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<document id="2FE1EE86615F7689E2580C976FC269FF" ID-DOI="10.11646/zootaxa.3889.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4952403" ID-ZooBank="1F3F076D-A4F1-4B4A-977D-4B99581C6779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730508465" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2014" docId="A7173150FC42FF9BFECC06AF828A989B" docLanguage="en" docName="zt03889p484.pdf" docOrigin="Zootaxa 3889 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ogyges mutenroshii Cano 2014, new species" docType="treatment" docVersion="4" lastPageNumber="471" masterDocId="5B2E4928FC50FF8FFE5B063880699A56" masterDocTitle="Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly" masterLastPageNumber="484" masterPageNumber="451" pageNumber="469" updateTime="1698967030688" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5A89CAA923760FB04A635993562B87EE">Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly</mods:title>
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<mods:namePart id="9C355F92B92F00A66B23C8F8E14A04BB">Cano, Enio B.</mods:namePart>
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<mods:date id="45499DB04E3785043A411A31F9ED8ACF">2014</mods:date>
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<mods:number id="F8592722DC6EB2E1B5EC7B00464DB24C">2014-12-03</mods:number>
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<mods:number id="B5BB0A1F8143D59BF8345E2DADCD654F">3889</mods:number>
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<mods:identifier id="6FB2A0AC48B7A391B937D14281F3CE42" type="DOI">10.11646/zootaxa.3889.4.1</mods:identifier>
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<treatment id="A7173150FC42FF9BFECC06AF828A989B" ID-DOI="http://doi.org/10.5281/zenodo.4953259" ID-Zenodo-Dep="4953259" LSID="urn:lsid:plazi:treatment:A7173150FC42FF9BFECC06AF828A989B" httpUri="http://treatment.plazi.org/id/A7173150FC42FF9BFECC06AF828A989B" lastPageId="20" lastPageNumber="471" pageId="18" pageNumber="469">
<subSubSection id="67A4D3CDFC42FF9DFECC06AF82189AE4" box="[151,625,151,178]" pageId="18" pageNumber="469" type="nomenclature">
<paragraph id="2F018046FC42FF9DFECC06AF82189AE4" blockId="18.[151,625,151,211]" box="[151,625,151,178]" pageId="18" pageNumber="469">
<heading id="7449372AFC42FF9DFECC06AF82189AE4" bold="true" box="[151,625,151,178]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="1DCA5C54FC42FF9DFECC06AF82189AE4" bold="true" box="[151,625,151,178]" pageId="18" pageNumber="469">
<taxonomicName id="E8BEFBC5FC42FF9DFECC06AF81BD9AE4" authority="Cano" authorityName="Cano" authorityYear="2014" box="[151,468,151,178]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="mutenroshii" status="sp. nov.">
<emphasis id="1DCA5C54FC42FF9DFECC06AF81E19AE7" bold="true" box="[151,392,151,177]" italics="true" pageId="18" pageNumber="469">Ogyges mutenroshii</emphasis>
Cano
</taxonomicName>
,
<taxonomicNameLabel id="06F9E12FFC42FF9DFFBA06A082189AE4" box="[481,625,152,178]" pageId="18" pageNumber="469" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="67A4D3CDFC42FF9DFECC068281279A85" box="[151,334,186,211]" pageId="18" pageNumber="469" type="description">
<paragraph id="2F018046FC42FF9DFECC068281279A85" blockId="18.[151,625,151,211]" box="[151,334,186,211]" pageId="18" pageNumber="469">
<figureCitation id="B7859CC3FC42FF9DFECC068281779A85" box="[151,286,186,211]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="18.[151,250,1713,1735]" captionStartId-1="19.[151,250,1967,1989]" captionTargetBox-0="[156,1436,407,1691]" captionTargetBox-1="[158,1429,1135,1940]" captionTargetId-0="figure-206@18.[151,1436,407,1691]" captionTargetId-1="figure-420@19.[151,1436,1129,1946]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURE 13. Holotype of Ogyges mutenroshii: a. lateral view. b. dorsal view. c. ventral view. d. detail of mentum. e. anterolateral view of head." captionText-1="FIGURE 14. Dorsal aspect of head of Ogyges mutenroshii." figureDoi-0="http://doi.org/10.5281/zenodo.4952437" figureDoi-1="http://doi.org/10.5281/zenodo.4952439" httpUri-0="https://zenodo.org/record/4952437/files/figure.png" httpUri-1="https://zenodo.org/record/4952439/files/figure.png" pageId="18" pageNumber="469">Figs. 1314</figureCitation>
,
<figureCitation id="B7859CC3FC42FF9DFF72068281279A85" box="[297,334,186,211]" captionStart="FIGURE 21" captionStartId="28.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure-318@28.[151,1436,193,1354]" captionTargetPageId="28" captionText="FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili." figureDoi="http://doi.org/10.5281/zenodo.4952455" httpUri="https://zenodo.org/record/4952455/files/figure.png" pageId="18" pageNumber="469">21r</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC42FF9DFECC073A85129B35" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="2F018046FC42FF9DFECC073A85129B35" blockId="18.[151,1436,258,355]" pageId="18" pageNumber="469">
<emphasis id="1DCA5C54FC42FF9DFECC073A81789B4D" bold="true" box="[151,273,258,283]" inLineHeading="true" pageId="18" pageNumber="469" reason="1">Diagnosis.</emphasis>
<taxonomicName id="E8BEFBC5FC42FF9DFF43073B81919B4D" authorityName="Cano" authorityYear="2014" box="[280,504,259,283]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="mutenroshii">
<emphasis id="1DCA5C54FC42FF9DFF43073B81919B4D" box="[280,504,259,283]" italics="true" pageId="18" pageNumber="469">Ogyges mutenroshii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="E8BEFBC5FC42FF9DFC9C073B83569B4D" authorityName="Schuster, Cano, &amp; Boucher" authorityYear="2005" box="[711,831,259,283]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="nahuali">
<emphasis id="1DCA5C54FC42FF9DFC9C073B83569B4D" box="[711,831,259,283]" italics="true" pageId="18" pageNumber="469">O. nahuali</emphasis>
</taxonomicName>
,
<taxonomicName id="E8BEFBC5FC42FF9DFD16073B83C29B4C" authorityName="Cano" authorityYear="2014" box="[845,939,259,282]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="cavei">
<emphasis id="1DCA5C54FC42FF9DFD16073B83C29B4C" box="[845,939,259,282]" italics="true" pageId="18" pageNumber="469">O. cavei</emphasis>
</taxonomicName>
, and
<taxonomicName id="E8BEFBC5FC42FF9DFDB3073B841E9B4C" authority="Cano, 2014" authorityName="Cano" authorityYear="2014" box="[1000,1143,259,282]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="toriyamai" status="sp. nov.">
<emphasis id="1DCA5C54FC42FF9DFDB3073B841E9B4C" box="[1000,1143,259,282]" italics="true" pageId="18" pageNumber="469">O. toriyamai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F9E12FFC42FF9DFA25073B85619B4D" box="[1150,1288,258,283]" pageId="18" pageNumber="469" rank="species">new species</taxonomicNameLabel>
, by the small dorsal punctures on elytra and strong punctures on metasternum. Is easily distinguished by the brilliant elytra, the very wide protibia, the form of central horn, the absence of postorbital pits, and the greater reduction in the eye.
</paragraph>
</subSubSection>
<caption id="7BC1D0CEFC42FF9DFECC008981CF9CB3" ID-DOI="http://doi.org/10.5281/zenodo.4952437" ID-Zenodo-Dep="4952437" httpUri="https://zenodo.org/record/4952437/files/figure.png" pageId="18" pageNumber="469" startId="18.[151,250,1713,1735]" targetBox="[156,1436,407,1691]" targetPageId="18">
<paragraph id="2F018046FC42FF9DFECC008981CF9CB3" blockId="18.[151,1436,1713,1765]" pageId="18" pageNumber="469">
<emphasis id="1DCA5C54FC42FF9DFECC0089814D9C90" bold="true" box="[151,292,1713,1735]" pageId="18" pageNumber="469">FIGURE 13.</emphasis>
Holotype of
<taxonomicName id="E8BEFBC5FC42FF9DFFE0008982E29C91" authorityName="Cano" authorityYear="2014" box="[443,651,1713,1735]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="mutenroshii">
<emphasis id="1DCA5C54FC42FF9DFFE0008982E29C91" box="[443,651,1713,1735]" italics="true" pageId="18" pageNumber="469">Ogyges mutenroshii</emphasis>
</taxonomicName>
: a. lateral view. b. dorsal view. c. ventral view. d. detail of mentum. e. anterolateral view of head.
</paragraph>
</caption>
<subSubSection id="67A4D3CDFC42FF9BFE9C013580A29B33" lastPageId="20" lastPageNumber="471" pageId="18" pageNumber="469" type="description">
<paragraph id="2F018046FC42FF9CFE9C013585FC98FE" blockId="18.[151,1437,1805,2013]" lastBlockId="19.[151,1437,151,1077]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<emphasis id="1DCA5C54FC42FF9DFE9C013581309D70" bold="true" box="[199,345,1805,1830]" inLineHeading="true" pageId="18" pageNumber="469" reason="1">Description.</emphasis>
<typeStatus id="F0053EE4FC42FF9DFF39013581A49D70" box="[354,461,1805,1830]" pageId="18" pageNumber="469" type="holotype">Holotype</typeStatus>
, sex unknown, black adult. Total length 34.56, elytral length 18.44, pronotal length 9.24, pronotal width 12.05, humeral width 10.95. Head: anterior border of labrum slightly concave. Clypeus inclined (45
<superScript id="D8CB2D0EFC42FF9DFF45016C814C9D34" attach="none" box="[286,293,1876,1890]" fontSize="6" pageId="18" pageNumber="469">o</superScript>
); anterior border straight and thin, with very small, central invagination, without suture separating it from the mediofrontal area (except an indication of slightly rugose area that runs between mediofrontal tubercles). Mediofrontal tubercles rounded and well defined. Mediofrontal area smooth, without internal tubercles and posterofrontal ridges. Laterofrontal areas roughened. Lateropostfrontal areas wide, glabrous and roughened at sides. Central horn long and very narrow, apical 1/3 declivous in lateral view, dorsally excavated on both sides up to the level of the lateroposterior tubercles, with apex free and slightly directed upward, without median longitudinal groove posteriorly; lateroposterior tubercles somewhat elongate, separated from the posterior keel of the central horn. Postfrontal groove laterally deep. Supraorbital ridge with equal anterior tubercles; posterior 1/2 bifurcate with the supraorbital fossae very deep; external ridge well marked. Ocular canthus with apex swollen, dorsally covering more than 2/3 of eye. Eyes reduced. Eye width =
<quantity id="E8462DA3FC43FF9CFDCC077483969B35" box="[919,1023,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="19" pageNumber="470" unit="mm" value="0.38">0.38 mm</quantity>
. Interocular width =
<quantity id="E8462DA3FC43FF9CFAB60774853C9B35" box="[1261,1365,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.92" pageId="19" pageNumber="470" unit="mm" value="6.92">6.92 mm</quantity>
. Head width =
<quantity id="E8462DA3FC43FF9CFEAF075781359BD1" box="[244,348,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="8.46" pageId="19" pageNumber="470" unit="mm" value="8.46">8.46 mm</quantity>
; ratio both eye widths/head width = 0.09. Postorbital pits, immediately behind the external ridge, punctate setose. Ligula almost flat basally, with eroded (also in the
<typeStatus id="F0053EE4FC43FF9CFD8907AC845D9BFB" box="[978,1076,404,429]" pageId="19" pageNumber="470" type="paratype">paratype</typeStatus>
) central tooth, with setaceous punctures medially; anterior ventral transversal carina clearly present. Lateral lobes of mentum with abundant setose punctures; lateral basal scars oval, opaque (shagreened), and punctate setose; medial basal mentum bare and shiny, with one-three punctures on each side. Hypostomal process elongate, without lateral depression, wide medially and narrow in apical third. Infraocular ridge present, short and smooth, surrounded by striate, setose punctures. Mandible with dorsal tooth occuppying 1/2 of the length; internal face of mandible in dorsal view, smooth. Antepenultimate antennomere of antennal club slightly longer (or subequal) than penultimate, less wide than the penultimate; antennal lamellae slightly concave dorsally (
<figureCitation id="B7859CC3FC43FF9CFD2604B7846F98FE" box="[893,1030,655,680]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="18.[151,250,1713,1735]" captionStartId-1="19.[151,250,1967,1989]" captionTargetBox-0="[156,1436,407,1691]" captionTargetBox-1="[158,1429,1135,1940]" captionTargetId-0="figure-206@18.[151,1436,407,1691]" captionTargetId-1="figure-420@19.[151,1436,1129,1946]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURE 13. Holotype of Ogyges mutenroshii: a. lateral view. b. dorsal view. c. ventral view. d. detail of mentum. e. anterolateral view of head." captionText-1="FIGURE 14. Dorsal aspect of head of Ogyges mutenroshii." figureDoi-0="http://doi.org/10.5281/zenodo.4952437" figureDoi-1="http://doi.org/10.5281/zenodo.4952439" httpUri-0="https://zenodo.org/record/4952437/files/figure.png" httpUri-1="https://zenodo.org/record/4952439/files/figure.png" pageId="19" pageNumber="470">Figs. 1314</figureCitation>
),
<quantity id="E8462DA3FC43FF9CFA4104B7841798F1" box="[1050,1150,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="19" pageNumber="470" unit="mm" value="1.69">1.69 mm</quantity>
wide and
<quantity id="E8462DA3FC43FF9CFAAA04B7853C98F1" box="[1265,1365,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="19" pageNumber="470" unit="mm" value="1.92">1.92 mm</quantity>
long.
</paragraph>
<paragraph id="2F018046FC43FF9CFE9C048C8114999E" blockId="19.[151,1437,151,1077]" pageId="19" pageNumber="470">Thorax: Lateral fossae of pronotum only with sparse and very shallow micropunctures, visible at great magnification. Pronotum with marginal groove narrow and smooth, laterally with scattered micropunctures of rugose aspect (striate), visible at great magnification, more abundant in front of lateral fossae; anterior angles rounded; disc smooth. Prosternellum brilliant at center, shagreened at anterior 2/5 and posterior (except on sides) 1/ 5. Mesosternum glabrous; lateral depressions elongate and rugose (shagreened). Mesepisternum with oval, elongate, shagreened area. Metasternum anterior angles glabrous; disc delimited by 38 and 40 well-marked punctures (some striate) on each side; marginal groove glabrous, narrow and sligthly striate, posteriorly two times wider than medially.</paragraph>
<paragraph id="2F018046FC43FF9CFE9C05EC83519E63" blockId="19.[151,1437,151,1077]" pageId="19" pageNumber="470">Elytra: Brilliant; striations well marked and with weakly defined punctures between striae 13, gradually wider between striae 410 and apical 1/3; junctions of striations 1 and 10 with some extra punctures. Vertical anterior border of elytra with sparse, minute setae on interstriae 26.</paragraph>
<caption id="7BC1D0CEFC43FF9CFECC0197829B9D93" ID-DOI="http://doi.org/10.5281/zenodo.4952439" ID-Zenodo-Dep="4952439" box="[151,754,1967,1989]" httpUri="https://zenodo.org/record/4952439/files/figure.png" pageId="19" pageNumber="470" startId="19.[151,250,1967,1989]" targetBox="[158,1429,1135,1940]" targetPageId="19">
<paragraph id="2F018046FC43FF9CFECC0197829B9D93" blockId="19.[151,754,1967,1989]" box="[151,754,1967,1989]" pageId="19" pageNumber="470">
<emphasis id="1DCA5C54FC43FF9CFECC019781769D92" bold="true" box="[151,287,1967,1989]" pageId="19" pageNumber="470">FIGURE 14.</emphasis>
Dorsal aspect of head of
<taxonomicName id="E8BEFBC5FC43FF9CFC79019782859D93" authorityName="Cano" authorityYear="2014" box="[546,748,1967,1989]" class="Insecta" family="Passallidae" genus="Ogyges" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="mutenroshii">
<emphasis id="1DCA5C54FC43FF9CFC79019782859D93" box="[546,748,1967,1989]" italics="true" pageId="19" pageNumber="470">Ogyges mutenroshii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="2F018046FC44FF9BFE9C06A081609A83" blockId="20.[151,1436,151,717]" pageId="20" pageNumber="471">Legs: Protibia very wide. Profemur with anterioventral groove marked; metafemur elongate; mesotibia with one spine.</paragraph>
<paragraph id="2F018046FC44FF9BFE9C06E7842B9AAE" blockId="20.[151,1436,151,717]" box="[199,1090,223,248]" pageId="20" pageNumber="471">Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.</paragraph>
<paragraph id="2F018046FC44FF9BFE9C073C80A29B33" blockId="20.[151,1436,151,717]" pageId="20" pageNumber="471">
Variation in
<typeStatus id="F0053EE4FC44FF9BFF02073C81D79B4A" box="[345,446,260,284]" pageId="20" pageNumber="471" type="paratype">paratype</typeStatus>
: Total length 33.19, elytral length 17.74, pronotal length 8.83, pronotal width 11.52, humeral width 10.18. Disc of metasternum delimited by 31 and 45 well-marked punctures (some striate) on each side.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC44FF9BFE9C075782D8986B" pageId="20" pageNumber="471" type="materials_examined">
<paragraph id="2F018046FC44FF9BFE9C075782219BDE" blockId="20.[151,1436,151,717]" box="[199,584,367,392]" pageId="20" pageNumber="471">
<emphasis id="1DCA5C54FC44FF9BFE9C075781D99BD1" box="[199,432,367,392]" inLineHeading="true" pageId="20" pageNumber="471" reason="1">
<emphasis id="1DCA5C54FC44FF9BFE9C075781C09BDE" bold="true" box="[199,425,367,392]" pageId="20" pageNumber="471">Material examined</emphasis>
.
</emphasis>
<specimenCount id="39B84BCFFC44FF9BFFEC0757822D9BDE" box="[439,580,367,392]" pageId="20" pageNumber="471" type="generic">2 specimens</specimenCount>
.
</paragraph>
<paragraph id="2F018046FC44FF9BFE9C07AC81CB984E" blockId="20.[151,1436,151,717]" pageId="20" pageNumber="471">
<emphasis id="1DCA5C54FC44FF9BFE9C07AC811C9BFD" box="[199,373,404,429]" inLineHeading="true" pageId="20" pageNumber="471" reason="1">
<emphasis id="1DCA5C54FC44FF9BFE9C07AC81079BFB" bold="true" box="[199,366,404,429]" pageId="20" pageNumber="471">Type material</emphasis>
.
</emphasis>
<materialsCitation id="9FD68A1BFC44FF9BFF2607AC83B09B86" collectingDate="2007-08-07" collectingDateMax="2007-08-11" collectingDateMin="2007-08-07" collectorName="Coll. S. Beynon." country="Honduras" elevation="1656" location="Parque Nacional Cusuco" pageId="20" pageNumber="471" specimenCount="1" stateProvince="Cortes" typeStatus="holotype">
<typeStatus id="F0053EE4FC44FF9BFF2607AC81839BFB" box="[381,490,404,429]" pageId="20" pageNumber="471" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="57A9C0D6FC44FF9BFFAF07AC82FD9BFB" box="[500,660,404,429]" name="Honduras" pageId="20" pageNumber="471">HONDURAS</collectingCountry>
:
<collectingRegion id="ED7A4EA4FC44FF9BFCC407AC82829BFB" box="[671,747,404,429]" country="Honduras" name="Cortes" pageId="20" pageNumber="471">Cortés</collectingRegion>
,
<location id="2A61D69DFC44FF9BFCAC07AC84799BFB" LSID="urn:lsid:plazi:treatment:A7173150FC42FF9BFECC06AF828A989B:2A61D69DFC44FF9BFCAC07AC84799BFB" box="[759,1040,404,429]" country="Honduras" name="Parque Nacional Cusuco" pageId="20" pageNumber="471" stateProvince="Cortes">Parque Nacional Cusuco</location>
, 2nd. broadleaved forest. Nr. UTM 16P 369210 1713408. Ca.
<quantity id="E8462DA3FC44FF9BFF9F078F82739B99" box="[452,538,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.656" pageId="20" pageNumber="471" unit="m" value="1656.0">
<elevation id="A4936775FC44FF9BFF9F078F82739B99" box="[452,538,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.656" pageId="20" pageNumber="471" unit="m" value="1656.0">1656 m</elevation>
</quantity>
alt.,
<date id="5B00A686FC44FF9BFC09078F837A9B86" box="[594,787,439,464]" pageId="20" pageNumber="471" value="2007-08-07" valueMax="2007-08-11" valueMin="2007-08-07">
<collectingDate id="4B445F6EFC44FF9BFC09078F837A9B86" box="[594,787,439,464]" pageId="20" pageNumber="471" value="2007-08-07" valueMax="2007-08-11" valueMin="2007-08-07">0711 VIII 2007</collectingDate>
</date>
.
<collectorName id="824BE590FC44FF9BFD46078F83B09B86" box="[797,985,439,464]" pageId="20" pageNumber="471">Coll. S. Beynon.</collectorName>
</materialsCitation>
<materialsCitation id="9FD68A1BFC44FF9BFDBB078081CB984E" collectingDate="2006-07-16" collectingDateMax="2006-07-20" collectingDateMin="2006-07-16" collectorName="Coll. E. Marabuto." country="Honduras" elevation="1550" location="Parque Nacional Cusuco" pageId="20" pageNumber="471" specimenCount="1" stateProvince="Cortes" typeStatus="paratype">
<typeStatus id="F0053EE4FC44FF9BFDBB0780842F9B86" box="[992,1094,440,464]" pageId="20" pageNumber="471" type="paratype">Paratype</typeStatus>
: One specimen, unknown sex, same data as holotype except: UTM 16P 369795 1714017. Alt. aprox.
<quantity id="E8462DA3FC44FF9BFDF207E484699BA5" box="[937,1024,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="471" unit="m" value="1550.0">
<elevation id="A4936775FC44FF9BFDF207E484699BA5" box="[937,1024,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="20" pageNumber="471" unit="m" value="1550.0">1550 m</elevation>
</quantity>
.
<date id="5B00A686FC44FF9BFA5007E484A99BA3" box="[1035,1216,476,501]" pageId="20" pageNumber="471" value="2006-07-16" valueMax="2006-07-20" valueMin="2006-07-16">
<collectingDate id="4B445F6EFC44FF9BFA5007E484A99BA3" box="[1035,1216,476,501]" pageId="20" pageNumber="471" value="2006-07-16" valueMax="2006-07-20" valueMin="2006-07-16">1620 VII 2006</collectingDate>
</date>
.
<collectingMethod id="F6FFF851FC44FF9BFA9107E480AF9841" pageId="20" pageNumber="471">Dung baited pitfall trap</collectingMethod>
.
<collectorName id="824BE590FC44FF9BFE8B07C781CB984E" box="[208,418,511,536]" pageId="20" pageNumber="471">Coll. E. Marabuto.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="2F018046FC44FF9BFE9C041C82D8986B" blockId="20.[151,1436,151,717]" box="[199,689,548,573]" pageId="20" pageNumber="471">
<materialsCitation id="9FD68A1BFC44FF9BFE9C041C82C4986B" box="[199,685,548,573]" collectionCode="UVGC" pageId="20" pageNumber="471" specimenCount="2" typeStatus="holotype">
<typeStatus id="F0053EE4FC44FF9BFE9C041C8146986B" box="[199,303,548,573]" pageId="20" pageNumber="471" type="holotype">Holotype</typeStatus>
and
<typeStatus id="F0053EE4FC44FF9BFF3E041C81AC986A" box="[357,453,548,572]" pageId="20" pageNumber="471" type="paratype">paratype</typeStatus>
deposited at
<collectionCode id="49AF1883FC44FF9BFC01041C82C4986B" box="[602,685,548,573]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="20" pageNumber="471">UVGC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC44FF9BFE9C047F827D98D3" pageId="20" pageNumber="471" type="etymology">
<paragraph id="2F018046FC44FF9BFE9C047F827D98D3" blockId="20.[151,1436,151,717]" pageId="20" pageNumber="471">
<emphasis id="1DCA5C54FC44FF9BFE9C047F81249809" box="[199,333,583,608]" inLineHeading="true" pageId="20" pageNumber="471" reason="1">
<emphasis id="1DCA5C54FC44FF9BFE9C047F812F9836" bold="true" box="[199,326,583,608]" pageId="20" pageNumber="471">Etymology</emphasis>
.
</emphasis>
Named after Muten Roshi (Master Roshi), a principal character of the Akira Toriyama´s series of manga and anime, “Dragon Ball”.
</paragraph>
</subSubSection>
<subSubSection id="67A4D3CDFC44FF9BFE9C04B7828A989B" pageId="20" pageNumber="471" type="distribution">
<paragraph id="2F018046FC44FF9BFE9C04B7828A989B" blockId="20.[151,1436,151,717]" pageId="20" pageNumber="471">
<emphasis id="1DCA5C54FC44FF9BFE9C04B7810998FE" bold="true" box="[199,352,655,680]" inLineHeading="true" pageId="20" pageNumber="471" reason="1">Distribution.</emphasis>
This species is known from the middle altitude cloud forest of the Cerro Azul Meambar National Park, in
<collectingRegion id="ED7A4EA4FC44FF9BFEAE048C8260989D" box="[245,521,692,717]" country="Honduras" name="Comayagua" pageId="20" pageNumber="471">Comayagua Department</collectingRegion>
,
<collectingCountry id="57A9C0D6FC44FF9BFC4F048C82E8989B" box="[532,641,692,717]" name="Honduras" pageId="20" pageNumber="471">Honduras</collectingCountry>
(
<figureCitation id="B7859CC3FC44FF9BFCCB048C82BC989B" box="[656,725,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1904,1926]" captionTargetBox="[171,1398,927,1870]" captionTargetId="figure-358@6.[149,1434,899,1893]" captionTargetPageId="6" captionText="FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi." figureDoi="http://doi.org/10.5281/zenodo.4952413" httpUri="https://zenodo.org/record/4952413/files/figure.png" pageId="20" pageNumber="471">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>