284 lines
36 KiB
XML
284 lines
36 KiB
XML
<document id="74E417A94DF8077204FAA182C7477F99" ID-DOI="10.11646/zootaxa.3889.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4952403" ID-ZooBank="1F3F076D-A4F1-4B4A-977D-4B99581C6779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730508465" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2014" docId="A7173150FC46FF98FECC035382949D26" docLanguage="en" docName="zt03889p484.pdf" docOrigin="Zootaxa 3889 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ogyges sandinoi Cano 2014, new species" docType="treatment" docVersion="4" lastPageNumber="474" masterDocId="5B2E4928FC50FF8FFE5B063880699A56" masterDocTitle="Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly" masterLastPageNumber="484" masterPageNumber="451" pageNumber="473" updateTime="1698967030688" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="41D26E4104423B68A85C9A1D4DF24CB2">Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly</mods:title>
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<mods:namePart id="10F3DFB1E29B88E9707869202C909310">Cano, Enio B.</mods:namePart>
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<mods:title id="8E8AA9F1862A8DCAC166283279CADC46">Zootaxa</mods:title>
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<mods:part id="DB3E50E3E0D046E6C61C991DBB8E02BA">
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<mods:date id="75A871B0090065C950156FB17C7D422D">2014</mods:date>
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<mods:number id="36D3C5B985599E11F2A732E2B2846AF5">2014-12-03</mods:number>
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<mods:number id="870A36423A50F652D86DEE34FB983C2B">3889</mods:number>
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<mods:identifier id="43CECFA0FB2877FEB6F3ECACFCE7E7BD" type="DOI">10.11646/zootaxa.3889.4.1</mods:identifier>
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<mods:identifier id="A2DBC3C4BD39A3389C0CA524B65F0713" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A7173150FC46FF98FECC035382949D26" ID-DOI="http://doi.org/10.5281/zenodo.4953263" ID-Zenodo-Dep="4953263" LSID="urn:lsid:plazi:treatment:A7173150FC46FF98FECC035382949D26" httpUri="http://treatment.plazi.org/id/A7173150FC46FF98FECC035382949D26" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
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<subSubSection id="67A4D3CDFC46FF99FECC035382569FD3" box="[151,575,1387,1413]" pageId="22" pageNumber="473" type="nomenclature">
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<paragraph id="2F018046FC46FF99FECC035382569FD3" blockId="22.[151,575,1387,1446]" box="[151,575,1387,1413]" pageId="22" pageNumber="473">
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<heading id="7449372AFC46FF99FECC035382569FD3" bold="true" box="[151,575,1387,1413]" fontSize="11" level="1" pageId="22" pageNumber="473" reason="1">
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<emphasis id="1DCA5C54FC46FF99FECC035382569FD3" bold="true" box="[151,575,1387,1413]" pageId="22" pageNumber="473">
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<taxonomicName id="E8BEFBC5FC46FF99FECC035381C19FD3" authority="Cano" authorityName="Cano" authorityYear="2014" box="[151,424,1387,1413]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="sandinoi" status="sp. nov.">
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<emphasis id="1DCA5C54FC46FF99FECC035381379FD3" bold="true" box="[151,350,1387,1413]" italics="true" pageId="22" pageNumber="473">Ogyges sandinoi</emphasis>
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Cano
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</taxonomicName>
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<taxonomicNameLabel id="06F9E12FFC46FF99FFEB035382569FD3" box="[432,575,1387,1413]" pageId="22" pageNumber="473" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="67A4D3CDFC46FF99FECC03B581309FF0" box="[151,345,1421,1446]" pageId="22" pageNumber="473" type="description">
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<paragraph id="2F018046FC46FF99FECC03B581309FF0" blockId="22.[151,575,1387,1446]" box="[151,345,1421,1446]" pageId="22" pageNumber="473">
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<figureCitation id="B7859CC3FC46FF99FECC03B581779FF0" box="[151,286,1421,1446]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="24.[151,250,1864,1886]" captionStartId-1="25.[151,250,1662,1684]" captionTargetBox-0="[155,1436,215,1837]" captionTargetBox-1="[162,1430,202,1630]" captionTargetId-0="figure-47@24.[151,1436,193,1842]" captionTargetId-1="figure-117@25.[150,1436,193,1641]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 17. Holotype of Ogyges sandinoi: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 18. Ogyges sandinoi: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952447" figureDoi-1="http://doi.org/10.5281/zenodo.4952449" httpUri-0="https://zenodo.org/record/4952447/files/figure.png" httpUri-1="https://zenodo.org/record/4952449/files/figure.png" pageId="22" pageNumber="473">
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Figs. 17–
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<geoCoordinate id="4A8AE681FC46FF99FF5903B581779FF0" box="[258,286,1421,1446]" degrees="18, 21" direction="west" orientation="longitude" pageId="22" pageNumber="473" precision="555" value="-18.21">18</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="B7859CC3FC46FF99FF7203B581309FF0" box="[297,345,1421,1446]" captionStart="FIGURE 21" captionStartId="28.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure-318@28.[151,1436,193,1354]" captionTargetPageId="28" captionText="FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili." figureDoi="http://doi.org/10.5281/zenodo.4952455" httpUri="https://zenodo.org/record/4952455/files/figure.png" pageId="22" pageNumber="473">21w</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="67A4D3CDFC46FF99FECC03ED816D9C0D" pageId="22" pageNumber="473" type="diagnosis">
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<paragraph id="2F018046FC46FF99FECC03ED816D9C0D" blockId="22.[151,1437,1493,2022]" pageId="22" pageNumber="473">
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<emphasis id="1DCA5C54FC46FF99FECC03ED81789FB8" bold="true" box="[151,273,1493,1518]" inLineHeading="true" pageId="22" pageNumber="473" reason="1">Diagnosis.</emphasis>
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Based on the form of clypeus, the central horn, and the mediopostfrontal structure, and the presence of setae on anterior corners of metasternum (sparse in
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<taxonomicName id="E8BEFBC5FC46FF99FC8603C283389C44" authorityName="Schuster & Reyes-Castillo" authorityYear="1990" box="[733,849,1530,1554]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="adamsi">
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<emphasis id="1DCA5C54FC46FF99FC8603C283389C44" box="[733,849,1530,1554]" italics="true" pageId="22" pageNumber="473">O. adamsi</emphasis>
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</taxonomicName>
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),
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<taxonomicName id="E8BEFBC5FC46FF99FD3303C283859C44" authorityName="Cano" authorityYear="2014" box="[872,1004,1530,1554]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="sandinoi">
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<emphasis id="1DCA5C54FC46FF99FD3303C283859C44" box="[872,1004,1530,1554]" italics="true" pageId="22" pageNumber="473">O. sandinoi</emphasis>
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</taxonomicName>
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seems closely allied to
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<taxonomicName id="E8BEFBC5FC46FF99FAA303C285059C44" authorityName="Schuster & Reyes-Castillo" authorityYear="1990" box="[1272,1388,1530,1554]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="adamsi">
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<emphasis id="1DCA5C54FC46FF99FAA303C285059C44" box="[1272,1388,1530,1554]" italics="true" pageId="22" pageNumber="473">O. adamsi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="E8BEFBC5FC46FF99FECC002681799C60" authorityName="Cano" authorityYear="2014" box="[151,272,1566,1590]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="handali">
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<emphasis id="1DCA5C54FC46FF99FECC002680C29C63" box="[151,171,1566,1589]" italics="true" pageId="22" pageNumber="473">O</emphasis>
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.
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<emphasis id="1DCA5C54FC46FF99FEE0002681799C60" box="[187,272,1566,1590]" italics="true" pageId="22" pageNumber="473">handali</emphasis>
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</taxonomicName>
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. This new species is easily distinguished by the smaller size (
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<quantity id="E8462DA3FC46FF99FD88002584399C60" box="[979,1104,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="22" pageNumber="473" unit="mm" value="30.5" valueMax="32.0" valueMin="29.0">29–32 mm</quantity>
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) and the presence of internal tubercles.
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</paragraph>
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</subSubSection>
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<subSubSection id="67A4D3CDFC46FF98FE9C005D84179E93" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473" type="description">
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<paragraph id="2F018046FC46FF98FE9C005D8084986B" blockId="22.[151,1437,1493,2022]" lastBlockId="23.[151,1437,151,1904]" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
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<emphasis id="1DCA5C54FC46FF99FE9C005D81309C28" bold="true" box="[199,345,1637,1662]" inLineHeading="true" pageId="22" pageNumber="473" reason="1">Description.</emphasis>
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<typeStatus id="F0053EE4FC46FF99FF39005D81A39C28" box="[354,458,1637,1662]" pageId="22" pageNumber="473" type="holotype">Holotype</typeStatus>
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male, black adult with dark-brown areas in venter. Total length 29.68, elytral length 16.00, pronotal length 8.26, pronotal width 10.27, humeral width 9.81. Head: anterior border of labrum concave or slightly concave. Clypeus thickened, convex, anterior border straight or slightly inclined laterally, with a small central indentation in form of “v”; with a moderately defined punctate sulcus separating it from the mediofrontal area. Mediofrontal tubercles rounded and well defined, with a well or barely defined anterofrontal ridges directed but not connected with the internal tubercles. Mediofrontal area smooth, with conical internal tubercles connected to the short and slightly curved posterofrontal ridges. A wide, bare, hemicircular smooth fossa present in front of the mediopostfrontal structure. Lateropostfrontal areas glabrous and slightly rugose towards lateral sides. Central horn very narrow and long, with apex not free, declivous towards the posterofrontal ridges, with dorsal micropuntations; without median longitudinal dorsal groove; lateroposterior tubercles short, not clearly delineated, rounded, directed slightly posteriorly, not separated from the central horn by a groove. Postfrontal groove shallow, deeper at sides. Supraorbital ridge with unequal anterior tubercles; posterior half not bifurcate with supraorbital fossa small; external ridge poorly marked; supraorbital fossa shallow and short. Ocular canthus with apex straight or slightly swollen, ventrally covering 3/4 of eye. Eyes reduced. Eye width =
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<quantity id="E8462DA3FC47FF98FA4A06E784139AA1" box="[1041,1146,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="23" pageNumber="474" unit="mm" value="0.38">0.38 mm</quantity>
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. Interocular width =
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<quantity id="E8462DA3FC47FF98FB3006E780AF9B4D" metricMagnitude="-3" metricUnit="m" metricValue="5.77" pageId="23" pageNumber="474" unit="mm" value="5.77">5.77 mm</quantity>
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. Head width =
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<quantity id="E8462DA3FC47FF98FFD0073C81909B4D" box="[395,505,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.38" pageId="23" pageNumber="474" unit="mm" value="7.38">7.38 mm</quantity>
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; ratio both eye widths/head width (measured between tips of canthi) = 0.10. Postorbital pits located immediately behind supraocular ridge and eye, punctate setose. Ligula almost flat basally, with small central apical tooth, ventral transversal anterior carina absent, with setaceous punctures medially. Lateral lobes of mentum setose punctate; lateral basal scars oval and brilliant; medial basal mentum bare and shiny. Hypostomal process elongate, with shallow lateral depression and with one or two minute setae, wide medially and narrow in apical third. Infraocular ridge present, short and smooth, surrounded by striate, setose punctures. Dorsal tooth of mandible occupies 1/2 length; internal dorsal face of mandible granular. Antennal club concave, measures
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<quantity id="E8462DA3FC47FF98FECC07C780929841" box="[151,251,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="23" pageNumber="474" unit="mm" value="1.92">1.92 mm</quantity>
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long and
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<quantity id="E8462DA3FC47FF98FF3007C781A69841" box="[363,463,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="23" pageNumber="474" unit="mm" value="2.15">2.15 mm</quantity>
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wide; antepenultimate antennomere of antennal club less wide than penultimate (
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<figureCitation id="B7859CC3FC47FF98FB3F07C78089986B" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="24.[151,250,1864,1886]" captionStartId-1="25.[151,250,1662,1684]" captionTargetBox-0="[155,1436,215,1837]" captionTargetBox-1="[162,1430,202,1630]" captionTargetId-0="figure-47@24.[151,1436,193,1842]" captionTargetId-1="figure-117@25.[150,1436,193,1641]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 17. Holotype of Ogyges sandinoi: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view." captionText-1="FIGURE 18. Ogyges sandinoi: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4952447" figureDoi-1="http://doi.org/10.5281/zenodo.4952449" httpUri-0="https://zenodo.org/record/4952447/files/figure.png" httpUri-1="https://zenodo.org/record/4952449/files/figure.png" pageId="23" pageNumber="474">Figs. 17–18</figureCitation>
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).
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</paragraph>
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<paragraph id="2F018046FC47FF98FE9C047F8549996E" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">Thorax: Lateral fossae of pronotum without punctures but, together with the marginal grooves, with sparse micropuntations of rugose aspect, visible with great magnification. Pronotum with marginal groove narrow and smooth; disc smooth, slightly opaque; anterior angles rounded; with a small, but present, posterior medial notch. Prosternellum shagreened on anterior half and brilliant posteriorly, except slightly opaque on sides. Mesosternum glabrous; lateral depressions wide, elongate and rugose (shagreened). Mesepisternum with oval, elongate, shagreened area. Metasternum anterior angles with moderate setae; disc not delimited by punctures; marginal groove with setose punctures anteriorly; narrow and smooth, posteriorly two times wider than medially.</paragraph>
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<paragraph id="2F018046FC47FF98FE9C057C813499F3" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">Elytra: Brilliant, striations marked, with small, defined, moderately deep punctures; junctions of striations 1 and 10 without extra punctures. Vertical anterior area smooth, with minute to small, scattered setae between portion of interstriae 1–8.</paragraph>
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<paragraph id="2F018046FC47FF98FE9C0588816099BB" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">Legs: Profemur with anterioventral groove poorly marked, but present; metafemur elongate; mesotibia with one spine.</paragraph>
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<paragraph id="2F018046FC47FF98FE9C05CF842B9E46" blockId="23.[151,1437,151,1904]" box="[199,1090,1015,1040]" pageId="23" pageNumber="474">Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.</paragraph>
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<paragraph id="2F018046FC47FF98FE9C0224831A9E0E" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
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Aedeagus: In ventral view phallus globose; parameres and phallobase scarcely separated laterally; ventrodorsal basal sclerotizations of the phallus present (
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<figureCitation id="B7859CC3FC47FF98FD480207830F9E0E" box="[787,870,1087,1112]" captionStart="FIGURE 18" captionStartId="25.[151,250,1662,1684]" captionTargetBox="[162,1430,202,1630]" captionTargetId="figure-117@25.[150,1436,193,1641]" captionTargetPageId="25" captionText="FIGURE 18. Ogyges sandinoi: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4952449" httpUri="https://zenodo.org/record/4952449/files/figure.png" pageId="23" pageNumber="474">Fig. 18</figureCitation>
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).
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</paragraph>
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<paragraph id="2F018046FC47FF98FE9C025C84179E93" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
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Variation in
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<typeStatus id="F0053EE4FC47FF98FF0E025C81A99E2A" box="[341,448,1124,1148]" pageId="23" pageNumber="474" type="paratype">paratypes</typeStatus>
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(
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<emphasis id="1DCA5C54FC47FF98FF89025D81899E2A" box="[466,480,1125,1148]" italics="true" pageId="23" pageNumber="474">n</emphasis>
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= 4): Total length 30.02–31.63 ( = 30.68), elytral length 15.62–17.62 ( = 16.60), pronotal length 7.76–8.91 ( = 8.20), pronotal width 10.23–11.19 ( = 10.58), humeral width 9.53–10.31 ( = 9.80). One specimen with the antennomeres of the left antennal club fused (a teratology).
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</paragraph>
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</subSubSection>
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<subSubSection id="67A4D3CDFC47FF98FE9C02F7843C9F03" pageId="23" pageNumber="474" type="materials_examined">
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<paragraph id="2F018046FC47FF98FE9C02F782219EBE" blockId="23.[151,1437,151,1904]" box="[199,584,1231,1256]" pageId="23" pageNumber="474">
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<emphasis id="1DCA5C54FC47FF98FE9C02F781D99EB1" box="[199,432,1231,1256]" inLineHeading="true" pageId="23" pageNumber="474" reason="1">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C02F781C09EBE" bold="true" box="[199,425,1231,1256]" pageId="23" pageNumber="474">Material examined</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="39B84BCFFC47FF98FFEC02F7822D9EBE" box="[439,580,1231,1256]" pageId="23" pageNumber="474" type="generic">5 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2F018046FC47FF98FE9C02CC838D9F66" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C02CC811D9F5D" box="[199,372,1268,1293]" inLineHeading="true" pageId="23" pageNumber="474" reason="1">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C02CC81049F5B" bold="true" box="[199,365,1268,1293]" pageId="23" pageNumber="474">Type material</emphasis>
|
||
.
|
||
</emphasis>
|
||
<materialsCitation id="9FD68A1BFC47FF98FF2002CC825A9F66" collectingDate="2011-05-28" collectingDateMax="2011-05-31" collectingDateMin="2011-05-28" collectorName="L. Saenz" country="Nicaragua" latitude="13.98079" location="Jalapa" longLatPrecision="1" longitude="-86.17922" pageId="23" pageNumber="474" specimenCount="1" stateProvince="Nueva Segovia" typeStatus="holotype">
|
||
<typeStatus id="F0053EE4FC47FF98FF2002CC81819F5B" box="[379,488,1268,1293]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="57A9C0D6FC47FF98FFAA02CC82F59F5B" box="[497,668,1268,1293]" name="Nicaragua" pageId="23" pageNumber="474">NICARAGUA</collectingCountry>
|
||
,
|
||
<collectingRegion id="ED7A4EA4FC47FF98FCFC02CC833D9F5B" box="[679,852,1268,1293]" country="Nicaragua" name="Nueva Segovia" pageId="23" pageNumber="474">Nueva Segovia</collectingRegion>
|
||
, nr
|
||
<location id="2A61D69DFC47FF98FD2602CC83AF9F5B" LSID="urn:lsid:plazi:treatment:A7173150FC46FF98FECC035382949D26:2A61D69DFC47FF98FD2602CC83AF9F5B" box="[893,966,1268,1293]" country="Nicaragua" latitude="13.98079" longLatPrecision="1" longitude="-86.17922" name="Jalapa" pageId="23" pageNumber="474" stateProvince="Nueva Segovia">Jalapa</location>
|
||
, Cerro Jesús.
|
||
<geoCoordinate id="4A8AE681FC47FF98FA3B02CC84A59F5B" box="[1120,1228,1268,1293]" degrees="13.98079" direction="north" orientation="latitude" pageId="23" pageNumber="474" precision="1" value="13.98079">13.98079</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4A8AE681FC47FF98FA8D02CC85249F5B" box="[1238,1357,1268,1293]" degrees="86.17922" direction="west" orientation="longitude" pageId="23" pageNumber="474" precision="1" value="-86.17922">-86.17922</geoCoordinate>
|
||
.
|
||
<date id="5B00A686FC47FF98FB0D02CC81629F66" pageId="23" pageNumber="474" value="2011-05-28" valueMax="2011-05-31" valueMin="2011-05-28">
|
||
<collectingDate id="4B445F6EFC47FF98FB0D02CC81629F66" pageId="23" pageNumber="474" value="2011-05-28" valueMax="2011-05-31" valueMin="2011-05-28">28–31 May 2011</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="824BE590FC47FF98FF4D032081119F66" box="[278,376,1303,1328]" pageId="23" pageNumber="474">L. Sáenz</collectorName>
|
||
collector (LSD)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="F0053EE4FC47FF98FC66032082C39F66" box="[573,682,1304,1328]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
|
||
(4): Same data as holotype.
|
||
</paragraph>
|
||
<paragraph id="2F018046FC47FF98FE9C0304843C9F03" blockId="23.[151,1437,151,1904]" box="[199,1109,1340,1365]" pageId="23" pageNumber="474">
|
||
<materialsCitation id="9FD68A1BFC47FF98FE9C030482719F03" box="[199,536,1340,1365]" collectionCode="UVGC" pageId="23" pageNumber="474" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="F0053EE4FC47FF98FE9C030481469F03" box="[199,303,1340,1365]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
|
||
deposited in
|
||
<collectionCode id="49AF1883FC47FF98FF9E030482719F03" box="[453,536,1340,1365]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="474">UVGC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9FD68A1BFC47FF98FC79030483A09F03" box="[546,969,1340,1365]" collectionCode="UVGC, IEXA" pageId="23" pageNumber="474" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="F0053EE4FC47FF98FC79030482E69F02" box="[546,655,1340,1364]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode id="49AF1883FC47FF98FD7D030483119F03" box="[806,888,1340,1365]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="474">UVGC</collectionCode>
|
||
,
|
||
<collectionCode id="49AF1883FC47FF98FDD8030483A09F03" box="[899,969,1340,1365]" pageId="23" pageNumber="474">IEXA</collectionCode>
|
||
</materialsCitation>
|
||
, and
|
||
<collectionCode id="49AF1883FC47FF98FA58030484389F03" box="[1027,1105,1340,1365]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="23" pageNumber="474">USAC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A4D3CDFC47FF98FE9C0367811E9FCB" pageId="23" pageNumber="474" type="etymology">
|
||
<paragraph id="2F018046FC47FF98FE9C0367811E9FCB" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C036781279F2E" bold="true" box="[199,334,1375,1400]" inLineHeading="true" pageId="23" pageNumber="474" reason="1">Etymology.</emphasis>
|
||
The name of this species is in honor of Augusto César Sandino, a Nicaraguan hero, born in the Segovia Mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A4D3CDFC47FF98FE9C039F81FB9FB3" pageId="23" pageNumber="474" type="distribution">
|
||
<paragraph id="2F018046FC47FF98FE9C039F81FB9FB3" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C039F81099F96" bold="true" box="[199,352,1447,1472]" inLineHeading="true" pageId="23" pageNumber="474" reason="1">Distribution.</emphasis>
|
||
Only known from Cerro Jesús, at the Nicaraguan-Honduran border (
|
||
<figureCitation id="B7859CC3FC47FF98FACA039F84B39F96" box="[1169,1242,1447,1472]" captionStart="FIGURE 4" captionStartId="6.[151,250,1904,1926]" captionTargetBox="[171,1398,927,1870]" captionTargetId="figure-358@6.[149,1434,899,1893]" captionTargetPageId="6" captionText="FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi." figureDoi="http://doi.org/10.5281/zenodo.4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="23" pageNumber="474">Fig. 4</figureCitation>
|
||
), at mid-altitude cloud forest (
|
||
<quantity id="E8462DA3FC47FF98FF7003F481EC9FB3" box="[299,389,1484,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="474" unit="m" value="1600.0">1600 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A4D3CDFC47FF98FE9C03D782949D26" pageId="23" pageNumber="474" type="discussion">
|
||
<paragraph id="2F018046FC47FF98FE9C03D782949D26" blockId="23.[151,1437,151,1904]" pageId="23" pageNumber="474">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C03D781269C51" box="[199,335,1519,1544]" inLineHeading="true" pageId="23" pageNumber="474" reason="1">
|
||
<emphasis id="1DCA5C54FC47FF98FE9C03D781219C5E" bold="true" box="[199,328,1519,1544]" pageId="23" pageNumber="474">Comments</emphasis>
|
||
.
|
||
</emphasis>
|
||
This is the first objective record of the genus
|
||
<taxonomicName id="E8BEFBC5FC47FF98FD3103C883D79C51" box="[874,958,1520,1543]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1DCA5C54FC47FF98FD3103C883D79C51" box="[874,958,1520,1543]" italics="true" pageId="23" pageNumber="474">Ogyges</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="57A9C0D6FC47FF98FA5D03D784149C5E" box="[1030,1149,1519,1544]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
|
||
. Previously, two species,
|
||
<taxonomicName id="E8BEFBC5FC47FF98FECC002C811B9C7A" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[151,370,1556,1580]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="laevissimus">
|
||
<emphasis id="1DCA5C54FC47FF98FECC002C811B9C7A" box="[151,370,1556,1580]" italics="true" pageId="23" pageNumber="474">Ogyges laevissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E8BEFBC5FC47FF98FFF3002C82569C7A" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[424,575,1556,1580]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="1DCA5C54FC47FF98FFF3002C82569C7A" box="[424,575,1556,1580]" italics="true" pageId="23" pageNumber="474">O. championi</emphasis>
|
||
</taxonomicName>
|
||
[as
|
||
<taxonomicName id="E8BEFBC5FC47FF98FC36002C82B79C7A" box="[621,734,1556,1580]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="laevior">
|
||
<emphasis id="1DCA5C54FC47FF98FC36002C82B79C7A" box="[621,734,1556,1580]" italics="true" pageId="23" pageNumber="474">O. laevior</emphasis>
|
||
</taxonomicName>
|
||
] have been cited from that country (summarized in
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FB44002C81C19C06" author="Schuster, J. C. & Reyes-Castillo, P." pageId="23" pageNumber="474" pagination="1 - 49" refId="ref19543" refString="Schuster, J. C. & Reyes-Castillo, P. (1990) Coleoptera, Passalidae: Ogyges Kaup, revision de un genero mesoamericano de montana. Acta Zoologica Mexicana (n. s.), 40, 1 - 49." type="journal article" year="1990">Schuster & Reyes-Castillo 1990: 19</bibRefCitation>
|
||
, 32–33). Because these citations are ambiguous, and following
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FA28000F80889C23" author="Schuster, J. C. & Reyes-Castillo, P." pageId="23" pageNumber="474" pagination="1 - 49" refId="ref19543" refString="Schuster, J. C. & Reyes-Castillo, P. (1990) Coleoptera, Passalidae: Ogyges Kaup, revision de un genero mesoamericano de montana. Acta Zoologica Mexicana (n. s.), 40, 1 - 49." type="journal article" year="1990">Schuster & Reyes-Castillo (1990)</bibRefCitation>
|
||
who restricted the species to
|
||
<collectingCountry id="57A9C0D6FC47FF98FC1A006482D39C23" box="[577,698,1628,1653]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
|
||
(
|
||
<taxonomicName id="E8BEFBC5FC47FF98FC950064831C9C22" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[718,885,1628,1652]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="laevissimus">
|
||
<emphasis id="1DCA5C54FC47FF98FC950064831C9C22" box="[718,885,1628,1652]" italics="true" pageId="23" pageNumber="474">O. laevissimus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<collectingCountry id="57A9C0D6FC47FF98FDE70064845C9C23" box="[956,1077,1628,1653]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
|
||
and
|
||
<collectingCountry id="57A9C0D6FC47FF98FA28006484A19C23" box="[1139,1224,1628,1653]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
|
||
(
|
||
<taxonomicName id="E8BEFBC5FC47FF98FA87006485119C22" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[1244,1400,1628,1652]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="1DCA5C54FC47FF98FA87006485119C22" box="[1244,1400,1628,1652]" italics="true" pageId="23" pageNumber="474">O. championi</emphasis>
|
||
</taxonomicName>
|
||
), I consider these or any other citation of
|
||
<taxonomicName id="E8BEFBC5FC47FF98FC1D00B882809CCE" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[582,745,1664,1688]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="laevissimus">
|
||
<emphasis id="1DCA5C54FC47FF98FC1D00B882809CCE" box="[582,745,1664,1688]" italics="true" pageId="23" pageNumber="474">O. laevissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E8BEFBC5FC47FF98FD7B00B883D19CCE" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[800,952,1664,1688]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="1DCA5C54FC47FF98FD7B00B883D19CCE" box="[800,952,1664,1688]" italics="true" pageId="23" pageNumber="474">O. championi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="57A9C0D6FC47FF98FDA6004784199CCE" box="[1021,1136,1663,1688]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
|
||
as erroneous. As evidence of this ambiguity, the records of
|
||
<taxonomicName id="E8BEFBC5FC47FF98FC4C009C82D49CEA" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[535,701,1700,1724]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="laevissimus">
|
||
<emphasis id="1DCA5C54FC47FF98FC4C009C82D49CEA" box="[535,701,1700,1724]" italics="true" pageId="23" pageNumber="474">O. laevissimus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="57A9C0D6FC47FF98FD52009C83159CEB" box="[777,892,1700,1725]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FDD6009C840D9CEB" author="Kuwert, A. F." box="[909,1124,1700,1725]" pageId="23" pageNumber="474" pagination="274 - 306" refId="ref19098" refString="Kuwert, A. F. (1897) Die Passaliden dichotomisch bearbeitet, die Arten. Novitates Zoologicae, 4, 274 - 306." type="journal article" year="1897">Kuwert 1897: 291</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FA28009C85FE9CEB" author="Hincks, W. D. & Dibb, J. R." box="[1139,1431,1700,1725]" pageId="23" pageNumber="474" refId="ref18872" refString="Hincks, W. D. & Dibb, J. R. (1935) Coleopterorum Catalogus, Pars 142: Passalidae. W. Junk, The Hague, Netherlands, 118 pp." type="book" year="1935">Hincks & Dibb 1935: 21</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FECC00FF81DA9CB6" author="Reyes-Castillo, P." box="[151,435,1735,1760]" pageId="23" pageNumber="474" pagination="1 - 240" refId="ref19215" refString="Reyes-Castillo, P. (1970) Coleoptera, Passalidae: morfologia y division en grandes grupos; generos americanos. Folia Entomologica Mexicana, 20 - 22, 1 - 240." type="journal article" year="1970">Reyes-Castillo 1970: 177</bibRefCitation>
|
||
) apparently were based on
|
||
<specimenCount id="39B84BCFFC47FF98FCB800F083159CB6" box="[739,892,1735,1760]" pageId="23" pageNumber="474" type="generic">one specimen</specimenCount>
|
||
at the MNHN, identified by Kuwert (
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FB4500FF81DB9D53" author="Schuster, J. C. & Reyes-Castillo, P." pageId="23" pageNumber="474" pagination="1 - 49" refId="ref19543" refString="Schuster, J. C. & Reyes-Castillo, P. (1990) Coleoptera, Passalidae: Ogyges Kaup, revision de un genero mesoamericano de montana. Acta Zoologica Mexicana (n. s.), 40, 1 - 49." type="journal article" year="1990">Schuster & Reyes-Castillo 1990: 20</bibRefCitation>
|
||
attribute the handwritten label to Kaup), with the locality data (in the same label) indicating Guatem.[ala]/
|
||
<collectingCountry id="57A9C0D6FC47FF98FFFC013782489D7E" box="[423,545,1807,1832]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
|
||
. The specimen of
|
||
<taxonomicName id="E8BEFBC5FC47FF98FCA8012883E49D7E" baseAuthorityName="Bates" baseAuthorityYear="1886" box="[755,909,1808,1832]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="1DCA5C54FC47FF98FCA8012883E49D7E" box="[755,909,1808,1832]" italics="true" pageId="23" pageNumber="474">O. championi</emphasis>
|
||
</taxonomicName>
|
||
from the MNHN, with the label “
|
||
<collectingCountry id="57A9C0D6FC47FF98FB48013785E49D7E" box="[1299,1421,1807,1832]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
|
||
”, cited by
|
||
<bibRefCitation id="4B2FFDB7FC47FF98FEA3010C821F9D1B" author="Schuster, J. C. & Reyes-Castillo, P." box="[248,630,1844,1869]" pageId="23" pageNumber="474" pagination="1 - 49" refId="ref19543" refString="Schuster, J. C. & Reyes-Castillo, P. (1990) Coleoptera, Passalidae: Ogyges Kaup, revision de un genero mesoamericano de montana. Acta Zoologica Mexicana (n. s.), 40, 1 - 49." type="journal article" year="1990">Schuster & Reyes-Castillo (1990)</bibRefCitation>
|
||
, by the barely marked lateral punctures on elytra, apparently belongs to the population of Alta and
|
||
<collectingRegion id="ED7A4EA4FC47FF98FF9F016F82319D39" box="[452,600,1879,1904]" country="Guatemala" name="Baja Verapaz" pageId="23" pageNumber="474">Baja Verapaz</collectingRegion>
|
||
in
|
||
<collectingCountry id="57A9C0D6FC47FF98FC27016F82909D26" box="[636,761,1879,1904]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |