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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1046.57052" ID-GBIF-Dataset="348e46a4-8819-4e3c-b4b9-b5e88bee10d7" ID-PMC="PMC8238928" ID-Pensoft-Pub="1313-2970-1046-1" ID-Pensoft-UUID="DBC42C98E4DA5074B86525CF2FB2FA64" ID-PubMed="34239337" ID-ZooBank="66E61C4EFAFE45DE9145DB38199BDEC3" ModsDocID="1313-2970-1046-1" checkinTime="1624307367557" checkinUser="pensoft" docAuthor="Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott &amp; Meyer, Marc De" docDate="2021" docId="2B80D8D091985DA382C8F97213F40741" docLanguage="en" docName="ZooKeys 1046: 1-141" docOrigin="ZooKeys 1046" docPubDate="2021-06-21" docSource="http://dx.doi.org/10.3897/zookeys.1046.57052" docTitle="Mesembrius simplicipes Curran 1929" docType="treatment" docVersion="4" id="DBC42C98E4DA5074B86525CF2FB2FA64" lastPageNumber="1" masterDocId="DBC42C98E4DA5074B86525CF2FB2FA64" masterDocTitle="Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data" masterLastPageNumber="141" masterPageNumber="1" pageNumber="1" updateTime="1668150539886" updateUser="ExternalLinkService">
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<mods:title>Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jordaens, Kurt</mods:namePart>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
<mods:nameIdentifier type="email">kurt.jordaens@africamuseum.be</mods:nameIdentifier>
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<mods:namePart>Goergen, Georg</mods:namePart>
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<mods:affiliation>International Institute for Tropical Agriculture (IITA), Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Skevington, Jeffrey H.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-9870</mods:nameIdentifier>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kelso, Scott</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Meyer, Marc De</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0755-2898</mods:nameIdentifier>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-21</mods:number>
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<mods:number>1046</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1046.57052</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1046.57052</mods:identifier>
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<treatment ID-GBIF-Taxon="183114815" LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741" httpUri="http://treatment.plazi.org/id/2B80D8D091985DA382C8F97213F40741" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="2B80D8D0-9198-5DA3-82C8-F97213F40741" authority="Curran, 1929" authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">Mesembrius simplicipes Curran, 1929</taxonomicName>
<figureCitation captionStart="Figures 17, 18" captionStartId="F9" captionText="Figures 17, 18. Mesembrius spp., habitus, lateral view 17 M. platytarsis Curran syn. nov. (♂) 18 M. regulus (Hull) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures17-18" httpUri="https://binary.pensoft.net/fig/557470" pageId="0" pageNumber="1">Figs 17</figureCitation>
<figureCitation captionStart="Figures 21, 22" captionStartId="F11" captionText="Figures 21, 22. 21 Mesembrius spp., habitus, lateral view. M. simplicipes Curran (♂) 22 M. strigilatus (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures21-22" httpUri="https://binary.pensoft.net/fig/557472" pageId="0" pageNumber="1">, 21</figureCitation>
<figureCitation captionStart="Figures 37, 38" captionStartId="F19" captionText="Figures 37, 38. Mesembrius spp., habitus, lateral view 37 M. platytarsis syn. nov. Curran (♀) 38 M. regulus (Hull) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures37-38" httpUri="https://binary.pensoft.net/fig/557480" pageId="0" pageNumber="1">, 37</figureCitation>
<figureCitation captionStart="Figures 41, 42" captionStartId="F21" captionText="Figures 41, 42. Mesembrius spp., habitus, lateral view 41 M. simplicipes Curran (♀) 42 M. strigilatus (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures41-42" httpUri="https://binary.pensoft.net/fig/557482" pageId="0" pageNumber="1">, 41</figureCitation>
<figureCitation captionStart="Figures 5863" captionStartId="F26" captionText="Figures 58 - 63. Mesembrius spp., head, frontal view 58 M. nigriceps Curran (♂) 59 M. perforatus (Speiser) (♂) 60 M. platytarsis Curran syn. nov. (♂) 61 M. regulus (Hull) (♂) 62 M. rex Curran (♂) 63 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures58-63" httpUri="https://binary.pensoft.net/fig/557487" pageId="0" pageNumber="1">, 60</figureCitation>
<figureCitation captionStart="Figures 6469" captionStartId="F27" captionText="Figures 64 - 69. Mesembrius spp., head, frontal view 64 M. simplicipes Curran (♂) 65 M. strigilatus (Bezzi) (♂) 66 M. sulcus sp. nov. (♂) 67 M. tarsatus (Bigot) (♂) 68 M. tibialis sp. nov. (♂) 69 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures64-69" httpUri="https://binary.pensoft.net/fig/557488" pageId="0" pageNumber="1">, 64</figureCitation>
<figureCitation captionStart="Figures 7578" captionStartId="F29" captionText="Figures 75 - 78. Mesembrius spp., head, frontal view 75 M. platytarsis Curran syn. nov. (♀) 76 M. regulus (Hull) (♂) 77 M. rex Curran (♂) 78 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures75-78" httpUri="https://binary.pensoft.net/fig/557490" pageId="0" pageNumber="1">, 75</figureCitation>
<figureCitation captionStart="Figures 7982" captionStartId="F30" captionText="Figures 79 - 82. Mesembrius spp., head, frontal view 79 M. simplicipes Curran (♂) 80 M. sulcus sp. nov. (♀) 81 M. tarsatus (Bigot) (♀) 82 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures79-82" httpUri="https://binary.pensoft.net/fig/557491" pageId="0" pageNumber="1">, 79</figureCitation>
<figureCitation captionStart="Figures 95100" captionStartId="F33" captionText="Figures 95 - 100. Mesembrius spp., abdomen, dorsal view 95 M. nigriceps Curran (♂) 96 M. perforatus (Speiser) (♂) 97 M. platytarsis Curran syn. nov. (♂) 98 M. regulus (Hull) (♂) 99 M. rex Curran (♂) 100 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures95-100" httpUri="https://binary.pensoft.net/fig/557494" pageId="0" pageNumber="1">, 97</figureCitation>
<figureCitation captionStart="Figures 101106" captionStartId="F34" captionText="Figures 101 - 106. Mesembrius spp., abdomen, dorsal view 101 M. simplicipes Curran (♂) 102 M. strigilatus (Bezzi) (♂) 103 M. sulcus sp. nov. (♂) 104 M. tarsatus (Bigot) (♂) 105 M. tibialis sp. nov. (♂) 106 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures101-106" httpUri="https://binary.pensoft.net/fig/557495" pageId="0" pageNumber="1">, 101</figureCitation>
<figureCitation captionStart="Figures 113118" captionStartId="F36" captionText="Figures 113 - 118. Mesembrius spp., abdomen, dorsal view 113 M. cyanipennis (Bezzi) (♀) 114 M. maculifer Hull (♀) 115 M. madagascariensis Keiser (♀) 116 M. minor (Bezzi) (♀) 117 M. morio (Bezzi) (♀) 118 M. platytarsis Curran syn. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures113-118" httpUri="https://binary.pensoft.net/fig/557497" pageId="0" pageNumber="1">, 118</figureCitation>
<figureCitation captionStart="Figures 119123" captionStartId="F37" captionText="Figures 119 - 123. Mesembrius spp., abdomen, dorsal view 119 M. regulus (Hull) (♀) 120 M. rex Curran (♀) 121 M. senegalensis (Macquart) (♀) 122 M. simplicipes Curran (♀) 123 M. strigilatus (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures119-123" httpUri="https://binary.pensoft.net/fig/557498" pageId="0" pageNumber="1">, 122</figureCitation>
<figureCitation captionStart="Figures 139142" captionStartId="F42" captionText="Figures 139 - 142. Mesembrius spp., right wing 139 M. nigriceps Curran (♂) 140 M. perforatus (Speiser) (♂) 141 M. platytarsis Curran syn. nov. (♂) 142 M. regulus (Hull) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures139-142" httpUri="https://binary.pensoft.net/fig/557503" pageId="0" pageNumber="1">, 141</figureCitation>
<figureCitation captionStart="Figures 143146" captionStartId="F43" captionText="Figures 143 - 146. Mesembrius spp., right wing 143 M. rex Curran (♂) 144 M. senegalensis (Macquart) (♂) 145 M. simplicipes Curran (♂) 146 M. strigilatus (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures143-146" httpUri="https://binary.pensoft.net/fig/557504" pageId="0" pageNumber="1">, 145</figureCitation>
<figureCitation captionStart="Figures 166171" captionStartId="F49" captionText="Figures 166 - 171. Mesembrius spp., proleg, dorsal view 166 M. simplicipes Curran (♂) 167 M. platytarsis Curran syn. nov. (♂) 168 M. tarsatus (Bigot) (♀) 169 M. chapini Curran (♀) 170 M. rex Curran (♀) 171 M. regulus (Hull) (♀). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures166-171" httpUri="https://binary.pensoft.net/fig/557509" pageId="0" pageNumber="1">, 166-167</figureCitation>
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">, 193</figureCitation>
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">, 194</figureCitation>
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">, 219</figureCitation>
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">, 223</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">Mesembrius simplicipes</taxonomicName>
Curran, 1929: 500.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">Mesembrius simplicipes</taxonomicName>
-
<bibRefCitation author="Keiser, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" pagination="223 - 318" refId="B34" refString="Keiser, F, 1971. Syrphidae von Madagascar (Dipt.). Verhandlungen der Naturforschenden Gesellschaft in Basel 81: 223 - 318" title="Syrphidae von Madagascar (Dipt.)." volume="81" year="1971">Keiser (1971)</bibRefCitation>
: 266 -
<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" pagination="488 - 510" refId="B49" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth (1980)</bibRefCitation>
: 504.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius platytarsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="platytarsis">Mesembrius platytarsis</taxonomicName>
Curran, 1929: 501. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius platytarsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="platytarsis">Mesembrius platytarsis</taxonomicName>
-
<bibRefCitation author="Hull, FM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="1" pagination="309 - 334" publicationUrl="https://www.jstor.org/stable/pdf/4064312.pdf" refId="B30" refString="Hull, FM, 1941. A study of syrphid flies from Madagascar (Diptera). Proceedings of the Academy of Natural Sciences of Philadelphia 92 (1940): 309 - 334, https://www.jstor.org/stable/pdf/4064312.pdf" title="A study of syrphid flies from Madagascar (Diptera)." url="https://www.jstor.org/stable/pdf/4064312.pdf" volume="92" year="1941">Hull (1941)</bibRefCitation>
: 330 -
<bibRefCitation author="Keiser, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" pagination="223 - 318" refId="B34" refString="Keiser, F, 1971. Syrphidae von Madagascar (Dipt.). Verhandlungen der Naturforschenden Gesellschaft in Basel 81: 223 - 318" title="Syrphidae von Madagascar (Dipt.)." volume="81" year="1971">Keiser (1971)</bibRefCitation>
: 265 -
<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" pagination="488 - 510" refId="B49" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth (1980)</bibRefCitation>
: 504.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius simplicipes</emphasis>
</taxonomicName>
males lack an apical pile brush on the profemur which is dorsoventrally flattened. The pro- and mesolegs are orange and with a darker area on the dorsal side of the femur. The probasitarsus is laterally expanded and has some long orange pile. The large yellow maculae on the abdomen lack short, black spines on the posterior edges. Scutum and scutellum are entirely yellow pilose. The male of this species cannot be confused with any other species by the lateral lobe on the probasitarsus. Females have a nearly black abdomen with a pair of vague lateral maculae on tergites II and III. The female of
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
can be distinguished from any other species (except from
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
) by the nearly black abdomen (clearly yellow to orange and black in other species). It can be distinguished from
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
by the pro- and mesolegs which are reddish-brown (extensively brown and black in
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Examined material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius simplicipes</emphasis>
</taxonomicName>
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Madagascar" location="Madagascar" specimenCount="1" typeStatus="Holotype">
Curran:
<typeStatus>Holotype</typeStatus>
, male: &quot;
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Mesembrius</taxonomicName>
// TYPE //
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">Mesembrius simplicipes</taxonomicName>
// Curran. // No.&quot; &quot;
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Mesembrius</taxonomicName>
//
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius simplicipes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplicipes">Mesembrius simplicipes</taxonomicName>
// Curran&quot; &quot;
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
// Great Oriental // Forest&quot; &quot;California Academy // of Sciences // Type No. 11230&quot; [CAS] [date and collector unknown; type studied from pictures].
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius platytarsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius platytarsis</emphasis>
</taxonomicName>
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Madagascar" location="Madagascar" specimenCount="1" typeStatus="Holotype">
Curran:
<typeStatus>Holotype</typeStatus>
, male: &quot;
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Mesembrius</taxonomicName>
// TYPE //
<taxonomicName authorityName="Curran" authorityYear="1929" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius platytarsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="platytarsis">Mesembrius platytarsis</taxonomicName>
// Curran. // No.&quot; &quot;
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
// Great Oriental // Forest&quot; &quot;California Academy // of Sciences // Type No. 11229&quot; [CAS] [date and collector unknown; type studied from pictures].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1957-12-24" collectorName="B. R. Stuckenberg" country="Madagascar" location="Station Agric." municipality="Alaotra" specimenCount="2" specimenCount-female="2">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingMunicipality>Alaotra</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:70989C7DA3E55AD8595363BB148413AD" country="Madagascar" municipality="Alaotra" name="Station Agric.">Station Agric.</location>
;
<collectingDate value="1957-12-24">24 Dec 1957</collectingDate>
;
<collectorName>B.R. Stuckenberg</collectorName>
leg.; NMSA
</materialsCitation>
<materialsCitation collectingDate="1958-03-30" collectorName="F. Keiser" country="Madagascar" location="Analvony" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:444D797688650B9D2559F9455DC1152F" country="Madagascar" name="Analvony">Analvony</location>
;
<collectingDate value="1958-03-30">30 Mar 1958</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1952-11" collectorName="E. S. Brown" country="Madagascar" location="Antananarivo" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1952-11">Nov 1952</collectingDate>
;
<collectorName>E.S. Brown</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation collectingDate="1957-10-18" collectorName="F. Keiser" country="Madagascar" location="Antananarivo" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1957-10-18">18 Oct 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1957-12-13" collectorName="F. Keiser" country="Madagascar" location="Antananarivo" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1957-12-13">13 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1957-12-14" collectorName="F. Keiser" country="Madagascar" location="Antananarivo" specimenCount="2" specimenCount-female="2" stateProvince="Antananarivo">
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1957-12-14">14 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1958-09-06" collectorName="F. Keiser" country="Madagascar" location="Antananarivo" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1958-09-06">6 Sep 1958</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1970-02-08" collectorName="L. and R. Blommers" country="Madagascar" location="Antananarivo" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1970-02-08">8 Feb 1970</collectingDate>
;
<collectorName>L. and R. Blommers</collectorName>
leg.; RMNH
</materialsCitation>
<materialsCitation collectingDate="1959-03-25" collectorName="F. Keiser" country="Madagascar" location="Lake Kavitaha" municipality="Ampefy" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<collectingMunicipality>Ampefy</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:AF199A1ADFD9C3AD30169770C0F810DE" country="Madagascar" municipality="Ampefy" name="Lake Kavitaha" stateProvince="Antananarivo">Lake Kavitaha</location>
;
<collectingDate value="1959-03-25">25 Mar 1959</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1959-03-29" collectorName="F. Keiser" country="Madagascar" location="Lake Kavitaha" municipality="Ampefy" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<collectingMunicipality>Ampefy</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:3820C79C792426E0C1E3085841C60C4C" country="Madagascar" municipality="Ampefy" name="Lake Kavitaha" stateProvince="Antananarivo">Lake Kavitaha</location>
;
<collectingDate value="1959-03-29">29 Mar 1959</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1968-02-02" collectorName="J. W. Boyes" country="Madagascar" location="Parc Tsimbazaza" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:B8967DE965F365EAB1D77DD0BBF2FE0D" country="Madagascar" name="Parc Tsimbazaza" stateProvince="Antananarivo">Parc Tsimbazaza</location>
;
<collectingDate value="1968-02-02">2 Feb 1968</collectingDate>
;
<collectorName>J.W. Boyes</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation collectingDate="1957-12-14" collectorName="F. Keiser" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:48EA7F256F210900DB0E7F85B55A5D85" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1957-12-14">14 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
; NMB
</materialsCitation>
<materialsCitation collectingDate="1993-10-15" collectingDateMax="1993-10-22" collectingDateMin="1993-10-15" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:0B1F32F0C992FE14073956981699A121" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-15" valueMax="1993-10-22" valueMin="1993-10-15">15-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation collectingDate="1993-10-16" collectingDateMax="1993-10-22" collectingDateMin="1993-10-16" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:4C12CC73B0A1D7231DBE6712B2B1EEB6" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-16" valueMax="1993-10-22" valueMin="1993-10-16">16-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation collectingDate="1993-10-16" collectingDateMax="1993-10-22" collectingDateMin="1993-10-16" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="2" specimenCount-male="2" stateProvince="Antananarivo">
<specimenCount type="male">2♂♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:70455C5A61816F386944B25101790458" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-16" valueMax="1993-10-22" valueMin="1993-10-16">16-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="1993-10-16" collectingDateMax="1993-10-22" collectingDateMin="1993-10-16" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:1C08B374075953402123149ADAB2EC47" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-16" valueMax="1993-10-22" valueMin="1993-10-16">16-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; NMK
</materialsCitation>
<materialsCitation collectingDate="1993-10-26" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:FF534678DF1B3667780995EA40C6929C" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-26">26 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation collectingDate="1993-10-26" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:04647918F3C508C5A1CB41F7165C043E" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-10-26">26 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="1993-11-06" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:567B89891EF91FFC17BFF828D9FCD743" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-11-06">6 Nov 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation collectingDate="1993-11-06" collectorName="C. Kassebeer" country="Madagascar" location="Park Tsimbazaza" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:5C1C9EC8AA616CB32D43417EF7303C75" country="Madagascar" name="Park Tsimbazaza" stateProvince="Antananarivo">Park Tsimbazaza</location>
;
<collectingDate value="1993-11-06">6 Nov 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation collectingDate="1957-09-30" collectorName="F. Keiser" country="Madagascar" location="Perinet" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:EEADAD48665D6B24D84E3F96F8D0E09F" country="Madagascar" name="Perinet" stateProvince="Antananarivo">Perinet</location>
;
<collectingDate value="1957-09-30">30 Sep 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Madagascar" location="Nosivola" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:2B80D8D091985DA382C8F97213F40741:CF6AFEDB4AD058156C333559EC7813BE" country="Madagascar" name="Nosivola">Nosivola</location>
; date and collector unknown; RMNH
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Re-description male</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation captionStart="Figures 17, 18" captionStartId="F9" captionText="Figures 17, 18. Mesembrius spp., habitus, lateral view 17 M. platytarsis Curran syn. nov. (♂) 18 M. regulus (Hull) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures17-18" httpUri="https://binary.pensoft.net/fig/557470" pageId="0" pageNumber="1">17</figureCitation>
,
<figureCitation captionStart="Figures 21, 22" captionStartId="F11" captionText="Figures 21, 22. 21 Mesembrius spp., habitus, lateral view. M. simplicipes Curran (♂) 22 M. strigilatus (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures21-22" httpUri="https://binary.pensoft.net/fig/557472" pageId="0" pageNumber="1">21</figureCitation>
).
</emphasis>
Body length: 12.7-13.4 mm. Wing length: 8.8-10.4 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
(Figs
<figureCitation captionStart="Figures 5863" captionStartId="F26" captionText="Figures 58 - 63. Mesembrius spp., head, frontal view 58 M. nigriceps Curran (♂) 59 M. perforatus (Speiser) (♂) 60 M. platytarsis Curran syn. nov. (♂) 61 M. regulus (Hull) (♂) 62 M. rex Curran (♂) 63 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures58-63" httpUri="https://binary.pensoft.net/fig/557487" pageId="0" pageNumber="1">60</figureCitation>
,
<figureCitation captionStart="Figures 6469" captionStartId="F27" captionText="Figures 64 - 69. Mesembrius spp., head, frontal view 64 M. simplicipes Curran (♂) 65 M. strigilatus (Bezzi) (♂) 66 M. sulcus sp. nov. (♂) 67 M. tarsatus (Bigot) (♂) 68 M. tibialis sp. nov. (♂) 69 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures64-69" httpUri="https://binary.pensoft.net/fig/557488" pageId="0" pageNumber="1">64</figureCitation>
). Eyes bare; holoptic, eye contiguity as long as length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black; black pilose; strongly yellow pollinose before anterior ocellus. Lateral ocelli nearly touching eye margin. Occiput black, yellow pilose with a few very short, stiff black setulae near dorsal eye margin; strongly white pollinose. Frontal triangle black; black pilose; strongly yellow pollinose. Frontal prominence shiny reddish-black. Antenna, scape and pedicel very dark reddish-black; postpedicel black; antennal arista reddish-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Scutum black with dorsally a pair of yellow pollinose vittae which are less well demarcated posteriorly; with a lateral yellow pollinose vitta; yellow pilose. Scutellum uniformly yellow-brown; yellow pilose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
Proleg (Figs
<figureCitation captionStart="Figures 166171" captionStartId="F49" captionText="Figures 166 - 171. Mesembrius spp., proleg, dorsal view 166 M. simplicipes Curran (♂) 167 M. platytarsis Curran syn. nov. (♂) 168 M. tarsatus (Bigot) (♀) 169 M. chapini Curran (♀) 170 M. rex Curran (♀) 171 M. regulus (Hull) (♀). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures166-171" httpUri="https://binary.pensoft.net/fig/557509" pageId="0" pageNumber="1">166</figureCitation>
,
<figureCitation captionStart="Figures 166171" captionStartId="F49" captionText="Figures 166 - 171. Mesembrius spp., proleg, dorsal view 166 M. simplicipes Curran (♂) 167 M. platytarsis Curran syn. nov. (♂) 168 M. tarsatus (Bigot) (♀) 169 M. chapini Curran (♀) 170 M. rex Curran (♀) 171 M. regulus (Hull) (♀). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures166-171" httpUri="https://binary.pensoft.net/fig/557509" pageId="0" pageNumber="1">167</figureCitation>
): Femur dorsoventrally flattened; dorsally brown, except proximal 1/5 and distal end; otherwise orange; yellow pile on posteroventral side longer than black pile on posterodorsal side; with a patch of very short black stiff spines on proximal ventral 1/5; pile entirely yellow anteriorly. Tibia orange; with long orange pile, but pile black on ventral distal end. Basitarsus whitish; laterally expanded; with a tuft of orange pile on the expansion; ventrally with some thick stiff pile on the expansion. Other tarsi orange; with orange and black short pile. Mesoleg: Femur as proleg, but without the posterodorsal black pile. Tibia orange; yellow pilose, with some short stiff black pile on distal end ventrally. Tarsi orange; with short black pile and some short stiff black pile ventrally. Metaleg (Figs
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">193</figureCitation>
,
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">194</figureCitation>
): Femur chocolate-brown, distal end orange; with loose yellow pile on proximal 1/2 to 2/3 and mostly black pilose on distal 1/3 to 1/2. Tibia orange; with short black pile. Metabasitarsus either deeply excavated anteriorly at proximal end and with a lobe (Fig.
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">193</figureCitation>
) or unmodified (Fig.
<figureCitation captionStart="Figures 193196" captionStartId="F55" captionText="Figures 193 - 196. Mesembrius spp., metaleg, dorsal view 193 M. platytarsis Curran syn. nov. (♂) 194 M. simplicipes Curran (♂). Metaleg, anterior view 195 M. caffer (Loew) (nominal morph) (♂) 196 M. caffer (Loew) (spined morph) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures193-196" httpUri="https://binary.pensoft.net/fig/557515" pageId="0" pageNumber="1">194</figureCitation>
); orange; black pilose. Other tarsi orange; dorsally black pilose, ventrally orange pilose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
(Figs
<figureCitation captionStart="Figures 139142" captionStartId="F42" captionText="Figures 139 - 142. Mesembrius spp., right wing 139 M. nigriceps Curran (♂) 140 M. perforatus (Speiser) (♂) 141 M. platytarsis Curran syn. nov. (♂) 142 M. regulus (Hull) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures139-142" httpUri="https://binary.pensoft.net/fig/557503" pageId="0" pageNumber="1">141</figureCitation>
,
<figureCitation captionStart="Figures 143146" captionStartId="F43" captionText="Figures 143 - 146. Mesembrius spp., right wing 143 M. rex Curran (♂) 144 M. senegalensis (Macquart) (♂) 145 M. simplicipes Curran (♂) 146 M. strigilatus (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures143-146" httpUri="https://binary.pensoft.net/fig/557504" pageId="0" pageNumber="1">145</figureCitation>
). Entire wing uniformly very dense microtrichose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Figs
<figureCitation captionStart="Figures 95100" captionStartId="F33" captionText="Figures 95 - 100. Mesembrius spp., abdomen, dorsal view 95 M. nigriceps Curran (♂) 96 M. perforatus (Speiser) (♂) 97 M. platytarsis Curran syn. nov. (♂) 98 M. regulus (Hull) (♂) 99 M. rex Curran (♂) 100 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures95-100" httpUri="https://binary.pensoft.net/fig/557494" pageId="0" pageNumber="1">97</figureCitation>
,
<figureCitation captionStart="Figures 101106" captionStartId="F34" captionText="Figures 101 - 106. Mesembrius spp., abdomen, dorsal view 101 M. simplicipes Curran (♂) 102 M. strigilatus (Bezzi) (♂) 103 M. sulcus sp. nov. (♂) 104 M. tarsatus (Bigot) (♂) 105 M. tibialis sp. nov. (♂) 106 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures101-106" httpUri="https://binary.pensoft.net/fig/557495" pageId="0" pageNumber="1">101</figureCitation>
). Tergite II with a pair of very large, yellow almost square maculae which expand into the anterolateral corners; black markings hourglass-shaped, posterior part reaching the lateral sides of the tergite; posterior part of black marking with some black pile in the centre which do not extend to the lateral tergite sides; otherwise yellow pilose. Tergite III with a yellow fascia which is almost interrupted by the medial broad black marking; the latter strongly white pollinose anteriorly; with a medial area of white pollinosity; posterior part of black marking with sparse black pile in the centre; yellow pilose otherwise. Tergite IV strongly white pollinose, except for a black medial area and a darker posterior border; yellow pilose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
(Figs
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">219</figureCitation>
,
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">223</figureCitation>
). Epandrium: Dorsal lobe of surstylus short, bent (as a boomerang); irregularly covered with short black spines which are denser at distal and proximal end and at dorsal bend. Ventral lobe of surstylus straight; bare.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Re-description female</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation captionStart="Figures 37, 38" captionStartId="F19" captionText="Figures 37, 38. Mesembrius spp., habitus, lateral view 37 M. platytarsis syn. nov. Curran (♀) 38 M. regulus (Hull) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures37-38" httpUri="https://binary.pensoft.net/fig/557480" pageId="0" pageNumber="1">37</figureCitation>
,
<figureCitation captionStart="Figures 41, 42" captionStartId="F21" captionText="Figures 41, 42. Mesembrius spp., habitus, lateral view 41 M. simplicipes Curran (♀) 42 M. strigilatus (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures41-42" httpUri="https://binary.pensoft.net/fig/557482" pageId="0" pageNumber="1">41</figureCitation>
).
</emphasis>
Body length: 14.2-14.9 mm. Wing length: 10.0-10.6 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
(Figs
<figureCitation captionStart="Figures 7578" captionStartId="F29" captionText="Figures 75 - 78. Mesembrius spp., head, frontal view 75 M. platytarsis Curran syn. nov. (♀) 76 M. regulus (Hull) (♂) 77 M. rex Curran (♂) 78 M. senegalensis (Macquart) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures75-78" httpUri="https://binary.pensoft.net/fig/557490" pageId="0" pageNumber="1">75</figureCitation>
,
<figureCitation captionStart="Figures 7982" captionStartId="F30" captionText="Figures 79 - 82. Mesembrius spp., head, frontal view 79 M. simplicipes Curran (♂) 80 M. sulcus sp. nov. (♀) 81 M. tarsatus (Bigot) (♀) 82 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures79-82" httpUri="https://binary.pensoft.net/fig/557491" pageId="0" pageNumber="1">79</figureCitation>
). Eyes bare; dichoptic. Face yellow-white with dark medial vitta; white pilose, white pollinose. Frons black in dorsal half and medial part of ventral half, yellow-white on lateral parts of ventral half; black pilose on black parts, white pilose on yellow-white parts. Distance between lateral ocellus and eye margin approx. width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black, orange-brown at distal end; scape black; pedicel orange-brown; postpedicel black; antennal arista reddish-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Scutum dark brown with one pair of dorsolateral yellow-white pollinose vittae which are vaguely connected posteriorly; with lateral, yellow-white pollinose vitta; short yellow pilose on anterior half, short yellow and black pilose on posterior half. Scutellum yellow-orange; yellow pilose with shorter, black pile interspersed in posterior half.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
Proleg: Femur orange brown with darker, central area on dorsal side; yellow-white pilose with black pile on posterodorsal side and posteroventral distal half. Tibia orange, slightly darkened on ventral distal end; yellow-white pilose. Tarsi orange-brown; basitarsus and second tarsomere yellow-white pilose, other tarsi yellow-white and black pilose. Femur orange brown with darker, central area on dorsal side; yellow-white pilose with black pile on ventral distal half. Tibia orange-brown; yellow-white pilose, with some short, thick black pile at ventral distal end. Basitarsus orange; yellow-white pilose, with short, thick black pile on ventral side. Other tarsi orange-brown; yellow-white and black pilose; with short, thick black pile ventrally, except on most distal tarsomere. Metaleg: Femur dark brown to black, reddish-brown at distal end; yellow-white pilose with shorter and thicker black pile on anteroventral and ventral distal 1/2. Tibia orange-brown; yellow-white and black pilose. Tarsi orange-brown; black and yellow-white pilose dorsally, yellow-orange pilose ventrally.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing.</emphasis>
Entire wing uniformly microtrichose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Figs
<figureCitation captionStart="Figures 113118" captionStartId="F36" captionText="Figures 113 - 118. Mesembrius spp., abdomen, dorsal view 113 M. cyanipennis (Bezzi) (♀) 114 M. maculifer Hull (♀) 115 M. madagascariensis Keiser (♀) 116 M. minor (Bezzi) (♀) 117 M. morio (Bezzi) (♀) 118 M. platytarsis Curran syn. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures113-118" httpUri="https://binary.pensoft.net/fig/557497" pageId="0" pageNumber="1">118</figureCitation>
,
<figureCitation captionStart="Figures 119123" captionStartId="F37" captionText="Figures 119 - 123. Mesembrius spp., abdomen, dorsal view 119 M. regulus (Hull) (♀) 120 M. rex Curran (♀) 121 M. senegalensis (Macquart) (♀) 122 M. simplicipes Curran (♀) 123 M. strigilatus (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures119-123" httpUri="https://binary.pensoft.net/fig/557498" pageId="0" pageNumber="1">122</figureCitation>
). Tergite II with a pair of large orange maculae; with an anterior and posterior black marking which are connected with a parallel-sided central black marking which is 1/5 the tergite width; yellow pilose on maculae and on anterior and central black marking, black pilose on posterior black marking; white pollinose on posterior black marking. Tergite III with orange fascia (approx. 1/2 of tergal length laterally; approx. 1/8 in medially); with posterior large black marking; yellow pilose on most of the orange fascia, black pilose on black marking and central area of orange fascia; strongly white pollinose on posterior half and medial anterior part. Tergite IV similar, but without orange fascia. Tergite V orange-brown with darker lateral sides; yellow-white pilose; white pollinose in anterolateral corners.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Madagascar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
Morphologically, the species is similar to
<taxonomicName lsidName="M. platytarsis" pageId="0" pageNumber="1" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">M. platytarsis</emphasis>
</taxonomicName>
syn. nov. Males of both species differ in the presence (
<taxonomicName lsidName="M. platytarsis" pageId="0" pageNumber="1" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">M. platytarsis</emphasis>
</taxonomicName>
syn. nov.) or absence (
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
) of a large lobe on the anterior side of the metabasitarsus. Male genitalia are also very similar (compare Fig.
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">219</figureCitation>
with Fig.
<figureCitation captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">223</figureCitation>
). Females are morphologically similar as well and the supposed difference in the extent of black pile amongst the yellow pile on the frons is unreliable. The mean interspecific
<emphasis italics="true" pageId="0" pageNumber="1">p</emphasis>
-distance between both species is very low (0.02%) and of what is usually as observed within species. Both species have been described from the same locality, i.e. the Eastern Forest of Madagascar.
<bibRefCitation author="Keiser, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" pagination="223 - 318" refId="B34" refString="Keiser, F, 1971. Syrphidae von Madagascar (Dipt.). Verhandlungen der Naturforschenden Gesellschaft in Basel 81: 223 - 318" title="Syrphidae von Madagascar (Dipt.)." volume="81" year="1971">Keiser (1971)</bibRefCitation>
also noted that both taxa often co-occur. We, therefore, consider the presence or absence of the lobe on the anterior side of the metabasitarsus in the male as a polymorphism and consider
<taxonomicName lsidName="M. platytarsis" pageId="0" pageNumber="1" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">M. platytarsis</emphasis>
</taxonomicName>
Curran, 1929 a junior synonym of
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
Curran, 1929.
</paragraph>
</subSubSection>
</treatment>
</document>