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<document id="ABCB0F6F7B88C9832DFDE2F4985F1C44" ENCODING="UTF-8" ID-GBIF-Dataset="0922c4b9-e8eb-4b0a-a266-679f0affe1ae" ModsDocID="z01605p033" checkinTime="1247317115952" checkinUser="thomas" docAuthor="Marcelo R. S. Melo, H. J. Walker, Jr. &amp; Cynthia Klepadlo" docDate="2007" docId="47CFF8659A766A41AFD144E692EE69E5" docLanguage="en" docName="2007_Melo_Walker_Klepadlo_gg1.xml" docOrigin="Zootaxa 1605" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:9562F15A-149A-41A7-9ECC-F2EEC1DD67A1" docTitle="Pseudoscopelus Luetken" docType="treatment" docVersion="13" lastPageNumber="33" masterDocId="F48720AFBF04EBE4BA50EA661DAB0FB6" masterDocTitle="Two new species of Pseudoscopelus (Teleostei: Chiasmodontidae), with a new diagnosis for the genus." masterLastPageNumber="46" masterPageNumber="33" pageNumber="33" updateTime="1701328240161" updateUser="ExternalLinkService">
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<mods:title id="17F63AB19DA215C95766D1566A42AC2B">Two new species of Pseudoscopelus (Teleostei: Chiasmodontidae), with a new diagnosis for the genus.</mods:title>
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<mods:namePart id="225FC983B7C84CD1589C5D25B900B943">Marcelo R. S. Melo</mods:namePart>
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<mods:namePart id="ABDAB11D2D2E6BF2A87BB4995B769929">H. J. Walker, Jr.</mods:namePart>
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<mods:namePart id="DB17542A189B5C9D6FA8422A88224F6A">Cynthia Klepadlo</mods:namePart>
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<mods:date id="2A954422D9AA44F4C574A176D0D7E148">2007</mods:date>
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<treatment id="47CFF8659A766A41AFD144E692EE69E5" ID-DOI="http://doi.org/10.5281/zenodo.6243401" ID-GBIF-Taxon="100121778" ID-Zenodo-Dep="6243401" LSID="urn:lsid:plazi:treatment:47CFF8659A766A41AFD144E692EE69E5" httpUri="http://treatment.plazi.org/id/47CFF8659A766A41AFD144E692EE69E5" lastPageNumber="33" pageNumber="33">
<subSubSection id="E88F57A8D55DF2374801C6134ADB6093" type="nomenclature">
<paragraph id="D94E3EF538DB7D426F5F82F4DB678CD7">
[[
<taxonomicName id="82B2F4B382DB3F7F6A4044F2A8989F4E" ID-CoL="73HD" ID-ENA="1306755" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">
Pseudoscopelus
<normalizedToken id="E089226EA36B75C40F31605C419EA02E" originalValue="Lütken">Luetken</normalizedToken>
</taxonomicName>
]]
</paragraph>
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<subSubSection id="8CB8B9B493F0718CD13032A7A1EE902A" type="discussion">
<paragraph id="A8041CF99564B4140E39EF2027E84B0B" pageNumber="33">
Four genera are currently recognized in Chiasmodontidae, one of the few families of bathy- and mesopelagic fishes in the large actinopterygian order Perciformes (Nelson, 2006):
<taxonomicName id="BDA77159C2AC4DB05F2B3F13CE1C80FC" LSID-ZBK="urn:lsid:zoobank.org:act:B9543C54-6827-40B2-8D80-EED6C11FB344" family="Chiasmodontidae" genus="Chiasmodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chiasmodon Johnson 1864:408" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Chiasmodon Johnson</taxonomicName>
(1864);
<taxonomicName id="FEBF3E632402F64DE5FE7A68741DDB89" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">
Pseudoscopelus
<normalizedToken id="728DC0C9B077942FD7507DB343E6A354" originalValue="Lütken">Luetken</normalizedToken>
</taxonomicName>
(1892);
<taxonomicName id="F88B950BEA7C25DDBC3E62B280D77B7F" LSID-ZBK="urn:lsid:zoobank.org:act:CBDF8830-9BA7-4116-81E6-11C317AFC452" family="Chiasmodontidae" genus="Dysalotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Dysalotus MacGilchrist 1905:268" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Dysalotus MacGilchrist</taxonomicName>
(1905); and
<taxonomicName id="39569F1204914448299DA04C4D4807C8" LSID-ZBK="urn:lsid:zoobank.org:act:10B4DB64-6B5C-448A-9FF0-523395D70A67" family="Chiasmodontidae" genus="Kali" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Kali Lloyd 1909:154" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Kali Lloyd</taxonomicName>
(1909).
<taxonomicName id="E36953C7BAFAB219E97E56F633A99255" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
is the most species rich genus with 15 nominal species, 12 of which are valid (Table 1). Most of these species have light organs, which are restricted to
<taxonomicName id="D0ACC9904F8BAD42D409CE958039063F" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
within the family. The genus has a worldwide distribution, being found in all major ocean basins, even off Antarctica, but is absent from the Mediterranean and Red Seas. In the present study, two new species of
<taxonomicName id="F1DF34D25491F7BBD4AC6AD2B857D3C4" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
are described,
<taxonomicName id="1CCBB55833BFCC72D89E9ADFBC5D933A" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="lavenbergi">P. lavenbergi</taxonomicName>
from the eastern Pacific, and
<taxonomicName id="1A5699B69E3D3576A3B589B461157BA8" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="bothrorrhinos">P. bothrorrhinos</taxonomicName>
from the western Pacific and Indian Oceans.
</paragraph>
<paragraph id="915F7DA052C87849C89393BF8DC30220" pageNumber="33">
The genus
<taxonomicName id="AB74E832E6DD8D794122C7062BBD36C4" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
was initially diagnosed by
<normalizedToken id="905B4F2D8003D85A6467F2BD749FB485" originalValue="Lütken">Luetken</normalizedToken>
(1892) based on the presence of lines of mucous pores [sic] along the body. This definition had been followed by subsequent authors (e.g., Tanaka, 1908; Norman, 1929), until Beebes (1932) discovery that instead of mucous pits, the lines over the body of
<taxonomicName id="1572A429AA26940036AD0B556C0B3890" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
were actually formed by photophores that produce a clear, bright light. Since Beebes work, the genus has been defined by the presence of discrete photophores along the body (e.g., Mooi and Paxton, 2001; Nelson, 2006). In 1974, Lavenberg revised the genus
<taxonomicName id="3DD7D0BAB803AA6750F9E7F792043675" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
for his PhD; he considered six species to be valid and listed six other new taxa, but never published his results according to the ICZN (1999) rules. The most recent works have been published by Prokofiev and Kukuev (2005, 2006a, b, c) who described four new species; two of which,
<taxonomicName id="6879C6885AA7EAD6DCE09E7C1AD6FDFF" LSID-ZBK="urn:lsid:zoobank.org:act:0A87D071-7FCE-4735-A63E-28D84313C501" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus aphos Prokofiev &amp; Kukuev 2005" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="aphos">P. aphos</taxonomicName>
and
<taxonomicName id="48E608615D0CD46A61A402CF399B47C0" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="parini">P. parini</taxonomicName>
, totally lack discrete photophores. Prokofiev and Kukuev (2006c), however, still regarded the presence of photophores as a generic character.
</paragraph>
<paragraph id="07FED134B0BB69E45FAD0ECE4A4A3D50" pageNumber="33">
<taxonomicName id="29713E09388FC39EF7563795E8625242" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
is diagnosed herein by two synapomorphies, present in all species. Using this as the diagnosis would also include
<taxonomicName id="13171FEEFE8654BB1808F5423F2382CE" LSID-ZBK="urn:lsid:zoobank.org:act:86B3784A-A207-483D-8652-DB48465F86B3" family="Chiasmodontidae" genus="Myersiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Myersiscus Fowler 1934:361" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Myersiscus Fowler</taxonomicName>
(1934), which was erected during the description of
<taxonomicName id="53135ACDC1A3188BF4AE022E2061CD2C" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="obtusifrons">P. obtusifrons</taxonomicName>
and was then considered a junior synonym of
<taxonomicName id="01D8F64408EF1D1DBA801698AEE6FAA4" LSID-ZBK="urn:lsid:zoobank.org:act:55535F07-62E9-4F6D-B657-E2A1C6E9463F" family="Chiasmodontidae" genus="Pseudoscopelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudoscopelus Lütken 1892:285" order="Perciformes" pageNumber="33" phylum="Chordata" rank="genus">Pseudoscopelus</taxonomicName>
exclusively because of the presence of photophores (Mooi and Paxton, 2001; Prokofiev and Kukuev, 2005, 2006c).
</paragraph>
</subSubSection>
</treatment>
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