479 lines
39 KiB
XML
479 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.65.7403" ID-PMC="PMC4957021" ID-Pensoft-Pub="1314-2003-65-35" ID-Pensoft-UUID="6224170CFFB3643C4C7AFF9EFFD8FFBF" ID-PubMed="27489487" ID-Zenodo-Dep="122985" ModsDocID="1314-2003-65-35" checkinTime="1466061625811" checkinUser="pensoft" docAuthor="Yahara, Tetsukazu, Tagane, Shuichiro, Mase, Keiko, Chhang, Phourin & Toyama, Hironori" docDate="2016" docId="99EAAAEDA731EC1D3F4471F17FF4CADF" docLanguage="en" docName="PhytoKeys 65: 35-46" docOrigin="PhytoKeys 65" docPubDate="2016-06-15" docSource="http://dx.doi.org/10.3897/phytokeys.65.7403" docTitle="Machilus bokorensis Yahara & Tagane 2016, sp. nov." docType="treatment" docVersion="7" id="6224170CFFB3643C4C7AFF9EFFD8FFBF" lastPageNumber="37" masterDocId="6224170CFFB3643C4C7AFF9EFFD8FFBF" masterDocTitle="Flora of Bokor National Park V: Two new species of Machilus (Lauraceae), M. bokorensis and M. brevipaniculata" masterLastPageNumber="46" masterPageNumber="35" pageNumber="37" updateTime="1668141723299" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Flora of Bokor National Park V: Two new species of Machilus (Lauraceae), M. bokorensis and M. brevipaniculata</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yahara, Tetsukazu</mods:namePart>
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<mods:affiliation>Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tagane, Shuichiro</mods:namePart>
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<mods:affiliation>Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
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<mods:nameIdentifier type="email">stagane29@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mase, Keiko</mods:namePart>
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<mods:affiliation>Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chhang, Phourin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Toyama, Hironori</mods:namePart>
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<mods:affiliation>Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-06-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>65</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.65.7403</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.65.7403</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-65-35</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">6224170CFFB3643C4C7AFF9EFFD8FFBF</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">122985</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-GBIF-Taxon="127901575" LSID="urn:lsid:plazi:treatment:99EAAAEDA731EC1D3F4471F17FF4CADF" httpUri="http://treatment.plazi.org/id/99EAAAEDA731EC1D3F4471F17FF4CADF" lastPageId="3" lastPageNumber="37" pageId="2" pageNumber="37">
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<subSubSection pageId="2" pageNumber="37" type="nomenclature">
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<paragraph pageId="2" pageNumber="37">
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<taxonomicName LSID="99EAAAED-A731-EC1D-3F44-71F17FF4CADF" authority="Yahara & Tagane" authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="bokorensis" status="sp. nov.">Machilus bokorensis Yahara & Tagane</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="37">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Machilus bokorensis Yahara & Tagane. a flowering branch, b abaxial lower surface of lamina c flowers d buds e holotype: Tagane et al. 5475 (KYO), scale bar = 10 cm f inflorescence, scale bar = 1 cm. Photographs: a-c by S. Tagane, d by K. Fuse." figureDoi="10.3897/phytokeys.65.7403.figure1" httpUri="https://binary.pensoft.net/fig/92599" pageId="2" pageNumber="37">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Machilus bokorensis Yahara & Tagane. a flowering branch b top of branch c flowers d stamen in 1 st whorl e stamen in 2 nd whorl with staminode f stamen in 3 rd whorl with glands g pistil h outer perianth lobes i inner perianth lobe j, k bud scales. Materials from Tagane et al. 5475 (KYO). Drawn by S. Tagane." figureDoi="10.3897/phytokeys.65.7403.figure2" httpUri="https://binary.pensoft.net/fig/92600" pageId="2" pageNumber="37">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="2" pageNumber="37" type="reference_group">
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<paragraph pageId="2" pageNumber="37">
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<taxonomicName authority="Yahara & Tagane & Mase & Chhang & Toyama, 2016" authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima" status="sp. nov.">Machilus odoratissima</taxonomicName>
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(non Nees) sensu H.Liou, Laurac. Chine & Indochine: 53 (1932), for the specimen collected from Nord de Kampot (P, no. 14625).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="37" type="diagnosis">
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<paragraph pageId="2" pageNumber="37">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="37">
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Similar to
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<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus odoratissima</emphasis>
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</taxonomicName>
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Nees in elliptic-lanceolate or oblanceolate, glabrous leaves and subterminal inflorescences, but differing in having sericeous inflorescence rachis (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Machilus bokorensis Yahara & Tagane. a flowering branch b top of branch c flowers d stamen in 1 st whorl e stamen in 2 nd whorl with staminode f stamen in 3 rd whorl with glands g pistil h outer perianth lobes i inner perianth lobe j, k bud scales. Materials from Tagane et al. 5475 (KYO). Drawn by S. Tagane." figureDoi="10.3897/phytokeys.65.7403.figure2" httpUri="https://binary.pensoft.net/fig/92600" pageId="2" pageNumber="37">2c</figureCitation>
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; vs. glabrous in
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<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus odoratissima</emphasis>
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</taxonomicName>
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), perianth lobes pubescent on both surfaces (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Machilus bokorensis Yahara & Tagane. a flowering branch b top of branch c flowers d stamen in 1 st whorl e stamen in 2 nd whorl with staminode f stamen in 3 rd whorl with glands g pistil h outer perianth lobes i inner perianth lobe j, k bud scales. Materials from Tagane et al. 5475 (KYO). Drawn by S. Tagane." figureDoi="10.3897/phytokeys.65.7403.figure2" httpUri="https://binary.pensoft.net/fig/92600" pageId="2" pageNumber="37">2c</figureCitation>
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; vs. outside almost glabrous), ca. 1 mm long staminodes (vs. ca. 2 mm long), and twigs drying blackish grey (vs. reddish). Similar to
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<taxonomicName authorityName="Meisn" authorityYear="1864" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus parviflora" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="parviflora">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus parviflora</emphasis>
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</taxonomicName>
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Meisn. among Indochinese species in having large terminal buds, but differing in bud scales densely puberulent outside (vs. densely golden hairy outside in
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<taxonomicName authorityName="Meisn" authorityYear="1864" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus parviflora" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="parviflora">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus parviflora</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<caption ID-Zenodo-Dep="976449" doi="10.3897/phytokeys.65.7403.figure1" httpUri="https://binary.pensoft.net/fig/92599" pageId="2" pageNumber="37" start="Figure 1" startId="F1">
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<paragraph pageId="2" pageNumber="37">
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<emphasis bold="true" pageId="2" pageNumber="37">Figure 1.</emphasis>
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<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="bokorensis">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus bokorensis</emphasis>
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</taxonomicName>
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Yahara & Tagane.
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<emphasis bold="true" pageId="2" pageNumber="37">a</emphasis>
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flowering branch,
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<emphasis bold="true" pageId="2" pageNumber="37">b</emphasis>
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abaxial lower surface of lamina
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<emphasis bold="true" pageId="2" pageNumber="37">c</emphasis>
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flowers
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<emphasis bold="true" pageId="2" pageNumber="37">d</emphasis>
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buds
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<emphasis bold="true" pageId="2" pageNumber="37">e</emphasis>
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holotype:
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<emphasis italics="true" pageId="2" pageNumber="37">Tagane et al. 5475</emphasis>
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(KYO), scale bar = 10 cm
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<emphasis bold="true" pageId="2" pageNumber="37">f</emphasis>
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inflorescence, scale bar = 1 cm. Photographs:
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<emphasis bold="true" pageId="2" pageNumber="37">a-c</emphasis>
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by S. Tagane,
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<emphasis bold="true" pageId="2" pageNumber="37">d</emphasis>
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by K. Fuse.
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="976451" doi="10.3897/phytokeys.65.7403.figure2" httpUri="https://binary.pensoft.net/fig/92600" pageId="2" pageNumber="37" start="Figure 2" startId="F2">
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<paragraph pageId="2" pageNumber="37">
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<emphasis bold="true" pageId="2" pageNumber="37">Figure 2.</emphasis>
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<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="bokorensis">
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<emphasis italics="true" pageId="2" pageNumber="37">Machilus bokorensis</emphasis>
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</taxonomicName>
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Yahara & Tagane.
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<emphasis bold="true" pageId="2" pageNumber="37">a</emphasis>
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flowering branch
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<emphasis bold="true" pageId="2" pageNumber="37">b</emphasis>
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top of branch
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<emphasis bold="true" pageId="2" pageNumber="37">c</emphasis>
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flowers
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<emphasis bold="true" pageId="2" pageNumber="37">d</emphasis>
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stamen in 1st whorl
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<emphasis bold="true" pageId="2" pageNumber="37">e</emphasis>
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stamen in 2nd whorl with staminode
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<emphasis bold="true" pageId="2" pageNumber="37">f</emphasis>
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stamen in 3rd whorl with glands
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<emphasis bold="true" pageId="2" pageNumber="37">g</emphasis>
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pistil
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<emphasis bold="true" pageId="2" pageNumber="37">h</emphasis>
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outer perianth lobes
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<emphasis bold="true" pageId="2" pageNumber="37">i</emphasis>
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inner perianth lobe
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<emphasis bold="true" pageId="2" pageNumber="37">j, k</emphasis>
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bud scales. Materials from
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<emphasis italics="true" pageId="2" pageNumber="37">Tagane et al. 5475</emphasis>
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(KYO). Drawn by S. Tagane.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="37" type="materials_examined">
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<paragraph pageId="2" pageNumber="37">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="37">
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<materialsCitation collectingDate="2013-02-16" collectorName="Tagane S., Toyama H., Wachi N., Ichihasi R., Mase K., Zhu M., Chhnang P." country="CAMBODIA" elevation="933" latitude="10.6554165" location="Mt. Bokor" longLatPrecision="1" longitude="104.06017" municipality="Bokor National Park" specimenCount="1" stateProvince="Kampot Province" typeStatus="holotype">
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<collectingCountry name="Cambodia">CAMBODIA</collectingCountry>
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.
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<collectingRegion country="Cambodia" name="Kampot">Kampot Province</collectingRegion>
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,
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<collectingMunicipality>Bokor National Park</collectingMunicipality>
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, dense evergreen forest on the plateau of
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<location LSID="urn:lsid:plazi:treatment:99EAAAEDA731EC1D3F4471F17FF4CADF:4AD0B50D96BAA15BA922D3ACF8E3BE79" country="CAMBODIA" latitude="10.6554165" longLatPrecision="1" longitude="104.06017" municipality="Bokor National Park" name="Mt. Bokor" stateProvince="Kampot Province">Mt. Bokor</location>
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,
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<geoCoordinate degrees="10" direction="north" minutes="39" orientation="latitude" precision="1" seconds="19.5" value="10.6554165">10°39'19.5"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="03" orientation="longitude" precision="1" seconds="36.6" value="104.06017">104°03'36.6"E</geoCoordinate>
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, alt.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="9.33" unit="m" value="933.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="9.33" unit="m" value="933.0">933 m</elevation>
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</quantity>
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,
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<collectingDate value="2013-02-16">16 Feb. 2013</collectingDate>
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, with flowers,
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<emphasis italics="true" pageId="2" pageNumber="37">
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<collectorName>Tagane S.</collectorName>
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,
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<collectorName>Toyama H.</collectorName>
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,
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<collectorName>Wachi N.</collectorName>
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,
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<collectorName>Ichihasi R.</collectorName>
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,
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<collectorName>Mase K.</collectorName>
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,
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<collectorName>Zhu M.</collectorName>
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,
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<collectorName>Chhnang P.</collectorName>
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5475
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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KYO!, isotypes BKF!, FU!, K, P, PNP!)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="37" type="description">
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<paragraph pageId="2" pageNumber="37">Description.</paragraph>
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<paragraph pageId="2" pageNumber="37">
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Small tree, 10 m tall. Branchlets glabrous, blackish when dry, sparsely lenticellate, old branches blackish grey or blackish grey brown. Terminal buds 3-7 mm long, bud scales broadly ovate to oblanceolate, densely puberulent outside, glabrous or very sparsely pubescent inside. Leaves alternate, blades elliptic-lanceolate, oblanceolate, (3-)7.5-11.5(-18.5)
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<normalizedToken originalValue="×">x</normalizedToken>
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(1.2-)2.5-4.0(-5.4) cm, thickly leathery, base cuneate, margin entire, slightly revolute, apex usually acute to obtuse, occasionally acuminate or rotund, developing leaves glabrous adaxially, sericeous abaxially, glabrous and densely foveolate on both sides when old, midrib concave adaxially, elevated abaxially, lateral veins 9-12 pairs, flat adaxially, slightly prominent abaxially, tertiary veins reticulate to scalariform-reticulate; petioles (0.5-)0.8-1.3(-2.5) cm long, glabrous. Inflorescences paniculate, subterminal, usually arising from near base of newly sprouted branchlets, (4.0-)6.0-13.0 cm long, sericeous, with 5-8 lateral branches. Pedicels 3-5 mm long, sericeous, bracts linear, 1-4 mm long, abaxially yellowish brown pubescent, adaxially glabrous, caducous. Flowers yellowish green; perianth lobes 6, oblong-elliptic, slightly unequal, outer ones, ca. 6.8
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<normalizedToken originalValue="×">x</normalizedToken>
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2.4 mm, inner ones ca. 7
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<normalizedToken originalValue="×">x</normalizedToken>
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2.5 mm, minutely pubescent on both surfaces. Stamens 9, subequal, 3.5-4 mm long, hairy at base, glands of 3rd series stipitate, ca. 0.4 mm long; staminodes 3, ca. 1 mm long, stipitate, glabrous. Ovary subglobose ca. 1.5 mm in diam., glabrous. Style 2.5-3 mm long, glabrous. Fruits subglobose, 7-8 mm in diam., glabrous, blackish when dry, perianth lobes spreading.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="37" type="materials_examined">
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<paragraph pageId="2" pageNumber="37">Ohter specimens examined.</paragraph>
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<paragraph pageId="2" pageNumber="37">
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<materialsCitation collectingDate="2001-03-09" collectingDateMax="2013-12-17" collectingDateMin="2001-03-09" collectorName="Middleton, Monyrak" country="Cambodia" elevation="1043" latitude="10.663555" location="Bokor National Park" longLatPrecision="1" longitude="104.04814" specimenCount="1">
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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||
<location LSID="urn:lsid:plazi:treatment:99EAAAEDA731EC1D3F4471F17FF4CADF:DADC1F81A9797A28A94A6E380D0376C0" country="Cambodia" latitude="10.663555" longLatPrecision="1" longitude="104.04814" name="Bokor National Park">Bokor National Park</location>
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[alt.
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.043" unit="m" value="1043.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.043" unit="m" value="1043.0">1043 m</elevation>
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</quantity>
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,
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<geoCoordinate degrees="10" direction="north" minutes="37" orientation="latitude" precision="1" seconds="16.8" value="10.621333">10°37'16.8"N</geoCoordinate>
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,
|
||
<geoCoordinate degrees="104" direction="east" minutes="01" orientation="longitude" precision="1" seconds="52.3" value="104.0312">104°01'52.3"E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2013-12-10">10 Dec. 2013</collectingDate>
|
||
,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 6258</emphasis>
|
||
(BKF, FU, PNP); alt.
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.042" unit="m" value="1042.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.042" unit="m" value="1042.0">1042 m</elevation>
|
||
</quantity>
|
||
, 10°38"N,
|
||
<geoCoordinate degrees="104" direction="east" minutes="01.4" orientation="longitude" precision="92" value="104.02333">104°01.4'E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2001-03-09">9 Mar. 2001</collectingDate>
|
||
, with fruits,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">
|
||
<collectorName>Middleton</collectorName>
|
||
&
|
||
<collectorName>Monyrak</collectorName>
|
||
643
|
||
</emphasis>
|
||
(P); alt.
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.014" unit="m" value="1014.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.014" unit="m" value="1014.0">1014 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="10" direction="north" minutes="38" orientation="latitude" precision="1" seconds="12.6" value="10.636833">10°38'12.6"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="02" orientation="longitude" precision="1" seconds="06.4" value="104.03511">104°02'06.4"E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2011-12-04">4 Dec. 2011</collectingDate>
|
||
,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 1556</emphasis>
|
||
(FU, PNP); ibidem,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 1575</emphasis>
|
||
(FU, PNP); alt.
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">1011 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="10" direction="north" minutes="38" orientation="latitude" precision="1" seconds="11.2" value="10.636444">10°38'11.2"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="02" orientation="longitude" precision="1" seconds="09.0" value="104.035835">104°02'09.0"E</geoCoordinate>
|
||
,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 1725</emphasis>
|
||
(FU, PNP); alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.28" unit="m" value="928.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.28" unit="m" value="928.0">928 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="10" direction="north" minutes="39" orientation="latitude" precision="1" seconds="19.5" value="10.6554165">10°39'19.5"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="03" orientation="longitude" precision="1" seconds="36.6" value="104.06017">104°03'36.6"E</geoCoordinate>
|
||
,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 2754</emphasis>
|
||
(FU, PNP); alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.03" unit="m" value="903.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.03" unit="m" value="903.0">903 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="10" direction="north" minutes="39" orientation="latitude" precision="1" seconds="35.4" value="10.659833">10°39'35.4"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="03" orientation="longitude" precision="1" seconds="03.1" value="104.050865">104°03'03.1"E</geoCoordinate>
|
||
,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 3167</emphasis>
|
||
(FU, PNP); alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.709999999999999" unit="m" value="871.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.709999999999999" unit="m" value="871.0">871 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="10" direction="north" minutes="39" orientation="latitude" precision="1" seconds="48.8" value="10.663555">10°39'48.8"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="02" orientation="longitude" precision="1" seconds="53.3" value="104.04814">104°02'53.3"E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2013-12-17">17 Dec. 2013</collectingDate>
|
||
, with flowers,
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Fuse et al. 6227</emphasis>
|
||
(BKF, FU, PNP)]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="37" type="distribution">
|
||
<paragraph pageId="2" pageNumber="37">Distribution.</paragraph>
|
||
<paragraph pageId="2" pageNumber="37">Cambodia (endemic to Mt. Bokor).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="37" type="habitat">
|
||
<paragraph pageId="2" pageNumber="37">Habitat and ecology.</paragraph>
|
||
<paragraph pageId="2" pageNumber="37">
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
is commonly found in moist evergreen forest on the top plateau of Mt. Bokor. Flowering specimens were collected in December and February and fruiting specimens in March.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="37" type="genbank accession no">
|
||
<paragraph pageId="2" pageNumber="37">GenBank accession No.</paragraph>
|
||
<paragraph pageId="2" pageNumber="37">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Toyama et al. 1575</emphasis>
|
||
: AB987667 (
|
||
<emphasis italics="true" pageId="2" pageNumber="37">rbcL</emphasis>
|
||
), AB987666 (
|
||
<emphasis italics="true" pageId="2" pageNumber="37">matK</emphasis>
|
||
), AB987665 (ITS).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="37" type="preliminary conservation assessment">
|
||
<paragraph pageId="2" pageNumber="37">Preliminary conservation assessment.</paragraph>
|
||
<paragraph pageId="2" pageNumber="37">
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
is commonly found in higher elevations in Mt. Bokor. Since many mature individuals were found in the protected areas, we suggest the status for this species as Least Concern (LC) according to IUCN Red List criteria (
|
||
<bibRefCitation author="IUCN" journalOrPublisher="IUCN Species Survival Commission, Gland / Cambridge" pageId="6" pageNumber="41" pagination="1 - 32" refId="B10" refString="IUCN, 2012. IUCN red list categories and criteria, version 3.1. IUCN Species Survival Commission, Gland/Cambridge: 1 - 32" title="IUCN red list categories and criteria, version 3.1" year="2012">IUCN 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37" type="notes">
|
||
<paragraph pageId="2" pageNumber="37">Note.</paragraph>
|
||
<paragraph lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37">
|
||
Among five species we collected in Mt. Bokor, this species is most common and agrees with the description of "
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
" in the key of
|
||
<bibRefCitation author="Liou, H" journalOrPublisher="Hermann & Cie, Paris" pageId="6" pageNumber="41" refId="B13" refString="Liou, H, 1934. Lauracees de Chine et d'Indochine. Hermann & Cie, Paris" title="Lauracees de Chine et d'Indochine" year="1934">Liou (1934)</bibRefCitation>
|
||
who recorded "
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
" from Nord de Kampot, corresponding to Mt. Bokor. Thus, we consider that this species is "
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
" recorded by
|
||
<bibRefCitation author="Liou, H" journalOrPublisher="Hermann & Cie, Paris" pageId="6" pageNumber="41" refId="B13" refString="Liou, H, 1934. Lauracees de Chine et d'Indochine. Hermann & Cie, Paris" title="Lauracees de Chine et d'Indochine" year="1934">Liou (1934)</bibRefCitation>
|
||
, although we could not examine the specimen of
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Poilane 14625</emphasis>
|
||
(P) cited by
|
||
<bibRefCitation author="Liou, H" journalOrPublisher="Hermann & Cie, Paris" pageId="6" pageNumber="41" refId="B13" refString="Liou, H, 1934. Lauracees de Chine et d'Indochine. Hermann & Cie, Paris" title="Lauracees de Chine et d'Indochine" year="1934">Liou (1934)</bibRefCitation>
|
||
because it is currently on loan. In Indo-china,
|
||
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus" order="Laurales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="2" pageNumber="37">Machilus</emphasis>
|
||
</taxonomicName>
|
||
plants similar to this
|
||
<pageBreakToken pageId="3" pageNumber="38" start="start">species</pageBreakToken>
|
||
have been identified as
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
for a long time (
|
||
<bibRefCitation author="Lecomte, H" journalOrPublisher="Flore generale de l'Indo-Chine" pageId="6" pageNumber="41" pagination="107 - 158" refId="B11" refString="Lecomte, H, 1914. Lauracees. Flore generale de l'Indo-Chine 5: 107 - 158" title="Lauracees" volume="5" year="1914">Lecomte 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Liou, H" journalOrPublisher="Hermann & Cie, Paris" pageId="6" pageNumber="41" refId="B13" refString="Liou, H, 1934. Lauracees de Chine et d'Indochine. Hermann & Cie, Paris" title="Lauracees de Chine et d'Indochine" year="1934">Liou 1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Phon, Dy" journalOrPublisher="Imprimerie Olympic, Phnom Penh" pageId="6" pageNumber="41" refId="B18" refString="Phon, Dy, 2000. Dictionary of Plants used in Cambodia. Imprimerie Olympic, Phnom Penh" title="Dictionary of Plants used in Cambodia" year="2000">Phon 2000</bibRefCitation>
|
||
). However,
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
is distinct from
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
distributed in Nepal, E Himalaya, Assam-Burma in sericeous inflorescence rachis (vs. glabrous in
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Pendry, CA" journalOrPublisher="Royal Botanic Garden Edinburgh, Edinburgh" pageId="6" pageNumber="41" refId="B17" refString="Pendry, CA, 2012. Flora of Nepal. Royal Botanic Garden Edinburgh, Edinburgh" title="Flora of Nepal" year="2012">Pendry 2012</bibRefCitation>
|
||
), perianth lobes pubescent on both surfaces (vs. outside almost glabrous), ca. 1 mm staminodes (vs. ca. 2 mm), and blackish grey twigs (vs. reddish). In Cambodia, we collected sterile specimens of
|
||
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus</emphasis>
|
||
</taxonomicName>
|
||
spp. morphologically similar to
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
in Central Cardamon and Seima protection forests but those are morphologically distinguishable and not sister to
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
in our unpublished phylogenetic tree constructed by
|
||
<emphasis italics="true" pageId="3" pageNumber="38">rbcL</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="3" pageNumber="38">matK</emphasis>
|
||
and ITS sequences. "
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
" is recorded from Mulu-prey, Preah Vihear (
|
||
<bibRefCitation author="Lecomte, H" journalOrPublisher="Flore generale de l'Indo-Chine" pageId="6" pageNumber="41" pagination="107 - 158" refId="B11" refString="Lecomte, H, 1914. Lauracees. Flore generale de l'Indo-Chine 5: 107 - 158" title="Lauracees" volume="5" year="1914">Lecomte 1914</bibRefCitation>
|
||
). The specimens from Mulu-prey (
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Harmand s.n.</emphasis>
|
||
, P) are, however, different from
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
in having naked terminal buds and scalariform tertiary veins.
|
||
<bibRefCitation author="Lecomte, H" journalOrPublisher="Flore generale de l'Indo-Chine" pageId="6" pageNumber="41" pagination="107 - 158" refId="B11" refString="Lecomte, H, 1914. Lauracees. Flore generale de l'Indo-Chine 5: 107 - 158" title="Lauracees" volume="5" year="1914">Lecomte (1914)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Liou, H" journalOrPublisher="Hermann & Cie, Paris" pageId="6" pageNumber="41" refId="B13" refString="Liou, H, 1934. Lauracees de Chine et d'Indochine. Hermann & Cie, Paris" title="Lauracees de Chine et d'Indochine" year="1934">Liou (1934)</bibRefCitation>
|
||
recorded "
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
" also from Laos and Vietnam, but we have not found specimens from Laos and Vietnam identical with
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
. As far as we know,
|
||
<taxonomicName authorityName="Yahara & Tagane & Mase & Chhang & Toyama" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus odoratissima" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="odoratissima">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus odoratissima</emphasis>
|
||
</taxonomicName>
|
||
is not distributed in Cambodia, Laos, Thailand and Vietnam. The relationship between
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
and the specimens of Mulu-prey, Laos and Vietnam remains to be further examined. At present, we consider that
|
||
<taxonomicName authorityName="Yahara & Tagane" authorityYear="2016" class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus bokorensis" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="bokorensis">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus bokorensis</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Bokor, Cambodia. According to our unpublished ITS tree, this species is sister to
|
||
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus rimosa" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="rimosa">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus rimosa</emphasis>
|
||
</taxonomicName>
|
||
(Blume) Blume distributed in Indonesia, but distinct from
|
||
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus rimosa" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="rimosa">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus rimosa</emphasis>
|
||
</taxonomicName>
|
||
in thickly leathery leaves densely foveolate on both sides (vs. thinner leaves not densely foveolate in
|
||
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Machilus rimosa" order="Laurales" pageId="3" pageNumber="38" phylum="Tracheophyta" rank="species" species="rimosa">
|
||
<emphasis italics="true" pageId="3" pageNumber="38">Machilus rimosa</emphasis>
|
||
</taxonomicName>
|
||
) and perianths larger (6.8-7 mm long vs. ca. 3.5 mm long).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |