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<document id="6CEA27EDA0A6B402ADA1AB2E6A042438" ID-DOI="10.11646/zootaxa.3889.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4952403" ID-ZooBank="1F3F076D-A4F1-4B4A-977D-4B99581C6779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730508465" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2014" docId="A7173150FC55FF86FECC02C5829D9BDE" docLanguage="en" docName="zt03889p484.pdf" docOrigin="Zootaxa 3889 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ogyges handali Cano 2014, new species" docType="treatment" docVersion="4" lastPageNumber="460" masterDocId="5B2E4928FC50FF8FFE5B063880699A56" masterDocTitle="Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly" masterLastPageNumber="484" masterPageNumber="451" pageNumber="456" updateTime="1698967030688" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1BDEEAA7C4F53CD1A7CB9EA9DE028452">Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly</mods:title>
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<mods:namePart id="8F83116CFBE5523326294045C204BF33">Cano, Enio B.</mods:namePart>
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<mods:date id="B5380850BB0EA5E7E795B256741BF451">2014</mods:date>
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<mods:number id="FAA3B684C79FE8191C9EABC70A066071">2014-12-03</mods:number>
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<mods:number id="38E5EC8139BF08BEE5772398C253F266">3889</mods:number>
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<mods:classification id="673FA16479389D0CE4C4D29368E2380E">journal article</mods:classification>
<mods:identifier id="EC2D63F71943EFEEBEC05CBE737AA7A7" type="DOI">10.11646/zootaxa.3889.4.1</mods:identifier>
<mods:identifier id="EFB5E5F1C58D7AE1F4EE86D43FC704C6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A7173150FC55FF86FECC02C5829D9BDE" ID-DOI="http://doi.org/10.5281/zenodo.4953255" ID-Zenodo-Dep="4953255" LSID="urn:lsid:plazi:treatment:A7173150FC55FF86FECC02C5829D9BDE" httpUri="http://treatment.plazi.org/id/A7173150FC55FF86FECC02C5829D9BDE" lastPageId="9" lastPageNumber="460" pageId="5" pageNumber="456">
<subSubSection id="67A4D3CDFC55FF8AFECC02C582559F41" box="[151,572,1277,1303]" pageId="5" pageNumber="456" type="nomenclature">
<paragraph id="2F018046FC55FF8AFECC02C582559F41" blockId="5.[151,572,1277,1337]" box="[151,572,1277,1303]" pageId="5" pageNumber="456">
<heading id="7449372AFC55FF8AFECC02C582559F41" bold="true" box="[151,572,1277,1303]" fontSize="11" level="1" pageId="5" pageNumber="456" reason="1">
<emphasis id="1DCA5C54FC55FF8AFECC02C582559F41" bold="true" box="[151,572,1277,1303]" pageId="5" pageNumber="456">
<taxonomicName id="E8BEFBC5FC55FF8AFECC02C581F79F41" authority="Cano" authorityName="Cano" authorityYear="2014" box="[151,414,1277,1303]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="handali" status="sp. nov.">
<emphasis id="1DCA5C54FC55FF8AFECC02C5813A9F41" bold="true" box="[151,339,1277,1303]" italics="true" pageId="5" pageNumber="456">Ogyges handali</emphasis>
Cano
</taxonomicName>
,
<taxonomicNameLabel id="06F9E12FFC55FF8AFFF702C582559F41" box="[428,572,1277,1303]" pageId="5" pageNumber="456" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC55FF8AFECC031881589F6F" box="[151,305,1312,1337]" pageId="5" pageNumber="456" type="description">
<paragraph id="2F018046FC55FF8AFECC031881589F6F" blockId="5.[151,572,1277,1337]" box="[151,305,1312,1337]" pageId="5" pageNumber="456">
<figureCitation id="B7859CC3FC55FF8AFECC031881689F6F" box="[151,257,1312,1337]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1947,1969]" captionStartId-1="8.[151,250,1899,1921]" captionTargetBox-0="[177,1436,628,1912]" captionTargetBox-1="[168,1425,204,1875]" captionTargetId-0="figure-233@7.[151,1436,628,1912]" captionTargetId-1="figure-51@8.[151,1437,193,1878]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Paratype of Ogyges handali: a. lateral view. b. dorsal view. c. ventral view. d. ventral view of head and prothorax. e. anterolateral view of head." captionText-1="FIGURE 6. Ogyges handali: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952415" figureDoi-1="http://doi.org/10.5281/zenodo.4952417" httpUri-0="https://zenodo.org/record/4952415/files/figure.png" httpUri-1="https://zenodo.org/record/4952417/files/figure.png" pageId="5" pageNumber="456">Figs. 5–6</figureCitation>
,
<figureCitation id="B7859CC3FC55FF8AFF56031881589F6F" box="[269,305,1312,1337]" captionStart="FIGURE 21" captionStartId="28.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure-318@28.[151,1436,193,1354]" captionTargetPageId="28" captionText="FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili." figureDoi="http://doi.org/10.5281/zenodo.4952455" httpUri="https://zenodo.org/record/4952455/files/figure.png" pageId="5" pageNumber="456">21i</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC55FF8AFECC035080919FBB" pageId="5" pageNumber="456" type="diagnosis">
<paragraph id="2F018046FC55FF8AFECC035080919FBB" blockId="5.[151,1437,1384,2021]" pageId="5" pageNumber="456">
<emphasis id="1DCA5C54FC55FF8AFECC035081789FD7" bold="true" box="[151,273,1384,1409]" inLineHeading="true" pageId="5" pageNumber="456" reason="1">Diagnosis.</emphasis>
<taxonomicName id="E8BEFBC5FC55FF8AFF42035181A39FD6" authorityName="Cano" authorityYear="2014" box="[281,458,1384,1408]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="handali">
<emphasis id="1DCA5C54FC55FF8AFF42035181A39FD6" box="[281,458,1384,1408]" italics="true" pageId="5" pageNumber="456">Ogyges handali</emphasis>
</taxonomicName>
resembles
<taxonomicName id="E8BEFBC5FC55FF8AFC100351839E9FD7" authority="Schuster &amp; Reyes-Castillo" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" box="[587,1015,1384,1409]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="adamsi">
<emphasis id="1DCA5C54FC55FF8AFC10035182A99FD6" box="[587,704,1384,1408]" italics="true" pageId="5" pageNumber="456">O. adamsi</emphasis>
Schuster &amp; Reyes-Castillo
</taxonomicName>
, by the absence of internal tubercles, the transversal convex clypeus, and the form of the central horn and lateroposterior tubercles. The new species is easily separated of
<taxonomicName id="E8BEFBC5FC55FF8AFF21038981989F9E" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" box="[378,497,1456,1480]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="adamsi">
<emphasis id="1DCA5C54FC55FF8AFF21038981989F9E" box="[378,497,1456,1480]" italics="true" pageId="5" pageNumber="456">O. adamsi</emphasis>
</taxonomicName>
by the abundant setae on anterior corners of metasternum (sparse or bare in
<taxonomicName id="E8BEFBC5FC55FF8AFBDA038980819FBA" authorityName="Schuster &amp; Reyes-Castillo" authorityYear="1990" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="adamsi">
<emphasis id="1DCA5C54FC55FF8AFBDA038980819FBA" italics="true" pageId="5" pageNumber="456">O. adamsi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC55FF88FE9C03C0808E9B16" lastPageId="7" lastPageNumber="458" pageId="5" pageNumber="456" type="description">
<paragraph id="2F018046FC55FF89FE9C03C082379B86" blockId="5.[151,1437,1384,2021]" lastBlockId="6.[151,1437,151,861]" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456">
<emphasis id="1DCA5C54FC55FF8AFE9C03C081309C47" bold="true" box="[199,345,1528,1553]" inLineHeading="true" pageId="5" pageNumber="456" reason="1">Description.</emphasis>
<typeStatus id="F0053EE4FC55FF8AFF3E03C081A49C47" box="[357,461,1528,1553]" pageId="5" pageNumber="456" type="holotype">Holotype</typeStatus>
male, black adult: Total length 33.67, elytral length 18.51, pronotal length 8.45, pronotal width 11.16, humeral width 10.31. Head: anterior border of labrum concave. Clypeus thickened, transversely convex, anterior border straight, with a central notch in form of “v”; with rugose sulcus separating from the inclined mediofrontal area. Mediofrontal tubercles small and rounded; internal tubercles absent. Laterofrontal areas roughened. Posterofrontal ridges absent; instead, a transversal ridge (tumose near the central horn) runs between the mediopostfrontal structure and the epicranial sutures, forming the anterior margin of lateropostfrontal areas. Mediofrontal area with several grooves confluent to the clypeus, with a wide, bare, rough fossae present in front of the mediopostfrontal structure. Lateropostfrontal areas glabrous, deep, with several grooves. Central horn long, with apex short and free; posterior 1/3 tumose and clearly differentiated from the lower lateroposterior tubercles and with a deep median longitudinal groove; lateroposterior tubercles lower than central horn, barely marked, separated from central horn and directed forward. Postfrontal groove shallow at middle and laterally deep. Supraorbital ridge brilliant and rugose, with unequal anterior tubercles; posterior 1/2 barely bifurcate but with supraorbital fossae elongate, well marked; external ridge rounded. Ocular canthus with apex slightly swollen, rounded, ventrally extended just to half of eye. Eyes reduced. Eye width =
<quantity id="E8462DA3FC55FF8AFAF501F4857F9DB2" box="[1198,1302,1996,2021]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="456" unit="mm" value="0.31">0.31 mm</quantity>
. Interocular width =
<quantity id="E8462DA3FC56FF89FEAE06AF81349AF9" box="[245,349,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.61" pageId="6" pageNumber="457" unit="mm" value="6.61">6.61 mm</quantity>
. Head (measured from tips of canthi)
<quantity id="E8462DA3FC56FF89FD5306AF83199AF9" box="[776,880,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="8.03" pageId="6" pageNumber="457" unit="mm" value="8.03">8.03 mm</quantity>
; ratio width of both eyes/head = 0.08. Postorbital pits punctuate setose. Ligula slightly protuberant basally, with apical central tooth small and anterior transversal carina absent; abundant setose punctures on the median area, basally sparse. Lateral lobes of mentum with moderate setose punctures, more abundant towards the brilliant oval lateral basal scars; medial basal mentum glabrous and smooth. Hypostomal process elongate, without lateral depression, wide medially and narrow in the apical third. Infraocular ridge present, short and smooth, surrounded by setose punctures. Mandible with dorsal tooth occupying 1/2 of the length; internal face of mandible in dorsal view granular. Antennal club (
<figureCitation id="B7859CC3FC56FF89FAAE0757855E9BDE" box="[1269,1335,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[177,1436,628,1912]" captionTargetId="figure-233@7.[151,1436,628,1912]" captionTargetPageId="7" captionText="FIGURE 5. Paratype of Ogyges handali: a. lateral view. b. dorsal view. c. ventral view. d. ventral view of head and prothorax. e. anterolateral view of head." figureDoi="http://doi.org/10.5281/zenodo.4952415" httpUri="https://zenodo.org/record/4952415/files/figure.png" pageId="6" pageNumber="457">Fig. 5</figureCitation>
) with all three antennomeres very wide and subequal measuring
<quantity id="E8462DA3FC56FF89FD6E07AC83C99BFD" box="[821,928,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="6" pageNumber="457" unit="mm" value="1.77">1.77 mm</quantity>
long and
<quantity id="E8462DA3FC56FF89FA7F07AC84E69BFD" box="[1060,1167,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.31" pageId="6" pageNumber="457" unit="mm" value="2.31">2.31 mm</quantity>
wide; antepenultimate antennomere narrower than penultimate.
</paragraph>
<paragraph id="2F018046FC56FF89FE9C07E483D598FE" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="457">Thorax: Lateral fossa of pronotum without punctures except sparse, rugose micropunctures ahead and behind, only visible at very great magnification. Pronotum with marginal groove narrow and smooth, anterior angles rounded, disc smooth at moderate magnification. Prosternellum wide and brilliant. Mesosternum bare and shiny, without definite opaque/rugose (shagreened) areas. Mesepisternum brilliant. Metasternum with anterior angles punctate setose; disc not delimited by punctures; marginal groove very narrow and rugose, with disperse setae in the anterior 4/5 of its length, posteriorly four times wider than medially.</paragraph>
<paragraph id="2F018046FC56FF89FE9C048C81CD9943" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="457">Elytra: Brilliant, with well-marked striations; with small, defined, moderately deep punctures; junctions of striations 1 and 10 without extra punctures. Anterior border of elytra vertical, smooth, with minute scattered setae between interstriae 2–8.</paragraph>
<paragraph id="2F018046FC56FF89FE9C05188543996E" blockId="6.[151,1437,151,861]" box="[199,1322,799,824]" pageId="6" pageNumber="457">Legs: Profemur with anterioventral groove marked; metafemur elongated; mesotibia with one spine.</paragraph>
<paragraph id="2F018046FC56FF89FE9C057C842B990B" blockId="6.[151,1437,151,861]" box="[199,1090,836,861]" pageId="6" pageNumber="457">Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.</paragraph>
<caption id="7BC1D0CEFC56FF89FECC0148828F9D95" ID-DOI="http://doi.org/10.5281/zenodo.4952413" ID-Zenodo-Dep="4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,250,1904,1926]" targetBox="[171,1398,927,1870]" targetPageId="6">
<paragraph id="2F018046FC56FF89FECC0148828F9D95" blockId="6.[151,1436,1904,1987]" pageId="6" pageNumber="457">
<emphasis id="1DCA5C54FC56FF89FECC0148817A9DD3" bold="true" box="[151,275,1904,1926]" pageId="6" pageNumber="457">FIGURE 4.</emphasis>
Map of northern Central America showing the locality of the new species. Light blue square,
<taxonomicName id="E8BEFBC5FC56FF89FA80014885FC9DD0" authorityName="Cano" authorityYear="2014" box="[1243,1429,1904,1926]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="menchuae">
<emphasis id="1DCA5C54FC56FF89FA80014885FC9DD0" box="[1243,1429,1904,1926]" italics="true" pageId="6" pageNumber="457">Ogyges menchuae</emphasis>
</taxonomicName>
; rose circle,
<taxonomicName id="E8BEFBC5FC56FF89FF5601B681109DF2" authorityName="Cano" authorityYear="2014" box="[269,377,1934,1956]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="handali">
<emphasis id="1DCA5C54FC56FF89FF5601B681109DF2" box="[269,377,1934,1956]" italics="true" pageId="6" pageNumber="457">O. handali</emphasis>
</taxonomicName>
; blue rhombus,
<taxonomicName id="E8BEFBC5FC56FF89FC4401B6821D9DF2" authorityName="Cano" authorityYear="2014" box="[543,628,1934,1956]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="cavei">
<emphasis id="1DCA5C54FC56FF89FC4401B6821D9DF2" box="[543,628,1934,1956]" italics="true" pageId="6" pageNumber="457">O. cavei</emphasis>
</taxonomicName>
; yellow triangle,
<taxonomicName id="E8BEFBC5FC56FF89FD7301B683E89DF2" authorityName="Cano" authorityYear="2014" box="[808,897,1934,1956]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="llama">
<emphasis id="1DCA5C54FC56FF89FD7301B683E89DF2" box="[808,897,1934,1956]" italics="true" pageId="6" pageNumber="457">O. llama</emphasis>
</taxonomicName>
,
<taxonomicName id="E8BEFBC5FC56FF89FDD501B6844C9DF2" authorityName="Cano" authorityYear="2014" box="[910,1061,1934,1956]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="mutenroshii">
<emphasis id="1DCA5C54FC56FF89FDD501B6844C9DF2" box="[910,1061,1934,1956]" italics="true" pageId="6" pageNumber="457">O. mutenroshii</emphasis>
</taxonomicName>
; purple cross,
<taxonomicName id="E8BEFBC5FC56FF89FAE701B685549DF2" authorityName="Cano" authorityYear="2014" box="[1212,1341,1934,1956]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="toriyamai">
<emphasis id="1DCA5C54FC56FF89FAE701B685549DF2" box="[1212,1341,1934,1956]" italics="true" pageId="6" pageNumber="457">O. toriyamai</emphasis>
</taxonomicName>
; red star,
<taxonomicName id="E8BEFBC5FC56FF89FECC019580919D95" authorityName="Cano" authorityYear="2014" box="[151,248,1965,1987]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="laurae">
<emphasis id="1DCA5C54FC56FF89FECC019580919D95" box="[151,248,1965,1987]" italics="true" pageId="6" pageNumber="457">O. laurae</emphasis>
</taxonomicName>
,
<taxonomicName id="E8BEFBC5FC56FF89FF5F019581139D95" authorityName="Cano" authorityYear="2014" box="[260,378,1965,1987]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="1DCA5C54FC56FF89FF5F019581139D95" box="[260,378,1965,1987]" italics="true" pageId="6" pageNumber="457">O. ratcliffei</emphasis>
</taxonomicName>
, and light green circle,
<taxonomicName id="E8BEFBC5FC56FF89FC31019582899D95" authorityName="Cano" authorityYear="2014" box="[618,736,1965,1987]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="sandinoi">
<emphasis id="1DCA5C54FC56FF89FC31019582899D95" box="[618,736,1965,1987]" italics="true" pageId="6" pageNumber="457">O. sandinoi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="2F018046FC57FF88FE9C06AF830C9A83" blockId="7.[151,1436,151,575]" pageId="7" pageNumber="458">
Aedeagus: In ventral view phallus globose; parameres and phallobase partially separated. In dorsal view, ventrodorsal basal sclerotizations of the phallus present (
<figureCitation id="B7859CC3FC57FF88FD480684833E9A83" box="[787,855,188,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1899,1921]" captionTargetBox="[168,1425,204,1875]" captionTargetId="figure-51@8.[151,1437,193,1878]" captionTargetPageId="8" captionText="FIGURE 6. Ogyges handali: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4952417" httpUri="https://zenodo.org/record/4952417/files/figure.png" pageId="7" pageNumber="458">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="2F018046FC57FF88FE9C06E7808E9B16" blockId="7.[151,1436,151,575]" pageId="7" pageNumber="458">
Variation in
<typeStatus id="F0053EE4FC57FF88FF0B06D881D29AAE" box="[336,443,224,248]" pageId="7" pageNumber="458" type="paratype">paratypes</typeStatus>
(
<emphasis id="1DCA5C54FC57FF88FF9006D981B09AAE" box="[459,473,225,248]" italics="true" pageId="7" pageNumber="458">n</emphasis>
= 19): Total length 30.54–35.58 ( = 33.00), elytral length = 16.06–19.17 ( = 17.97), pronotal length = 7.5–8.76 ( = 8.44), pronotal width = 9.7–11.54 ( = 11.03), humeral width = 9.16–11.27 ( = 10.36).
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC57FF86FE9C077485E99AF9" lastPageId="9" lastPageNumber="460" pageId="7" pageNumber="458" type="materials_examined">
<paragraph id="2F018046FC57FF88FE9C0774823E9B33" blockId="7.[151,1436,151,575]" box="[199,599,332,357]" pageId="7" pageNumber="458">
<emphasis id="1DCA5C54FC57FF88FE9C077481D99B35" box="[199,432,332,357]" inLineHeading="true" pageId="7" pageNumber="458" reason="1">
<emphasis id="1DCA5C54FC57FF88FE9C077481C09B33" bold="true" box="[199,425,332,357]" pageId="7" pageNumber="458">Material examined</emphasis>
.
</emphasis>
<specimenCount id="39B84BCFFC57FF88FFEC0774823A9B33" box="[439,595,332,357]" pageId="7" pageNumber="458" type="generic">49 specimens</specimenCount>
.
</paragraph>
<paragraph id="2F018046FC57FF88FE9C075783679869" blockId="7.[151,1436,151,575]" pageId="7" pageNumber="458">
<emphasis id="1DCA5C54FC57FF88FE9C0757811D9BD1" box="[199,372,367,392]" inLineHeading="true" pageId="7" pageNumber="458" reason="1">
<emphasis id="1DCA5C54FC57FF88FE9C075781079BDE" bold="true" box="[199,366,367,392]" pageId="7" pageNumber="458">Type material</emphasis>
.
</emphasis>
<materialsCitation id="9FD68A1BFC57FF88FF27075784B09BF9" collectingDate="2011-06-11" collectorName="El Duraznal &amp; Plan de la Arada &amp; Col. E. B. Cano." country="Guatemala" county="Santa Rosalia" elevation="1668" latitude="14.525111" location="Plan de la Arada" longLatPrecision="1" longitude="-89.37981" municipality="El Duraznal" pageId="7" pageNumber="458" specimenCount="1" stateProvince="Chiquimula" typeStatus="holotype">
<typeStatus id="F0053EE4FC57FF88FF270757818D9BDE" box="[380,484,367,392]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="57A9C0D6FC57FF88FC6B0757828A9BDE" box="[560,739,367,392]" name="Guatemala" pageId="7" pageNumber="458">GUATEMALA</collectingCountry>
:
<collectingRegion id="ED7A4EA4FC57FF88FCB60757831D9BDE" box="[749,884,367,392]" country="Guatemala" name="Chiquimula" pageId="7" pageNumber="458">Chiquimula</collectingRegion>
, aldea
<collectingCounty id="C660F8CAFC57FF88FD99075784369BDE" box="[962,1119,367,392]" pageId="7" pageNumber="458">Santa Rosalía</collectingCounty>
,
<collectorName id="824BE590FC57FF88FA320757849A9BDE" box="[1129,1267,367,392]" pageId="7" pageNumber="458">
<collectingMunicipality id="CF651A3CFC57FF88FA320757849A9BDE" box="[1129,1267,367,392]" pageId="7" pageNumber="458">El Duraznal</collectingMunicipality>
</collectorName>
, cerca del
<collectorName id="824BE590FC57FF88FB31075781749BF9" pageId="7" pageNumber="458">
<location id="2A61D69DFC57FF88FB31075781749BF9" LSID="urn:lsid:plazi:treatment:A7173150FC55FF86FECC02C5829D9BDE:2A61D69DFC57FF88FB31075781749BF9" country="Guatemala" county="Santa Rosalia" latitude="14.525111" longLatPrecision="1" longitude="-89.37981" municipality="El Duraznal" name="Plan de la Arada" pageId="7" pageNumber="458" stateProvince="Chiquimula">Plan de la Arada</location>
</collectorName>
,
<date id="5B00A686FC57FF88FF7007AE81C49BF9" box="[299,429,406,431]" pageId="7" pageNumber="458" value="2011-06-11">
<collectingDate id="4B445F6EFC57FF88FF7007AE81C49BF9" box="[299,429,406,431]" pageId="7" pageNumber="458" value="2011-06-11">11 VI 2011</collectingDate>
</date>
,
<geoCoordinate id="4A8AE681FC57FF88FFE007AE82389BF9" box="[443,593,403,431]" degrees="14" direction="north" minutes="31" orientation="latitude" pageId="7" pageNumber="458" precision="1" seconds="30.4" value="14.525111">
14
<superScript id="D8CB2D0EFC57FF88FF8D07AB81B49BF7" attach="none" box="[470,477,403,417]" fontSize="6" pageId="7" pageNumber="458">o</superScript>
31'30.4''N
</geoCoordinate>
,
<geoCoordinate id="4A8AE681FC57FF88FC0507AE82919BF9" box="[606,760,403,431]" degrees="89" direction="west" minutes="22" orientation="longitude" pageId="7" pageNumber="458" precision="1" seconds="47.3" value="-89.37981">
89
<superScript id="D8CB2D0EFC57FF88FC2107AB82E89BF7" attach="none" box="[634,641,403,417]" fontSize="6" pageId="7" pageNumber="458">o</superScript>
22'47.3''W
</geoCoordinate>
,
<quantity id="E8462DA3FC57FF88FD5E07AE83379BF8" box="[773,862,406,431]" metricMagnitude="3" metricUnit="m" metricValue="1.668" pageId="7" pageNumber="458" unit="m" value="1668.0">
<elevation id="A4936775FC57FF88FD5E07AE83379BF8" box="[773,862,406,431]" metricMagnitude="3" metricUnit="m" metricValue="1.668" pageId="7" pageNumber="458" unit="m" value="1668.0">1668 m</elevation>
</quantity>
, bosque nuboso,
<collectorName id="824BE590FC57FF88FA7F07AE84B09BF9" box="[1060,1241,406,431]" pageId="7" pageNumber="458">Col. E.B. Cano.</collectorName>
</materialsCitation>
<materialsCitation id="9FD68A1BFC57FF88FABA07AF80929BA1" collectingDate="2011-04-01" collectingDateMax="2011-04-04" collectingDateMin="2011-04-01" collectorName="Same &amp; Col. C. Suchite" country="Guatemala" county="Santa Rosalia" elevation="1700" latitude="14.525111" location="Same" longLatPrecision="1" longitude="-89.37981" municipality="El Duraznal" pageId="7" pageNumber="458" specimenCount="27" specimenCount-female="15" specimenCount-male="12" stateProvince="Chiquimula" typeStatus="paratype">
<typeStatus id="F0053EE4FC57FF88FABA07AF853B9BF9" box="[1249,1362,407,431]" pageId="7" pageNumber="458" type="paratype">Paratypes</typeStatus>
:
<collectorName id="824BE590FC57FF88FB0507AE85F59BF9" box="[1374,1436,406,431]" pageId="7" pageNumber="458">Same</collectorName>
data as holotype (
<specimenCount id="39B84BCFFC57FF88FF3A078281C79B85" box="[353,430,442,467]" pageId="7" pageNumber="458" type="male">1 male</specimenCount>
,
<specimenCount id="39B84BCFFC57FF88FFE1078282489B85" box="[442,545,442,467]" pageId="7" pageNumber="458" type="female">1 female</specimenCount>
).
<location id="2A61D69DFC57FF88FC6F0782821B9B85" LSID="urn:lsid:plazi:treatment:A7173150FC55FF86FECC02C5829D9BDE:2A61D69DFC57FF88FC6F0782821B9B85" box="[564,626,442,467]" name="Same" pageId="7" pageNumber="458">Same</location>
data except
<date id="5B00A686FC57FF88FD5A078283F89B85" box="[769,913,442,467]" pageId="7" pageNumber="458" value="2011-04-01" valueMax="2011-04-04" valueMin="2011-04-01">
<collectingDate id="4B445F6EFC57FF88FD5A078283F89B85" box="[769,913,442,467]" pageId="7" pageNumber="458" value="2011-04-01" valueMax="2011-04-04" valueMin="2011-04-01">1–4 IV 2011</collectingDate>
</date>
, aprox.
<quantity id="E8462DA3FC57FF88FDB70782842A9B84" box="[1004,1091,442,467]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="458" unit="m" value="1700.0">
<elevation id="A4936775FC57FF88FDB70782842A9B84" box="[1004,1091,442,467]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="458" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<collectorName id="824BE590FC57FF88FA0B078284949B85" box="[1104,1277,442,467]" pageId="7" pageNumber="458">Col. C. Suchité</collectorName>
(
<specimenCount id="39B84BCFFC57FF88FB560782851D9B85" box="[1293,1396,442,467]" pageId="7" pageNumber="458" type="male">11 males</specimenCount>
,
<specimenCount id="39B84BCFFC57FF88FBDB0782809A9BA1" pageId="7" pageNumber="458" type="female">14 females</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="9FD68A1BFC57FF88FF5307E68149984D" collectingDate="1998-08-20" collectorName="de La Mesilla &amp; Cols. &amp; Cano &amp; J. Monzon" country="Guatemala" elevation="1630" location="El Duraznal" municipality="Same" pageId="7" pageNumber="458" specimenCount="1" stateProvince="Chiquimula" typeStatus="paratype">
<collectingMunicipality id="CF651A3CFC57FF88FF5307E6812F9BA1" box="[264,326,478,503]" pageId="7" pageNumber="458">Same</collectingMunicipality>
data except aldea
<location id="2A61D69DFC57FF88FC7B07E682C49BA1" LSID="urn:lsid:plazi:treatment:A7173150FC55FF86FECC02C5829D9BDE:2A61D69DFC57FF88FC7B07E682C49BA1" box="[544,685,478,503]" country="Guatemala" municipality="Same" name="El Duraznal" pageId="7" pageNumber="458" stateProvince="Chiquimula">El Duraznal</location>
, cerca
<collectorName id="824BE590FC57FF88FCA407E683C89BA1" box="[767,929,478,503]" pageId="7" pageNumber="458">de La Mesilla</collectorName>
,
<quantity id="E8462DA3FC57FF88FDF407E684609BA0" box="[943,1033,478,503]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="458" unit="m" value="1630.0">
<elevation id="A4936775FC57FF88FDF407E684609BA0" box="[943,1033,478,503]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="458" unit="m" value="1630.0">1630 m</elevation>
</quantity>
,
<date id="5B00A686FC57FF88FA4307E684D89BA1" box="[1048,1201,478,503]" pageId="7" pageNumber="458" value="1998-08-20">
<collectingDate id="4B445F6EFC57FF88FA4307E684D89BA1" box="[1048,1201,478,503]" pageId="7" pageNumber="458" value="1998-08-20">20 VIII 1998</collectingDate>
</date>
,
<collectorName id="824BE590FC57FF88FA9B07E684939BA1" box="[1216,1274,478,503]" pageId="7" pageNumber="458">Cols.</collectorName>
E.
<collectorName id="824BE590FC57FF88FB7E07E685099BA1" box="[1317,1376,478,503]" pageId="7" pageNumber="458">Cano</collectorName>
&amp;
<collectorName id="824BE590FC57FF88FBD107E7809D984D" pageId="7" pageNumber="458">J. Monzón</collectorName>
(5)
</materialsCitation>
.
<materialsCitation id="9FD68A1BFC57FF88FF77043A82BB9869" collectingDate="1998-06-24" collectionCode="VI" collectorName="Las Presas &amp; el Plan de la Arada &amp; Col. E. Cano" country="Guatemala" county="San Jose" elevation="1900" location="Bosque" municipality="Minas" pageId="7" pageNumber="458" specimenCount="1" stateProvince="Guatemala" typeStatus="paratype">
<collectingRegion id="ED7A4EA4FC57FF88FF77043A81C3984D" box="[300,426,514,539]" country="Guatemala" name="Guatemala" pageId="7" pageNumber="458">Guatemala</collectingRegion>
:
<collectingRegion id="ED7A4EA4FC57FF88FFED043A8254984D" box="[438,573,514,539]" country="Guatemala" name="Chiquimula" pageId="7" pageNumber="458">Chiquimula</collectingRegion>
,
<collectingCounty id="C660F8CAFC57FF88FC12043A82C5984D" box="[585,684,514,539]" pageId="7" pageNumber="458">San José</collectingCounty>
las
<collectingMunicipality id="CF651A3CFC57FF88FC87043A834C984D" box="[732,805,514,539]" pageId="7" pageNumber="458">Minas</collectingMunicipality>
, camino entre caserío
<collectorName id="824BE590FC57FF88FA7F043B84F5984C" box="[1060,1180,515,538]" pageId="7" pageNumber="458">Las Presas</collectorName>
y
<collectorName id="824BE590FC57FF88FAE0043A85F1984D" box="[1211,1432,514,539]" pageId="7" pageNumber="458">el Plan de la Arada</collectorName>
,
<location id="2A61D69DFC57FF88FECC041F80829868" LSID="urn:lsid:plazi:treatment:A7173150FC55FF86FECC02C5829D9BDE:2A61D69DFC57FF88FECC041F80829868" box="[151,235,551,574]" country="Guatemala" county="San Jose" municipality="Minas" name="Bosque" pageId="7" pageNumber="458" stateProvince="Guatemala">Bosque</location>
nuboso.
<quantity id="E8462DA3FC57FF88FF0A041E81C19868" box="[337,424,550,575]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="458" unit="m" value="1900.0">
<elevation id="A4936775FC57FF88FF0A041E81C19868" box="[337,424,550,575]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="458" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="5B00A686FC57FF88FFEF041E825F9869" box="[436,566,550,575]" pageId="7" pageNumber="458" value="1998-06-24">
<collectingDate id="4B445F6EFC57FF88FFEF041E825F9869" box="[436,566,550,575]" pageId="7" pageNumber="458" value="1998-06-24">
24
<collectionCode id="49AF1883FC57FF88FF8D041F819A9868" box="[470,499,551,574]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="458">VI</collectionCode>
1998
</collectingDate>
</date>
,
<collectorName id="824BE590FC57FF88FC1A041E82BB9869" box="[577,722,550,575]" pageId="7" pageNumber="458">Col. E. Cano</collectorName>
</materialsCitation>
(16).
</paragraph>
<caption id="7BC1D0CEFC57FF88FECC01A381D49D86" ID-DOI="http://doi.org/10.5281/zenodo.4952415" ID-Zenodo-Dep="4952415" httpUri="https://zenodo.org/record/4952415/files/figure.png" pageId="7" pageNumber="458" startId="7.[151,250,1947,1969]" targetBox="[177,1436,628,1912]" targetPageId="7">
<paragraph id="2F018046FC57FF88FECC01A381D49D86" blockId="7.[151,1437,1947,2000]" pageId="7" pageNumber="458">
<emphasis id="1DCA5C54FC57FF88FECC01A3817A9DE6" bold="true" box="[151,275,1947,1969]" pageId="7" pageNumber="458">FIGURE 5.</emphasis>
Paratype of
<taxonomicName id="E8BEFBC5FC57FF88FFCF01A3825D9DE7" authorityName="Cano" authorityYear="2014" box="[404,564,1947,1969]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="handali">
<emphasis id="1DCA5C54FC57FF88FFCF01A3825D9DE7" box="[404,564,1947,1969]" italics="true" pageId="7" pageNumber="458">Ogyges handali</emphasis>
</taxonomicName>
: a. lateral view. b. dorsal view. c. ventral view. d. ventral view of head and prothorax. e. anterolateral view of head.
</paragraph>
</caption>
<caption id="7BC1D0CEFC58FF87FECC015380929DF6" ID-DOI="http://doi.org/10.5281/zenodo.4952417" ID-Zenodo-Dep="4952417" httpUri="https://zenodo.org/record/4952417/files/figure.png" pageId="8" pageNumber="459" startId="8.[151,250,1899,1921]" targetBox="[168,1425,204,1875]" targetPageId="8">
<paragraph id="2F018046FC58FF87FECC015380929DF6" blockId="8.[151,1435,1899,1952]" pageId="8" pageNumber="459">
<emphasis id="1DCA5C54FC58FF87FECC0153817B9DD6" bold="true" box="[151,274,1899,1921]" pageId="8" pageNumber="459">FIGURE 6.</emphasis>
<taxonomicName id="E8BEFBC5FC58FF87FF43015381DE9DD7" authorityName="Cano" authorityYear="2014" box="[280,439,1899,1921]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="handali">
<emphasis id="1DCA5C54FC58FF87FF43015381DE9DD7" box="[280,439,1899,1921]" italics="true" pageId="8" pageNumber="459">Ogyges handali</emphasis>
</taxonomicName>
: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus.
</paragraph>
</caption>
<paragraph id="2F018046FC59FF86FE9C06AF85E99AF9" blockId="9.[151,1437,151,392]" box="[199,1408,151,176]" pageId="9" pageNumber="460">
<materialsCitation id="9FD68A1BFC59FF86FE9C06AF85E99AF9" box="[199,1408,151,176]" collectionCode="UVGC, USAC, IEXA, IBUNAM, SNM, MNHN" pageId="9" pageNumber="460" specimenCount="2" typeStatus="holotype">
<typeStatus id="F0053EE4FC59FF86FE9C06AF81469AE6" box="[199,303,151,176]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
deposited at
<collectionCode id="49AF1883FC59FF86FF9F06AF827F9AE6" box="[452,534,151,176]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="9" pageNumber="460">UVGC</collectionCode>
,
<typeStatus id="F0053EE4FC59FF86FC7A06A082E59AE6" box="[545,652,152,176]" pageId="9" pageNumber="460" type="paratype">paratypes</typeStatus>
deposited in
<collectionCode id="49AF1883FC59FF86FD7906AF831D9AE6" box="[802,884,151,176]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="9" pageNumber="460">UVGC</collectionCode>
,
<collectionCode id="49AF1883FC59FF86FDDB06AF83A79AE6" box="[896,974,151,176]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="9" pageNumber="460">USAC</collectionCode>
,
<collectionCode id="49AF1883FC59FF86FD8306AF84779AE6" box="[984,1054,151,176]" pageId="9" pageNumber="460">IEXA</collectionCode>
,
<collectionCode id="49AF1883FC59FF86FA7206AF84F79AE6" box="[1065,1182,151,176]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="9" pageNumber="460" type="University or college">IBUNAM</collectionCode>
,
<collectionCode id="49AF1883FC59FF86FAF206AF84819AE6" box="[1193,1256,151,176]" country="Slovakia" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="460">SNM</collectionCode>
, and
<collectionCode id="49AF1883FC59FF86FB7806A085159AF9" box="[1315,1404,152,175]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC59FF86FE9C068480969B4B" pageId="9" pageNumber="460" type="etymology">
<paragraph id="2F018046FC59FF86FE9C068480969B4B" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="460">
<emphasis id="1DCA5C54FC59FF86FE9C068481249A85" box="[199,333,188,213]" inLineHeading="true" pageId="9" pageNumber="460" reason="1">
<emphasis id="1DCA5C54FC59FF86FE9C0684812F9A83" bold="true" box="[199,326,188,213]" pageId="9" pageNumber="460">Etymology</emphasis>
.
</emphasis>
Named in honor of Commander Dr. Schafik Handal, officially recognized as “Hijo Meritísimo de la Ciudad de
<collectingRegion id="ED7A4EA4FC59FF86FF7106E781A89AAE" box="[298,449,223,248]" country="El Salvador" name="San Salvador" pageId="9" pageNumber="460">San Salvador</collectingRegion>
” and “Honor al Mérito Centroamericano”, for his efforts in favor of the peace in Central America.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC59FF86FE9C071F829D9BDE" pageId="9" pageNumber="460" type="distribution">
<paragraph id="2F018046FC59FF86FE9C071F829D9BDE" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="460">
<emphasis id="1DCA5C54FC59FF86FE9C071F81099B69" box="[199,352,295,320]" inLineHeading="true" pageId="9" pageNumber="460" reason="1">
<emphasis id="1DCA5C54FC59FF86FE9C071F81309B16" bold="true" box="[199,345,295,320]" pageId="9" pageNumber="460">Distribution</emphasis>
.
</emphasis>
The species is known only from the fragmented cloud forests of Cerro Montecristo between
<quantity id="E8462DA3FC59FF86FECC0774815A9B35" box="[151,307,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.77" metricValueMax="1.9" metricValueMin="1.64" pageId="9" pageNumber="460" unit="m" value="1770.0" valueMax="1900.0" valueMin="1640.0">1640–1900 m</quantity>
in
<collectingCountry id="57A9C0D6FC59FF86FF03077481B89B33" box="[344,465,332,357]" name="Guatemala" pageId="9" pageNumber="460">Guatemala</collectingCountry>
(
<figureCitation id="B7859CC3FC59FF86FFB9077482419B33" box="[482,552,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1904,1926]" captionTargetBox="[171,1398,927,1870]" captionTargetId="figure-358@6.[149,1434,899,1893]" captionTargetPageId="6" captionText="FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi." figureDoi="http://doi.org/10.5281/zenodo.4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="9" pageNumber="460">Fig. 4</figureCitation>
). The new species should be present in the protected forest of the
<collectingCountry id="57A9C0D6FC59FF86FB43077485F49B33" box="[1304,1437,332,357]" name="El Salvador" pageId="9" pageNumber="460">El Salvador</collectingCountry>
portion and the bigger forest of the Honduran portion.
</paragraph>
</subSubSection>
</treatment>
</document>
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