" pageNumber="465">Honduras</collectingCountry>
:
<collectingRegion id="ED7A4EA4FC5EFF81FCF00294829E9E93" box="[683,759,1196,1221]" country="Honduras" name="Cortes" pageId="14" pageNumber="465">Cortés</collectingRegion>
, cerca de San Pedro
<location id="2A61D69DFC5EFF81FD840294847A9E93" LSID="urn:lsid:plazi:treatment:A7173150FC5BFF81FECC0114809B9C5E:2A61D69DFC5EFF81FD840294847A9E93" box="[991,1043,1196,1221]" country="Honduras" name="Sula" pageId="14" pageNumber="465" stateProvince="Cortes">Sula</location>
, 15.512598°, -88.113660°
<quantity id="E8462DA3FC5EFF81FB1B029485FE9E95" box="[1344,1431,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="14" pageNumber="465" unit="m" value="1580.0">
<elevation id="A4936775FC5EFF81FB1B029485FE9E95" box="[1344,1431,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="14" pageNumber="465" unit="m" value="1580.0">1580 m</elevation>
</quantity>
,
<date id="5B00A686FC5EFF81FECC02F781649EBE" box="[151,269,1231,1256]" pageId="14" pageNumber="465" value="2011-10-02">
<collectingDate id="4B445F6EFC5EFF81FECC02F781649EBE" box="[151,269,1231,1256]" pageId="14" pageNumber="465" value="2011-10-02">2 X 2011</collectingDate>
</date>
,
<collectorName id="824BE590FC5EFF81FF4502E8811B9EB1" box="[286,370,1232,1255]" pageId="14" pageNumber="465">Bosque</collectorName>
nuboso.
<collectorName id="824BE590FC5EFF81FFBE02F782BC9EBE" box="[485,725,1231,1256]" pageId="14" pageNumber="465">Coll. F. Camposeco.</collectorName>
</materialsCitation>
<materialsCitation id="9FD68A1BFC5EFF81FCB902E881659F5B" collectingDate="2011-10-02" collectorName="Bosque &amp; Coll. F. Camposeco." country="Honduras" elevation="1580" location="Same" pageId="14" pageNumber="465" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Cortes" typeStatus="paratype">
<typeStatus id="F0053EE4FC5EFF81FCB902E8833A9EBE" box="[738,851,1232,1256]" pageId="14" pageNumber="465" type="paratype">Paratypes</typeStatus>
:
<location id="2A61D69DFC5EFF81FD3F02F783CB9EBE" LSID="urn:lsid:plazi:treatment:A7173150FC5BFF81FECC0114809B9C5E:2A61D69DFC5EFF81FD3F02F783CB9EBE" box="[868,930,1231,1256]" name="Same" pageId="14" pageNumber="465">Same</location>
data as holotype (
<specimenCount id="39B84BCFFC5EFF81FAD302F784819EBE" box="[1160,1256,1231,1256]" pageId="14" pageNumber="465" type="male">5 males</specimenCount>
,
<specimenCount id="39B84BCFFC5EFF81FAA202E885079EBE" box="[1273,1390,1231,1256]" pageId="14" pageNumber="465" type="female">7 females</specimenCount>
, 30 unknown)
</materialsCitation>
.
<materialsCitation id="9FD68A1BFC5EFF81FF4302CC829A9F66" collectingDate="1987-03-20" collectingDateMax="1987-03-21" collectingDateMin="1987-03-20" collectionCode="VI" collectorName="S. Pedro Sula &amp; J. C. Schuster &amp; Type C" country="Honduras" elevation="1550" location="Cortes" pageId="14" pageNumber="465" specimenCount="3" specimenCount-female="3" stateProvince="Cortes" typeStatus="paratype">
<collectingCountry id="57A9C0D6FC5EFF81FF4302CC81E09F5B" box="[280,393,1268,1293]" name="Honduras" pageId="14" pageNumber="465">Honduras</collectingCountry>
:
<collectingRegion id="ED7A4EA4FC5EFF81FFCD02CC81B69F5B" box="[406,479,1268,1293]" country="Honduras" name="Cortes" pageId="14" pageNumber="465">Cortés</collectingRegion>
Dept.
<quantity id="E8462DA3FC5EFF81FC7502CC821E9F5B" box="[558,631,1268,1293]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="465" unit="km" value="30.0">30 km</quantity>
W of
<collectorName id="824BE590FC5EFF81FC9802CC830B9F5B" box="[707,866,1268,1293]" pageId="14" pageNumber="465">S. Pedro Sula</collectorName>
,
<quantity id="E8462DA3FC5EFF81FD3502CC83AC9F5D" box="[878,965,1268,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="465" unit="m" value="1550.0">
<elevation id="A4936775FC5EFF81FD3502CC83AC9F5D" box="[878,965,1268,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="465" unit="m" value="1550.0">1550 m</elevation>
</quantity>
alt.,
<date id="5B00A686FC5EFF81FA5A02CC84C39F5B" box="[1025,1194,1268,1293]" pageId="14" pageNumber="465" value="1987-03-20" valueMax="1987-03-21" valueMin="1987-03-20">
<collectingDate id="4B445F6EFC5EFF81FA5A02CC84C39F5B" box="[1025,1194,1268,1293]" pageId="14" pageNumber="465" value="1987-03-20" valueMax="1987-03-21" valueMin="1987-03-20">20-21 III 1987</collectingDate>
</date>
, bosque nuboso,
<collectorName id="824BE590FC5EFF81FB2A02CC80909F66" pageId="14" pageNumber="465">J.C. Schuster</collectorName>
, #
<collectionCode id="49AF1883FC5EFF81FF42032F81519F66" box="[281,312,1303,1328]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="465">VI</collectionCode>
<specimenCount id="39B84BCFFC5EFF81FF63032F81F29F66" box="[312,411,1303,1328]" pageId="14" pageNumber="465" type="female">3 female</specimenCount>
,
<collectorName id="824BE590FC5EFF81FFFD032081919F66" box="[422,504,1303,1328]" pageId="14" pageNumber="465">Type C</collectorName>
disturbance sound (1)
</materialsCitation>
.
</paragraph>
<paragraph id="2F018046FC5EFF81FE9C0304853B9F03" blockId="14.[151,1437,151,1544]" box="[199,1362,1340,1365]" pageId="14" pageNumber="465">
<materialsCitation id="9FD68A1BFC5EFF81FE9C0304827E9F03" box="[199,535,1340,1365]" collectionCode="UVGC" pageId="14" pageNumber="465" specimenCount="1" typeStatus="holotype">
<typeStatus id="F0053EE4FC5EFF81FE9C030481469F03" box="[199,303,1340,1365]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
deposited at
<collectionCode id="49AF1883FC5EFF81FF9F0304827E9F03" box="[452,535,1340,1365]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="14" pageNumber="465">UVGC</collectionCode>
</materialsCitation>
.
<materialsCitation id="9FD68A1BFC5EFF81FC7A030484F49F03" box="[545,1181,1340,1365]" collectionCode="UVGC, IEXA, USAC, RC, SNM" pageId="14" pageNumber="465" specimenCount="1" typeStatus="paratype">
<typeStatus id="F0053EE4FC5EFF81FC7A030482E79F02" box="[545,654,1340,1364]" pageId="14" pageNumber="465" type="paratype">Paratypes</typeStatus>
deposited in
<collectionCode id="49AF1883FC5EFF81FD7F0304831F9F03" box="[804,886,1340,1365]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="14" pageNumber="465">UVGC</collectionCode>
,
<collectionCode id="49AF1883FC5EFF81FDDA030483AE9F03" box="[897,967,1340,1365]" pageId="14" pageNumber="465">IEXA</collectionCode>
,
<collectionCode id="49AF1883FC5EFF81FD89030484769F03" box="[978,1055,1340,1365]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="14" pageNumber="465">USAC</collectionCode>
,
<collectionCode id="49AF1883FC5EFF81FA700304843A9F03" box="[1067,1107,1340,1365]" pageId="14" pageNumber="465">RC</collectionCode>
,
<collectionCode id="49AF1883FC5EFF81FA05030484F49F03" box="[1118,1181,1340,1365]" country="Slovakia" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="14" pageNumber="465">SNM</collectionCode>
</materialsCitation>
, and
<collectionCode id="49AF1883FC5EFF81FA83030485249F03" box="[1240,1357,1340,1365]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="14" pageNumber="465" type="University or college">IBUNAM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF81FE9C036781A79F96" pageId="14" pageNumber="465" type="etymology">
<paragraph id="2F018046FC5EFF81FE9C036781A79F96" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FE9C036781279F2E" bold="true" box="[199,334,1375,1400]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">Etymology.</emphasis>
Named after colleagues of the National Science Foundation project LLAMA (Leaf Litter Arthropods of Mesoamerica), led by Jack Longino and Bob Anderson. Several
<taxonomicNameLabel id="06F9E12FFC5EFF81FA4403BC84C19FCB" box="[1055,1192,1412,1437]" pageId="14" pageNumber="465" rank="species">new species</taxonomicNameLabel>
were collected under the umbrella of this project.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC5EFF81FE9C03F4809B9C5E" pageId="14" pageNumber="465" type="distribution">
<paragraph id="2F018046FC5EFF81FE9C03F4809B9C5E" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="465">
<emphasis id="1DCA5C54FC5EFF81FE9C03F481099FB3" bold="true" box="[199,352,1484,1509]" inLineHeading="true" pageId="14" pageNumber="465" reason="1">Distribution.</emphasis>
This species is known from cloud forests of middle altitude near
<collectingCountry id="57A9C0D6FC5EFF81FA0303F484A59FB3" box="[1112,1228,1484,1509]" name="Saint Pierre and Miquelon" pageId="14" pageNumber="465">San Pedro</collectingCountry>
Sula in
<collectingCountry id="57A9C0D6FC5EFF81FB7403F485F59FB3" box="[1327,1436,1484,1509]" name="Honduras" pageId="14" pageNumber="465">Honduras</collectingCountry>
(
<figureCitation id="B7859CC3FC5EFF81FEC403D7808C9C5E" box="[159,229,1519,1544]" captionStart="FIGURE 4" captionStartId="6.[151,250,1904,1926]" captionTargetBox="[171,1398,927,1870]" captionTargetId="figure-358@6.[149,1434,899,1893]" captionTargetPageId="6" captionText="FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi." figureDoi="http://doi.org/10.5281/zenodo.4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="14" pageNumber="465">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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