263 lines
33 KiB
XML
263 lines
33 KiB
XML
<document id="131279CE933ED928E322DD25A049764B" ID-CLB-Dataset="29797" ID-DOI="10.11646/zootaxa.4442.1.8" ID-GBIF-Dataset="e30f7fa7-414f-45e4-8478-9f0a64a1b405" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301308" ID-ZooBank="7E50F2C6-4BBC-4D37-809A-5357877BFFBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530294012709" checkinUser="plazi" docAuthor="Bukejs, Andris & Alekseev, Vitalii I." docDate="2018" docId="013A87AFEB49CA4139D4FA39B76BFE49" docLanguage="en" docName="zootaxa.4442.1.8.pdf" docOrigin="Zootaxa 4442 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ataenius damzeni Bukejs & Alekseev 2018, new species" docType="treatment" docUuid="1D52A7C1-3DCB-427B-9A8C-C1663667E803" docUuidSource="ZooBank" docVersion="8" lastPageNumber="158" masterDocId="FD03FFD7EB48CA443943FFFCB606FFE5" masterDocTitle="A new extinct species of Ataenius Harold from Baltic amber (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="160" masterPageNumber="153" pageNumber="154" updateTime="1698551771271" updateUser="ExternalLinkService">
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<mods:mods id="CF115FDFCC3F2C9AC3EFA5CFF57E90BB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F97D4CA6BF00377FFB57F6F15E6BC044">
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<mods:title id="5F6136C52718FBE9AE922C315FC92633">A new extinct species of Ataenius Harold from Baltic amber (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:name id="FE299C22932177AE9A9140E2DF0339EB" type="personal">
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<mods:namePart id="B21F64A6B8D949C3623942E78F3A8238">Bukejs, Andris</mods:namePart>
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<mods:roleTerm id="A08F9DA688BA943D6ED12B1ADFFCCA42">Author</mods:roleTerm>
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<mods:namePart id="BDE3E25938DE9784CD226AB7648036F7">Alekseev, Vitalii I.</mods:namePart>
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<mods:typeOfResource id="8C063FE2899EA20057678A4F0F4284D1">text</mods:typeOfResource>
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<mods:title id="15FC8A632D072CB2C34CD6FCCF03791D">Zootaxa</mods:title>
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<mods:part id="0CC70474CA6537DE23B4B345D3E2C3F8">
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<mods:date id="0FE1278D266A7347318F53CD3E3D52E0">2018</mods:date>
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<mods:number id="FA848A9A849B0413D82B431556ACEDDA">2018-06-29</mods:number>
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<mods:number id="95C8AED1AD1A703F195A197E7CACA9A7">4442</mods:number>
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<mods:identifier id="06C1B274916F998CA51868B223F9F6CB" type="DOI">10.11646/zootaxa.4442.1.8</mods:identifier>
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<mods:identifier id="48EF72C7CCF45FA44534972278DFD54A" type="GBIF-Dataset">e30f7fa7-414f-45e4-8478-9f0a64a1b405</mods:identifier>
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<treatment id="013A87AFEB49CA4139D4FA39B76BFE49" ID-DOI="http://doi.org/10.5281/zenodo.5977191" ID-GBIF-Taxon="146043465" ID-Zenodo-Dep="5977191" LSID="urn:lsid:zoobank.org:act:1D52A7C1-3DCB-427B-9A8C-C1663667E803" httpUri="http://treatment.plazi.org/id/013A87AFEB49CA4139D4FA39B76BFE49" lastPageId="5" lastPageNumber="158" pageId="1" pageNumber="154">
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<subSubSection id="C1896532EB49CA4539D4FA39B714F9E5" pageId="1" pageNumber="154" type="nomenclature">
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<paragraph id="892C36B9EB49CA4539D4FA39B4F8FA3A" blockId="1.[151,766,1477,1536]" box="[151,766,1477,1503]" pageId="1" pageNumber="154">
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<heading id="D26481D5EB49CA4539D4FA39B4F8FA3A" bold="true" box="[151,766,1477,1503]" fontSize="11" level="1" pageId="1" pageNumber="154" reason="1">
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<emphasis id="BBE7EAABEB49CA4539D4FA39B4F8FA3A" bold="true" box="[151,766,1477,1503]" pageId="1" pageNumber="154">
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<taxonomicName id="4E934D3AEB49CA4539D4FA39B464FA3A" ID-CoL="J7JF" authority="Bukejs & Alekseev" authorityName="Bukejs & Alekseev" authorityYear="2018" box="[151,610,1477,1503]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
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<emphasis id="BBE7EAABEB49CA4539D4FA39B768FA3A" bold="true" box="[151,366,1477,1503]" italics="true" pageId="1" pageNumber="154">Ataenius damzeni</emphasis>
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Bukejs & Alekseev
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</taxonomicName>
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,
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<taxonomicNameLabel id="A0D457D0EB49CA453B2DFA39B4F8FA3A" box="[622,766,1477,1503]" pageId="1" pageNumber="154" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="892C36B9EB49CA4539D4FA1BB714F9E5" blockId="1.[151,766,1477,1536]" box="[151,274,1511,1536]" pageId="1" pageNumber="154">
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(
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<figureCitation id="11A82A3CEB49CA4539DCFA1BB70FF9E5" box="[159,265,1511,1536]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStart-2="FIGURES 5–7" captionStartId-0="2.[151,264,1969,1992]" captionStartId-1="3.[151,264,1683,1706]" captionStartId-2="4.[151,264,1617,1640]" captionTargetBox-0="[384,1202,916,1946]" captionTargetBox-1="[384,1202,589,1662]" captionTargetBox-2="[384,1202,481,1597]" captionTargetId-0="figure@2.[384,1202,916,1946]" captionTargetId-1="figure@3.[384,1202,589,1662]" captionTargetId-2="figure@4.[384,1202,481,1597]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–2. Ataenius damzeni Bukejs & Alekseev, new species, holotype “5953-01”. 1, habitus, dorsofrontal view; 2, habitus, ventrolateral view. Scale bar = 1 mm." captionText-1="FIGURES 3–4. Ataenius damzeni Bukejs & Alekseev, new species, paratype “5953-02”. 3, habitus, lateral view; 4, habitus, laterocaudal view. Scale bar = 1 mm." captionText-2="FIGURES 5–7. Ataenius damzeni Bukejs & Alekseev, new species. 5, general view of amber piece “5953” with specimen enumeration (1—holotype; 2—paratype; 3‒4—non-type specimens); 6, details of pronotum and head, dorsofrontal view (holotype “5953-01”); 7, details of abdomen and hind left leg (holotype “5953-01”). Scale bar = 5 mm for Fig. 5; 0.25 mm for Figs. 6 and 7." httpUri-0="https://zenodo.org/record/1301310/files/figure.png" httpUri-1="https://zenodo.org/record/1301312/files/figure.png" httpUri-2="https://zenodo.org/record/1301314/files/figure.png" pageId="1" pageNumber="154">Figs. 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C1896532EB49CA4539D4F9D3B350F8C5" pageId="1" pageNumber="154" type="materials_examined">
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<paragraph id="892C36B9EB49CA4539D4F9D3B290F988" blockId="1.[151,1437,1583,2005]" pageId="1" pageNumber="154">
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<emphasis id="BBE7EAABEB49CA4539D4F9D3B741F9AD" bold="true" box="[151,327,1583,1608]" pageId="1" pageNumber="154">Type material.</emphasis>
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Holotype: “5953-01” (MAIG), adult, complete, sex unknown. Ventral side of specimen partially obscured by “milky” opacity. Paratype: “5953-02” (MAIG), adult, complete, sex unknown.
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</paragraph>
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<paragraph id="892C36B9EB49CA453984F98BB2C8F93D" blockId="1.[151,1437,1583,2005]" pageId="1" pageNumber="154">
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<emphasis id="BBE7EAABEB49CA453984F98BB43FF975" bold="true" box="[199,569,1655,1680]" pageId="1" pageNumber="154">Additional material examined.</emphasis>
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“5953-03” (
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<collectionCode id="EF82AE7CEB49CA453B8AF984B51FF975" box="[713,793,1656,1680]" pageId="1" pageNumber="154">MAIG</collectionCode>
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), adult, damaged specimen without head, sex unknown. Dorsal side of specimen partially and ventral side of specimen almost completely obscured by “milky” opacity. “5953-04” (
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<collectionCode id="EF82AE7CEB49CA45385FF93CB76BF93D" box="[284,365,1728,1752]" pageId="1" pageNumber="154">MAIG</collectionCode>
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), adult, damaged specimen with separated pronotum and head, sex unknown.
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</paragraph>
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<paragraph id="892C36B9EB49CA453984F918B54DF919" blockId="1.[151,1437,1583,2005]" box="[199,843,1764,1789]" pageId="1" pageNumber="154">
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<materialsCitation id="39FB3CE4EB49CA453984F918B54DF919" ID-GBIF-Occurrence="1852084114" box="[199,843,1764,1789]" location="Baltic" municipality="Upper Eocene." pageId="1" pageNumber="154" specimenCount="1" typeStatus="holotype">
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<emphasis id="BBE7EAABEB49CA453984F918B753F918" bold="true" box="[199,341,1764,1789]" pageId="1" pageNumber="154">
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<typeStatus id="5628881BEB49CA453984F918B704F918" box="[199,258,1764,1789]" pageId="1" pageNumber="154">Type</typeStatus>
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strata.
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</emphasis>
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<location id="8C4C6062EB49CA45381FF918B799F918" LSID="urn:lsid:plazi:treatment:013A87AFEB49CA4139D4FA39B76BFE49:8C4C6062EB49CA45381FF918B799F918" box="[348,415,1764,1789]" municipality="Upper Eocene." name="Baltic" pageId="1" pageNumber="154">Baltic</location>
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amber, mid-Eocene to
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<collectingMunicipality id="6948ACC3EB49CA453BE7F919B54DF919" box="[676,843,1765,1788]" pageId="1" pageNumber="154">Upper Eocene.</collectingMunicipality>
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</materialsCitation>
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</paragraph>
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<paragraph id="892C36B9EB49CA453984F8FBB350F8C5" blockId="1.[151,1437,1583,2005]" box="[199,1366,1799,1824]" pageId="1" pageNumber="154">
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<materialsCitation id="39FB3CE4EB49CA453984F8FBB350F8C5" ID-GBIF-Occurrence="1852084113" box="[199,1366,1799,1824]" country="Russia" county="Palmnicken" location="Baltic Sea" municipality="Yantarny" pageId="1" pageNumber="154" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="BBE7EAABEB49CA453984F8FBB763F8C5" bold="true" box="[199,357,1799,1824]" pageId="1" pageNumber="154">
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<typeStatus id="5628881BEB49CA453984F8FBB704F8C5" box="[199,258,1799,1824]" pageId="1" pageNumber="154">Type</typeStatus>
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locality.
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</emphasis>
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<location id="8C4C6062EB49CA45382FF8FBB7D9F8C5" LSID="urn:lsid:plazi:treatment:013A87AFEB49CA4139D4FA39B76BFE49:8C4C6062EB49CA45382FF8FBB7D9F8C5" box="[364,479,1799,1824]" country="Russia" county="Palmnicken" municipality="Yantarny" name="Baltic Sea" pageId="1" pageNumber="154" stateProvince="Kaliningrad">Baltic Sea</location>
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coast,
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<collectingMunicipality id="6948ACC3EB49CA453B6EF8F4B494F8C5" box="[557,658,1800,1824]" pageId="1" pageNumber="154">Yantarny</collectingMunicipality>
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settlement [formerly
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<collectingCounty id="604D4E35EB49CA453AC5F8FBB216F8C5" box="[902,1040,1799,1824]" pageId="1" pageNumber="154">Palmnicken</collectingCounty>
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],
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<collectingRegion id="4B57F85BEB49CA453D62F8FBB2A1F8C5" box="[1057,1191,1799,1824]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="154">Kaliningrad</collectingRegion>
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region,
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<collectingCountry id="F1847629EB49CA453C40F8FBB354F8C5" box="[1283,1362,1799,1824]" name="Russia" pageId="1" pageNumber="154">Russia</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1896532EB49CA403984F8D0B5ACFE80" lastPageId="4" lastPageNumber="157" pageId="1" pageNumber="154" type="description">
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<paragraph id="892C36B9EB49CA453984F8D0B351F882" blockId="1.[151,1437,1583,2005]" pageId="1" pageNumber="154">
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<emphasis id="BBE7EAABEB49CA453984F8D0B75FF8A0" bold="true" box="[199,345,1836,1861]" pageId="1" pageNumber="154">Description.</emphasis>
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<emphasis id="BBE7EAABEB49CA453829F8D0B7CCF8A0" box="[362,458,1836,1861]" italics="true" pageId="1" pageNumber="154">Habitus.</emphasis>
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Body moderately convex, elongate oval, subparallel sided, slightly alutaceous, unicolorous black (as preserved), glabrous. Total body length
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<quantity id="4E6B9B5CEB49CA453A0FF8ACB5A4F882" box="[844,930,1872,1896]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="154" unit="mm" value="2.8">2.8 mm</quantity>
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(excluding head); body width
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<quantity id="4E6B9B5CEB49CA453DB9F8B3B355F882" box="[1274,1363,1871,1896]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="154" unit="mm" value="1.5">1.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="892C36B9EB49CA463984F888B5C9FF1D" blockId="1.[151,1437,1583,2005]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="155" pageId="1" pageNumber="154">
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<emphasis id="BBE7EAABEB49CA453984F888B702F868" box="[199,260,1908,1933]" italics="true" pageId="1" pageNumber="154">Head</emphasis>
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. Head transverse, convex, without transverse carina, slightly narrower than pronotal anterior margin; anterior half of head with microreticulation and sparse, very fine punctures, and posterior half with dense, fine punctures. Distance between punctures approximately 0.5–1.5× diameter of one puncture. Clypeus dilated, covering mouthparts; clypeal margin narrowly reflexed, rounded on each side of shallow, wide, median emargination; sides almost straight anterior to genae. Frontal suture shallow, and trisinuate. Compound eyes not visible from directly above. Antennae not visible in examined specimens.
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</paragraph>
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<paragraph id="892C36B9EB4ACA463984FEF8B2E1FE10" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="155">
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<emphasis id="BBE7EAABEB4ACA463984FEF8B71EFEF8" box="[199,280,260,285]" italics="true" pageId="2" pageNumber="155">Thorax</emphasis>
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. Pronotum transverse, about 1.5× as wide as long, and not explanate. Disc evenly and slightly convex, anterolateral region with slightly expanded area. Pronotal punctation distinctly unequal in size: dense and small in anterior one-third of pronotal length, and distinctly larger and sparser posteriorly. Distance between punctures approximately 0.5–3.0× diameter of one puncture, interspaces with secondary punctation and microreticulation. Posterior pronotal margin slightly convex, narrowly bordered, and crenulate. Lateral margins slightly and evenly convex, slightly upturned, narrowly bordered, crenulate, without setae or scales. Anterior margin straight and without bordering. Anterior angles obtusely rounded, protruding; posterior angles widely rounded.
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</paragraph>
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<paragraph id="892C36B9EB4ACA463984FE03B382FDF2" blockId="2.[151,1437,151,861]" box="[199,1412,511,536]" pageId="2" pageNumber="155">Scutellar shield relatively small, elongate, rounded apically, about 1.7× as long as wide, with fine punctures.</paragraph>
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<paragraph id="892C36B9EB4ACA463984FDD8B55CFD0A" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="155">Elytra subparallel-sided, slightly tapering posteriorly, moderately convex, elongate, about 1.6× as long as their combined width, 2.0× as long as pronotum, and widest at mid-length. Anterior margin of elytra slightly narrower than pronotum; lateral margins without setae. Humerus apparently without distinct denticle. Each elytron with 10 deep, punctate striae; punctures small, shallow, apparently elongate. Interstriae distinctly convex, with microreticulation and minute secondary punctation; elytral interstriae equal in height. Epipleura wide at humeri and gradually narrowing toward elytral apex, reaching elytral apex.</paragraph>
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<paragraph id="892C36B9EB4ACA463984FD01B214FCB8" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="155">Hypomeron finely and densely punctate. Mesosternum carinate between coxae; mesocoxae distinctly separated from epipleura. Metasternum slightly convex; disc almost flat with midline impressed, with sparse, small punctures; lateral area and lateral metasternal triangle impunctate and scabrous.</paragraph>
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<caption id="DDEC6631EB4ACA4639D4F84DB46FF803" httpUri="https://zenodo.org/record/1301310/files/figure.png" pageId="2" pageNumber="155" startId="2.[151,264,1969,1992]" targetBox="[384,1202,916,1946]" targetPageId="2">
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<paragraph id="892C36B9EB4ACA4639D4F84DB46FF803" blockId="2.[151,1435,1969,2022]" pageId="2" pageNumber="155">
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<emphasis id="BBE7EAABEB4ACA4639D4F84DB73BF823" bold="true" box="[151,317,1969,1992]" pageId="2" pageNumber="155">FIGURES 1–2.</emphasis>
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<taxonomicName id="4E934D3AEB4ACA463804F84EB4D5F822" authority="Bukejs & Alekseev" authorityName="Bukejs & Alekseev" authorityYear="2018" box="[327,723,1969,1991]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
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<emphasis id="BBE7EAABEB4ACA463804F84EB7FBF822" box="[327,509,1969,1991]" italics="true" pageId="2" pageNumber="155">Ataenius damzeni</emphasis>
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Bukejs & Alekseev
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</taxonomicName>
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,
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<taxonomicNameLabel id="A0D457D0EB4ACA463BA3F84EB566F822" box="[736,864,1969,1991]" pageId="2" pageNumber="155" rank="species">
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<emphasis id="BBE7EAABEB4ACA463BA3F84EB566F822" bold="true" box="[736,864,1969,1991]" pageId="2" pageNumber="155">new species</emphasis>
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</taxonomicNameLabel>
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, holotype “5953-01”. 1, habitus, dorsofrontal view; 2, habitus, ventrolateral view. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="892C36B9EB4BCA473984FF65B563FDFD" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="156">
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<emphasis id="BBE7EAABEB4BCA473984FF65B6FAFF55" box="[199,252,153,176]" italics="true" pageId="3" pageNumber="156">Legs</emphasis>
|
||
. Profemur wide, about 1.6× as long as wide, with few fine punctures; perimarginal groove distinct; posterior margin with short, erect seta medially. Meso- and metafemora narrower than profemur, nearly equal in size, with few fine punctures, and sharp posterior femoral line. Protibia rather short, flattened and strongly expanded in apical portion, impunctate, nearly as long as profemur; protibia with terminal spur, slightly arcuate ventrally; with three large outer denticles in apical half. Meso- and metatibiae weakly expanded apically, without oblique carinae; with two terminal spurs of unequal length, located close together and below tarsal insertion, superior spur about as long as tarsomere 1 and about 2× as long as inferior spur; apex of meso- and metatibia bearing fringe of spine-like accessory setae that are unequal in size, and accessory spine apparently lacking. Tarsi slender, 5-segmented, without dense setae ventrally, and with few setae apically. Meso- and metatarsi nearly as long as adjacent tibiae; meso- and metatarsomere 1 about as long as superior tibial spur, and about 1.2–1.4× as long as tarsomeres 2–5 combined. Claws small, simple, symmetrical.
|
||
</paragraph>
|
||
<caption id="DDEC6631EB4BCA4739D4F96FB409F92D" httpUri="https://zenodo.org/record/1301312/files/figure.png" pageId="3" pageNumber="156" startId="3.[151,264,1683,1706]" targetBox="[384,1202,589,1662]" targetPageId="3">
|
||
<paragraph id="892C36B9EB4BCA4739D4F96FB409F92D" blockId="3.[151,1436,1683,1736]" pageId="3" pageNumber="156">
|
||
<emphasis id="BBE7EAABEB4BCA4739D4F96FB73AF94C" bold="true" box="[151,316,1683,1706]" pageId="3" pageNumber="156">FIGURES 3–4.</emphasis>
|
||
<taxonomicName id="4E934D3AEB4BCA473800F968B4CEF94F" authority="Bukejs & Alekseev" authorityName="Bukejs & Alekseev" authorityYear="2018" box="[323,712,1684,1706]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
|
||
<emphasis id="BBE7EAABEB4BCA473800F968B7FEF94F" box="[323,504,1684,1706]" italics="true" pageId="3" pageNumber="156">Ataenius damzeni</emphasis>
|
||
Bukejs & Alekseev
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A0D457D0EB4BCA473B97F968B555F94F" box="[724,851,1684,1706]" pageId="3" pageNumber="156" rank="species">
|
||
<emphasis id="BBE7EAABEB4BCA473B97F968B555F94F" bold="true" box="[724,851,1684,1706]" pageId="3" pageNumber="156">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype “5953-02”. 3, habitus, lateral view; 4, habitus, laterocaudal view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="892C36B9EB4BCA473984F909B362F827" blockId="3.[151,1437,1781,2022]" pageId="3" pageNumber="156">
|
||
<emphasis id="BBE7EAABEB4BCA473984F909B737F8EB" box="[199,305,1781,1806]" italics="true" pageId="3" pageNumber="156">Abdomen</emphasis>
|
||
. Shiny, with five visible, coalesced ventrites. Ventrite 1 impunctate, scabrous, with slightly elevated triangular area anteromedially; ventrites 2–5 with small, sparse punctures; punctation of ventrites 2 and 3 slightly sparser. Ventrites 2–5 fluted along anterior margin; ventrites 2 and 5 finely fluted, and ventrites 3 and 4 with stronger fluting. Ventrite 5 with two erect setae on disc. Relative length ratios of ventrites 1–5 (medially) equal to 9:10:10:10:10. Pygidium partially exposed, and apparently with short median transverse carina; pygidial surface scabrous and slightly semicircularly impressed in apical portion; bearing two erect setae near anterior margin.
|
||
</paragraph>
|
||
<paragraph id="892C36B9EB4BCA403984F831B4A8FF1D" blockId="3.[151,1437,1781,2022]" lastBlockId="4.[151,1436,151,429]" lastPageId="4" lastPageNumber="157" pageId="3" pageNumber="156">
|
||
<emphasis id="BBE7EAABEB4BCA473984F831B74BF803" bold="true" box="[199,333,1997,2022]" pageId="3" pageNumber="156">Variability.</emphasis>
|
||
Body length excluding head (head turned downwards)
|
||
<quantity id="4E6B9B5CEB4BCA473A9AF832B263F800" box="[985,1125,1998,2022]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="3" pageNumber="156" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4E6B9B5CEB4BCA473C7FF831B39EF800" box="[1340,1432,1997,2022]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="156" unit="mm" value="1.5">1.5 mm</quantity>
|
||
.
|
||
<typeStatus id="5628881BEB4CCA4039D4FF6BB702FF55" box="[151,260,151,176]" pageId="4" pageNumber="157" type="holotype">Holotype</typeStatus>
|
||
: “5953-01”
|
||
<quantity id="4E6B9B5CEB4CCA4038C8FF64B7E4FF4A" box="[395,482,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="157" unit="mm" value="2.8">2.8 mm</quantity>
|
||
long excluding head, and
|
||
<quantity id="4E6B9B5CEB4CCA403A4AFF6BB559FF4A" box="[777,863,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="157" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide;
|
||
<typeStatus id="5628881BEB4CCA403AEFFF64B217FF55" box="[940,1041,152,176]" pageId="4" pageNumber="157" type="paratype">paratype</typeStatus>
|
||
: “5953-02”
|
||
<quantity id="4E6B9B5CEB4CCA403DDBFF64B2E8FF4A" box="[1176,1262,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="157" unit="mm" value="2.8">2.8 mm</quantity>
|
||
long excluding head, and
|
||
<quantity id="4E6B9B5CEB4CCA403848FF40B764FF31" box="[267,354,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="157" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide; additional specimens: “5953-03”
|
||
<quantity id="4E6B9B5CEB4CCA403A6AFF40B579FF31" box="[809,895,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="157" unit="mm" value="2.9">2.9 mm</quantity>
|
||
long excluding head, and
|
||
<quantity id="4E6B9B5CEB4CCA403DE8FF40B307FF31" box="[1195,1281,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="157" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide; “5953- 04” not measured because of poor preservation.
|
||
</paragraph>
|
||
<paragraph id="892C36B9EB4CCA403984FEF9B5ACFE80" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="157">
|
||
<typeStatus id="5628881BEB4CCA403984FEF9B72FFEF8" box="[199,297,261,285]" pageId="4" pageNumber="157" type="paratype">Paratype</typeStatus>
|
||
and two additional specimens slightly differs in body length; otherwise apparently similar in all visible morphological characters to the
|
||
<typeStatus id="5628881BEB4CCA403B11FEDBB4BEFEA5" box="[594,696,295,320]" pageId="4" pageNumber="157" type="holotype">holotype</typeStatus>
|
||
. Two additional specimens are not sufficiently visible or damaged (see material description) therefore they not included in the
|
||
<typeStatus id="5628881BEB4CCA403A75FEB1B560FE80" box="[822,870,333,357]" pageId="4" pageNumber="157">type</typeStatus>
|
||
serie.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1896532EB4CCA403984FE93B7D1FE48" pageId="4" pageNumber="157" type="etymology">
|
||
<paragraph id="892C36B9EB4CCA403984FE93B7D1FE48" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="157">
|
||
<emphasis id="BBE7EAABEB4CCA403984FE93B748FE6D" bold="true" box="[199,334,367,392]" pageId="4" pageNumber="157">Etymology.</emphasis>
|
||
The specific epithet is a patronym dedicated to Mr. Jonas Damzen (
|
||
<collectingRegion id="4B57F85BEB4CCA403DC8FE93B2E6FE6D" box="[1163,1248,367,392]" country="Lithuania" name="Vilniaus" pageId="4" pageNumber="157">Vilnius</collectingRegion>
|
||
,
|
||
<collectingCountry id="F1847629EB4CCA403DB2FE93B367FE6D" box="[1265,1377,367,392]" name="Lithuania" pageId="4" pageNumber="157">Lithuania</collectingCountry>
|
||
), the collector of the amber piece.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DDEC6631EB4CCA4039D4F9ADB726F921" httpUri="https://zenodo.org/record/1301314/files/figure.png" pageId="4" pageNumber="157" startId="4.[151,264,1617,1640]" targetBox="[384,1202,481,1597]" targetPageId="4">
|
||
<paragraph id="892C36B9EB4CCA4039D4F9ADB726F921" blockId="4.[151,1436,1617,1732]" pageId="4" pageNumber="157">
|
||
<emphasis id="BBE7EAABEB4CCA4039D4F9ADB73BF982" bold="true" box="[151,317,1617,1640]" pageId="4" pageNumber="157">FIGURES 5–7.</emphasis>
|
||
<taxonomicName id="4E934D3AEB4CCA403806F9AFB4C8F98D" authority="Bukejs & Alekseev" authorityName="Bukejs & Alekseev" authorityYear="2018" box="[325,718,1618,1640]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
|
||
<emphasis id="BBE7EAABEB4CCA403806F9AFB7FCF98D" box="[325,506,1618,1640]" italics="true" pageId="4" pageNumber="157">Ataenius damzeni</emphasis>
|
||
Bukejs & Alekseev
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BBE7EAABEB4CCA403B99F9AEB559F98D" bold="true" box="[730,863,1618,1640]" pageId="4" pageNumber="157">
|
||
<taxonomicNameLabel id="A0D457D0EB4CCA403B99F9AEB55DF98D" box="[730,859,1618,1640]" pageId="4" pageNumber="157" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</emphasis>
|
||
5, general view of amber piece “5953” with specimen enumeration (1—holotype; 2—paratype; 3‒4—non-type specimens); 6, details of pronotum and head, dorsofrontal view (holotype “5953-01”); 7, details of abdomen and hind left leg (holotype “5953-01”). Scale bar = 5 mm for Fig. 5; 0.25 mm for Figs. 6 and 7.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C1896532EB4CCA413984F90CB76BFE49" lastPageId="5" lastPageNumber="158" pageId="4" pageNumber="157" type="diagnosis">
|
||
<paragraph id="892C36B9EB4CCA403984F90CB353F804" blockId="4.[151,1437,1776,2017]" pageId="4" pageNumber="157">
|
||
<emphasis id="BBE7EAABEB4CCA403984F90CB7D7F8EC" bold="true" box="[199,465,1776,1801]" pageId="4" pageNumber="157">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4E934D3AEB4CCA40389FF90DB4A0F8EC" authority="Bukejs & Alekseev, 2018" authorityName="Bukejs & Alekseev" authorityYear="2018" box="[476,678,1776,1801]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
|
||
<emphasis id="BBE7EAABEB4CCA40389FF90DB4A0F8EC" box="[476,678,1776,1801]" italics="true" pageId="4" pageNumber="157">Ataenius damzeni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A0D457D0EB4CCA403BFBF90DB545F8EC" box="[696,835,1776,1801]" pageId="4" pageNumber="157" rank="species">new species</taxonomicNameLabel>
|
||
differs from scarabaeids previously described from Baltic amber (
|
||
<taxonomicName id="4E934D3AEB4CCA40387AF8E9B411F8C8" authorityName="Quiel" authorityYear="1910" box="[313,535,1813,1837]" class="Insecta" family="Scarabaeidae" genus="Ataenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="europaeus">
|
||
<emphasis id="BBE7EAABEB4CCA40387AF8E9B411F8C8" box="[313,535,1813,1837]" italics="true" pageId="4" pageNumber="157">Ataenius europaeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E934D3AEB4CCA403B66F8E9B4F6F8C8" baseAuthorityName="Zang" baseAuthorityYear="1905" box="[549,752,1813,1837]" class="Insecta" family="Scarabaeidae" genus="Saprosites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="succini">
|
||
<emphasis id="BBE7EAABEB4CCA403B66F8E9B4F6F8C8" box="[549,752,1813,1837]" italics="true" pageId="4" pageNumber="157">Saprosites succini</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4E934D3AEB4CCA403A73F8E9B20AF8C8" authorityName="Tamutis, Alekseev & Bukejs" authorityYear="2017" box="[816,1036,1812,1837]" class="Insecta" family="Scarabaeidae" genus="Airapus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="lithuanicus">
|
||
<emphasis id="BBE7EAABEB4CCA403A73F8E9B20AF8C8" box="[816,1036,1812,1837]" italics="true" pageId="4" pageNumber="157">Airapus lithuanicus</emphasis>
|
||
</taxonomicName>
|
||
) in possessing: (1) glabrous elytral interstriae without granulation and setation; (2) metatarsomere 1 slightly shorter than metatarsomeres 2–5 combined; (3) meso- and metatibiae with two terminal spurs of unequal length; (4) anterior portion of head with sparse and very fine punctation; (5) pronotal base narrowly bordered; (6) pronotal punctation distinctly unequal in size, with dense and small punctures anteriorly, and larger and sparser punctures posteriorly; (7) pronotal lateral margins crenulate, without setae or scales; and (8) smaller body size. See also details provided in key below.
|
||
</paragraph>
|
||
<paragraph id="892C36B9EB4DCA413984FF6BB76BFE49" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="158">
|
||
This new fossil species differs from extant congeners of the genus (
|
||
<bibRefCitation id="ED024B48EB4DCA413A93FF6BB289FF55" box="[976,1167,151,176]" pageId="5" pageNumber="158" refId="ref4099" refString="Cartwright, O. L. (1974) Ataenius, Aphotaenius and Pseudataenius of the United States and Canada. Smithsonian Contributions to Zoology, 154, 1 - 106." type="journal article">Cartwright 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="ED024B48EB4DCA413DDEFF6BB348FF55" box="[1181,1358,151,176]" pageId="5" pageNumber="158" refId="ref4697" refString="Stebnicka, Z. T. (2001) Aphodiinae (Insecta: Coleoptera: Scarabaeidae). Fauna of New Zealand. Vol. 42. Manaaki Whenua Press, Landcare Research, Lincoln, Canterbury, 64 pp." type="book">Stebnicka 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="ED024B48EB4DCA413C1FFF64B391FF55" box="[1372,1431,152,176]" pageId="5" pageNumber="158" refId="ref4738" refString="Stebnicka, A. T. (2003) The New World species of Ataenius Harold, 1867. III. Revision of the A. imbricatus - group sensu lato (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, 46 (3), 219 - 249." type="journal article">2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="ED024B48EB4DCA4139D4FF40B791FF31" box="[151,407,188,213]" pageId="5" pageNumber="158" refId="ref4610" refString="Smith, A. B. T. & Skelley, P. E. (2007) A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa, 1458 (1), 1 - 80. https: // doi. org / 10.11646 / zootaxa. 1458.1.1" type="journal article">Smith & Skelley 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="ED024B48EB4DCA4138E0FF40B4B5FF30" box="[419,691,188,213]" pageId="5" pageNumber="158" refId="ref4888" refString="Stebnicka, Z. T. & Lago, P. K. (2005) The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi, 19 (1 - 2), 55 - 83." type="journal article">Stebnicka & Lago 2005</bibRefCitation>
|
||
) in the combination of the following characters: (1) body without argillaceous coating; (2) clypeus broadly rounded each side of shallow median emargination (without trace of clypeal denticles); (3) head finely punctate; (4) pronotal punctation heterogeneous; (5) pronotal disc evenly convex; (6) lateral pronotal margins slightly upturned, narrowly bordered, crenulate, without setae or scales; (7) scutellar shield oblong; (8) mesosternum carinate between coxae; (9) elytra subparallel-sided; (10) elytral interstrial intervals convex, equal in height; (11) body ventral surface shiny; and (12) abdominal ventrites 2–5 distinctly punctate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |