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<mods:title id="0BC938D2006F0AD6867F5EDCAF589683">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:namePart id="CF3B62C5B4AA1D2C3253714D0FCD7BEA">ALEKSEEV, VITALII</mods:namePart>
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<mods:affiliation id="D0775B9490533DFCF75E3977DB0D3587">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia.</mods:affiliation>
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<mods:affiliation id="7C1D67C8F571749E2D44D1E22F462CB5">Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<subSubSection id="C3473DA5FFFD665EFF506F8AB8D553E2" box="[151,926,1483,1510]" pageId="29" pageNumber="230" type="nomenclature">
<paragraph id="8BE26E2EFFFD665EFF506F8AB8D553E2" blockId="29.[151,926,1483,1546]" box="[151,926,1483,1510]" pageId="29" pageNumber="230">
<heading id="D0AAD942FFFD665EFF506F8AB8D553E2" bold="true" box="[151,926,1483,1510]" fontSize="11" level="1" pageId="29" pageNumber="230" reason="1">
<taxonomicName id="4C5D15ADFFFD665EFF506F8AB8D553E2" authority="Alekseev et Nabozhenko, 2023" authorityName="Alekseev et Nabozhenko" authorityYear="2023" box="[151,926,1483,1510]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="232" phylum="Arthropoda" rank="species" species="auriborussiensis">
<emphasis id="B929B23CFFFD665EFF506F8AB8D553E2" bold="true" box="[151,926,1483,1510]" pageId="29" pageNumber="230">
<emphasis id="B929B23CFFFD665EFF506F8ABA0453E1" bold="true" box="[151,335,1483,1510]" italics="true" pageId="29" pageNumber="230">Zopheromimus</emphasis>
<emphasis id="B929B23CFFFD665EFE906F8AB95553E1" bold="true" box="[343,542,1483,1510]" italics="true" pageId="29" pageNumber="230">auriborussiensis</emphasis>
<bibRefCitation id="EFCC13DFFFFD665EFDE46F8AB8D553E2" author="Alekseev, V. I. &amp; Nabozhenko, M. V." box="[547,926,1483,1510]" pageId="29" pageNumber="230" pagination="435 - 445" refId="ref24250" refString="Alekseev, V. I. &amp; Nabozhenko, M. V. (2023) News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae). Zootaxa, 5297 (3), 435 - 445. https: // doi. org / 10.11646 / zootaxa. 5297.3.8" type="journal article" year="2023">Alekseev et Nabozhenko, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3473DA5FFFD665EFF506FAEBA63500D" box="[151,296,1519,1546]" pageId="29" pageNumber="230" type="description">
<paragraph id="8BE26E2EFFFD665EFF506FAEBA63500D" blockId="29.[151,926,1483,1546]" box="[151,296,1519,1546]" pageId="29" pageNumber="230">
(
<figureCitation id="136672ABFFFD665EFF586FAEBA54500D" box="[159,287,1519,1546]" captionStart="FIGURES 4244" captionStartId="30.[151,264,1174,1198]" captionTargetBox="[151,1435,179,1148]" captionTargetId="figure-21@30.[151,1435,179,1148]" captionTargetPageId="30" captionText="FIGURES 4244. Zopheromimus auriborussiensis in Baltic amber, specimen No. JDC-11965 [JDC]: 42—habitus in dorsal view; 43—idem, habitus in ventral view; 44—idem, habitus in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14033681" httpUri="https://zenodo.org/record/14033681/files/figure.png" pageId="29" pageNumber="230">Figs 4244</figureCitation>
)
</paragraph>
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<subSubSection id="C3473DA5FFFD665EFF506C76BF1B509E" pageId="29" pageNumber="230" type="materials_examined">
<paragraph id="8BE26E2EFFFD665EFF506C76BF1B509E" blockId="29.[151,1437,1591,1869]" pageId="29" pageNumber="230">
<emphasis id="B929B23CFFFD665EFF506C76BACA5056" bold="true" box="[151,385,1591,1617]" pageId="29" pageNumber="230">Material examined.</emphasis>
<materialsCitation id="3B356473FFFD665EFE4E6C79BED65056" box="[393,1437,1591,1618]" collectionCode="JDC" location="Baltic" pageId="29" pageNumber="230" specimenCode="JDC-11965" specimenCount="2" specimenCount-adult="1">
<specimenCount id="9D5BA5A7FFFD665EFE4E6C79BAF35055" box="[393,440,1592,1618]" pageId="29" pageNumber="230">One</specimenCount>
specimen with collection number
<specimenCode id="DBFBC655FFFD665EFCF96C76B8F55055" box="[830,958,1591,1618]" pageId="29" pageNumber="230">JDC-11965</specimenCode>
[
<collectionCode id="ED4CF6EBFFFD665EFC0A6C79BF4D5055" box="[973,1030,1592,1618]" pageId="29" pageNumber="230">JDC</collectionCode>
],
<location id="8E8238F5FFFD665EFBDE6C76BF175056" LSID="urn:lsid:plazi:treatment:03F4DF38FFFD665EFF506F8ABA5B514A:8E8238F5FFFD665EFBDE6C76BF175056" box="[1049,1116,1591,1617]" name="Baltic" pageId="29" pageNumber="230">Baltic</location>
amber;
<specimenCount id="9D5BA5A7FFFD665EFB7E6C76BFBF5056" box="[1209,1268,1591,1617]" pageId="29" pageNumber="230" type="adult">adult</specimenCount>
, sex unknown.
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A complete beetle is included in a transparent, yellow amber piece with dimensions of 20×
<quantity id="4CA5C3CBFFFD665EFB546C1DBFAA5071" box="[1171,1249,1628,1654]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="230" unit="mm" value="17.0">17 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFFD665EFEE26C3EBA23509D" box="[293,360,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="230" unit="mm" value="5.0">5 mm</quantity>
, preserved without supplementary fixation. Syninclusions: absent.
</paragraph>
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<subSubSection id="C3473DA5FFFD665EFF006CE5BA5B514A" pageId="29" pageNumber="230" type="discussion">
<paragraph id="8BE26E2EFFFD665EFF006CE5BA5B514A" blockId="29.[151,1437,1591,1869]" pageId="29" pageNumber="230">
<emphasis id="B929B23CFFFD665EFF006CE5BA4D50B9" bold="true" box="[199,262,1700,1726]" pageId="29" pageNumber="230">Note.</emphasis>
Body length of beetle JDC-11965 (including visible part of head)
<quantity id="4CA5C3CBFFFD665EFC206CE5BF7450B9" box="[999,1087,1700,1726]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="29" pageNumber="230" unit="mm" value="4.9">4.9 mm</quantity>
. This specimen has 5-5-4 tarsal formula, protruding hemispherical eyes, and 11-segmented antennae with weakly developed 2-segmented club. Specimen JDC-11965 shares with the
<typeStatus id="54E6D08CFFFD665EFD8C6CAAB9E65102" box="[587,685,1771,1797]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5D15ADFFFD665EFD116CADB8E55102" authorityName="Alekseev et Nabozhenko" authorityYear="2023" box="[726,942,1771,1797]" genus="Zopheromimus" pageId="29" pageNumber="230" rank="species" species="auriborussiensis">
<emphasis id="B929B23CFFFD665EFD116CADB8E55102" box="[726,942,1771,1797]" italics="true" pageId="29" pageNumber="230">Z. auriborussiensis</emphasis>
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other external characters and is considered conspecific. The smaller body size (
<quantity id="4CA5C3CBFFFD665EFDF46D51B9C6512D" box="[563,653,1808,1834]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="29" pageNumber="230" unit="mm" value="4.9">4.9 mm</quantity>
in contrast to
<quantity id="4CA5C3CBFFFD665EFCF26D4EB8C6512D" box="[821,909,1807,1834]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="29" pageNumber="230" unit="mm" value="5.3">5.3 mm</quantity>
in the
<typeStatus id="54E6D08CFFFD665EFC186D4EBF08512E" box="[991,1091,1807,1833]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
) is interpreted as intraspecific variability.
</paragraph>
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