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<document id="66107D2526F34E88C66A2A621007681A" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFEB501DFF23FBD6FB664682" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus barbatus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="28" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="955043A9EE32E0D3E4824DFA8725E132" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9D3074D704284E4EE0AAC3FC63FD8E3F">
<mods:title id="957A025EEC97B24FE8F6E11E6105B0FA">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="86843CCBF863A179A47C74A5FD87CC22" type="personal">
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<mods:namePart id="9E38C32DEBAD1DDFF6766DCF4B242EB4">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="2535C07CABE57A2404BC54D6158C1A5F">text</mods:typeOfResource>
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<mods:title id="3F1DE45D64624AAA95A77FCBE570878D">Zootaxa</mods:title>
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<mods:part id="98ADF8BC36A69A048FAEAE43078B02F0">
<mods:date id="9D0461CF90801DDB1DA3B742536C3962">2024</mods:date>
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<mods:number id="5B258349787F540CE0F43CC19877E9A4">2024-09-04</mods:number>
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<mods:number id="4122523146793091A140D81D81B638A6">5505</mods:number>
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<mods:url id="CBB8E194D0F9579CEDFA1E07F3983A15">http://dx.doi.org/10.11646/zootaxa.5505.1.1</mods:url>
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<mods:classification id="825B27513C58E0CCAFC1F65E9E6C5B0D">journal article</mods:classification>
<mods:identifier id="7CFC3FD4739C70AD3428B3E259346708" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
<mods:identifier id="F34B91D806D14AE32B5F14498167AEA8" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFEB501DFF23FBD6FB664682" ID-DOI="http://doi.org/10.5281/zenodo.13746035" ID-Zenodo-Dep="13746035" LSID="urn:lsid:plazi:treatment:03E7FF50FFEB501DFF23FBD6FB664682" httpUri="http://treatment.plazi.org/id/03E7FF50FFEB501DFF23FBD6FB664682" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="C3541DCDFFEB501BFF23FBD6FD094046" box="[151,716,1050,1077]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BF14E46FFEB501BFF23FBD6FD094046" blockId="27.[151,716,1050,1113]" box="[151,716,1050,1077]" pageId="27" pageNumber="28">
<heading id="D0B9F92AFFEB501BFF23FBD6FD094046" bold="true" box="[151,716,1050,1077]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B93A9254FFEB501BFF23FBD6FD094046" bold="true" box="[151,716,1050,1077]" pageId="27" pageNumber="28">
<taxonomicName id="4C4E35C5FFEB501BFF23FBD6FD914047" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,596,1050,1077]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="barbatus" status="comb. nov.">
<emphasis id="B93A9254FFEB501BFF23FBD6FE284046" bold="true" box="[151,493,1050,1077]" italics="true" pageId="27" pageNumber="28">Anhoraeomorphus barbatus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFEB501BFDD5FBD7FD094046" box="[609,716,1051,1077]" pageId="27" pageNumber="28" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FBF3FEF0402A" box="[151,309,1087,1113]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BF14E46FFEB501BFF23FBF3FEF0402A" blockId="27.[151,716,1050,1113]" box="[151,309,1087,1113]" pageId="27" pageNumber="28">
(
<figureCitation id="137552C3FFEB501BFF2BFBF3FEEB402A" box="[159,302,1087,1113]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 99–106</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BF14E46FFEB501BFF23FB4FFDA140E8" blockId="27.[151,612,1155,1179]" box="[151,612,1155,1179]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB5E6968FFEB501BFF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="27" pageNumber="28">
<treatmentCitation id="0AEF6857FFEB501BFF23FB4FFDA540E8" author="Franz, H." box="[151,608,1155,1179]" page="167" pageId="27" pageNumber="28" year="1986">
<taxonomicName id="4C4E35C5FFEB501BFF23FB4FFDA540E8" ID-CoL="3MNJH" authority="Franz, 1986 b: 167" authorityName="Franz" authorityPageNumber="167" authorityYear="1986" box="[151,608,1155,1179]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFEB501BFF23FB4FFE5B40E8" box="[151,414,1155,1179]" italics="true" pageId="27" pageNumber="28">Horaeomorphus barbatus</emphasis>
<bibRefCitation id="EFDF33B7FFEB501BFE10FB48FDA540E8" author="Franz, H." box="[420,608,1155,1179]" pageId="27" pageNumber="28" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 167</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FB05FB004254" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BF14E46FFEB501BFF23FB05FD3A413C" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF23FB05FE5A4090" bold="true" box="[151,415,1225,1251]" pageId="27" pageNumber="28">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFEB501BFE10FB05FD164158" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="27" pageNumber="28" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFEB501BFE10FB05FD784090" bold="true" box="[420,701,1225,1251]" pageId="27" pageNumber="28">
<typeStatus id="54F5F0E4FFEB501BFE10FB05FDD44090" box="[420,529,1225,1251]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFEB501BFDA9FB05FD714090" box="[541,692,1225,1251]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFEB501BFD7DFB06FD254097" box="[713,736,1226,1252]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
, four labels: “Madagaskar,Plateau / du Tampoketsa / lg.
<collectorName id="26BB2B90FFEB501BFA8AFB05FA594090" box="[1342,1436,1225,1251]" pageId="27" pageNumber="28">H.Franz</collectorName>
1969” [white, printed], “
<specimenCount id="9D4885CFFFEB501BFE1EFB21FE054174" box="[426,448,1261,1287]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
” [white, printed], “
<taxonomicName id="4C4E35C5FFEB501BFD23FB21FC8B4174" box="[663,846,1261,1287]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ barbatus m. / det.
<determinerName id="7E2DA282FFEB501BFB90FB21FB474174" box="[1060,1154,1261,1287]" pageId="27" pageNumber="28">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFEB501BFDDCFADDFD0E4158" box="[616,715,1297,1323]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFEB501BFD69FADDFD3E413C" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFEB501BFD69FADDFC904158" bold="true" box="[733,853,1297,1323]" pageId="27" pageNumber="28">
<typeStatus id="54F5F0E4FFEB501BFD69FADDFC904158" box="[733,853,1297,1323]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
(see Remarks):
<specimenCount id="9D4885CFFFEB501BFBBEFADDFBF3415F" box="[1034,1078,1297,1324]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D4885CFFFEB501BFBF5FADDFBAE415F" box="[1089,1131,1297,1324]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
, same data as for holotype, except for yellowish “
<typeStatus id="54F5F0E4FFEB501BFE38FAF9FDF5413C" box="[396,560,1333,1359]" pageId="27" pageNumber="28" type="paratype">PARATYPUS</typeStatus>
” labels (
<collectionCode id="ED5FD683FFEB501BFD24FAF9FD36413C" box="[656,755,1333,1359]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FA95FB004254" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FA95FE5A4100" bold="true" box="[199,415,1369,1395]" pageId="27" pageNumber="28">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFEB501BFE5AFA95FDF84100" box="[494,573,1369,1395]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) slender; head (
<figureCitation id="137552C3FFEB501BFD5EFA95FC464100" box="[746,899,1369,1395]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 100–101</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFEB501BFEA2FAB1FEAC41E4" box="[278,361,1405,1431]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large antebasal pits; antennomeres 3–7 (
<figureCitation id="137552C3FFEB501BFF66FA6DFEE041C8" box="[210,293,1441,1467]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="27" pageNumber="28">Fig. 94</figureCitation>
) each slightly elongate; all femora similarly slender; hind tibiae in male unmodified; aedeagus (
<figureCitation id="137552C3FFEB501BFADEFA6DFF3F41AC" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 103–106</figureCitation>
) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view strongly curved dorsally, distolateral endophallic sclerites narrowly separated, each elongate and with subtriangular apex, each paramere with group of numerous subapical and apical setae occupying about half of its length.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501DFF73F9FDFDAB45A2" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8BF14E46FFEB501BFF73F9FDFEAE4203" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73F9FDFEB04238" bold="true" box="[199,373,1585,1611]" pageId="27" pageNumber="28">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFEB501BFDA8F9FDFDAE4238" box="[540,619,1585,1611]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) moderately dark brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFEB501BFF75F999FEA24203" box="[193,359,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="2.3049999999999997" metricValueMax="2.31" metricValueMin="2.3" pageId="27" pageNumber="28" unit="mm" value="2.3049999999999997" valueMax="2.31" valueMin="2.3">2.30–2.31 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73F9B5FDC243FC" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
Head (
<figureCitation id="137552C3FFEB501BFEA7F9B5FEB642E0" box="[275,371,1657,1683]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 100</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFEB501BFC07F9B5FB9F42E7" box="[947,1114,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="27" pageNumber="28" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFEB501BFB28F9B5FA8142E7" box="[1180,1348,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="27" pageNumber="28" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFEB501BFEF8F951FE6842C4" box="[332,429,1693,1719]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 101</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval with distinct posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFEB501BFF2BF8E1FF3A4334" box="[159,255,1837,1863]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 102</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFEB501BFC3EF8E1FBF5433B" box="[906,1072,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.1" metricValueMin="1.08" pageId="27" pageNumber="28" unit="mm" value="1.09" valueMax="1.1" valueMin="1.08">1.08–1.10 mm</quantity>
, antennomeres 1–7 each weakly elongate, 8–9 each about as long as broad, 10 slightly transverse, 11 shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<caption id="DF311ECEFFEC501CFF23FA05FE7F425A" ID-DOI="http://doi.org/10.5281/zenodo.13745957" ID-Zenodo-Dep="13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1481,1505]" targetBox="[151,1428,181,1441]" targetPageId="28" targetType="figure">
<paragraph id="8BF14E46FFEC501CFF23FA05FE7F425A" blockId="28.[151,1436,1481,1578]" pageId="28" pageNumber="29">
<emphasis id="B93A9254FFEC501CFF23FA05FEA14192" bold="true" box="[151,356,1481,1505]" pageId="28" pageNumber="29">FIGURES 99–106.</emphasis>
<taxonomicName id="4C4E35C5FFEC501CFED9FA05FD214191" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[365,740,1481,1506]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFEC501CFED9FA05FD4B4192" box="[365,654,1481,1505]" italics="true" pageId="28" pageNumber="29">Anhoraeomorphus barbatus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFEC501CFBEFFA05FBB14192" bold="true" box="[1115,1140,1481,1505]" pageId="28" pageNumber="29">99</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEC501CFF14FA21FF034276" bold="true" box="[160,198,1517,1541]" pageId="28" pageNumber="29">100</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEC501CFC23FA21FC784276" bold="true" box="[919,957,1517,1541]" pageId="28" pageNumber="29">101</emphasis>
); antenna (
<emphasis id="B93A9254FFEC501CFB9BFA21FB904276" bold="true" box="[1071,1109,1517,1541]" pageId="28" pageNumber="29">102</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEC501CFA88FA21FA514276" bold="true" box="[1340,1428,1517,1541]" pageId="28" pageNumber="29">103, 105</emphasis>
) and lateral (
<emphasis id="B93A9254FFEC501CFEA4F9DDFEAC425A" bold="true" box="[272,361,1553,1577]" pageId="28" pageNumber="29">104, 106</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEC501CFF73F9B0FEEA4355" blockId="28.[151,1437,1659,1974]" pageId="28" pageNumber="29">
Pronotum bell-shaped, broadest near anterior third; PL
<quantity id="4CB6E3A3FFEC501CFCF2F9B0FC2B42E5" box="[838,1006,1660,1686]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" metricValueMax="6.3" metricValueMin="6.1" pageId="28" pageNumber="29" unit="mm" value="0.62" valueMax="0.63" valueMin="0.61">0.61–0.63 mm</quantity>
, PW
<quantity id="4CB6E3A3FFEC501CFB84F9B7FB1C42E5" box="[1072,1241,1659,1686]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="28" pageNumber="29" unit="mm" value="0.52" valueMax="0.54" valueMin="0.5">0.50–0.54 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large round pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFEC501CFF73F8FCFA5E43E1" blockId="28.[151,1437,1659,1974]" pageId="28" pageNumber="29">
Elytra together oval, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFEC501CFC2CF8FCFB844339" box="[920,1089,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="28" pageNumber="29" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.25–1.30 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEC501CFB32F8FCFB354339" box="[1158,1264,1840,1866]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="28" pageNumber="29" unit="mm" value="0.88">0.88 mm</quantity>
, EI 1.43–1.49; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures very dense but extremely shallow and diffuse, so that surface of elytra is slightly less glossy than pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEC501CFF73F850FB3943C5" blockId="28.[151,1437,1659,1974]" box="[199,1276,1948,1974]" pageId="28" pageNumber="29">Legs moderately long, slender, unmodified except weakly curved apices of pro- and mesotibiae.</paragraph>
<paragraph id="8BF14E46FFED501DFF73FF5BFB2045FA" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
Aedeagus (
<figureCitation id="137552C3FFED501DFEF0FF5BFE2544C2" box="[324,480,151,177]" captionStart="FIGURES 99–106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99–106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="29" pageNumber="30">Figs 103–106</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFED501DFD37FF5BFD2F44C2" box="[643,746,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="29" pageNumber="30" unit="mm" value="0.38">0.38 mm</quantity>
; in ventral view broadest in sub-basal third, distally gradually narrowing and in subapical region rapidly narrowing towards broadly subtriangular apex with rounded tip; in lateral view apical region of dorsal wall strongly curved dorsally; endophallus with nearly symmetrical and conspicuously broad distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites narrowly separated, each with subtriangular apex; parameres broad and their apices slightly projecting beyond aedeagal apex, each paramere with twisted short distal region appearing in ventral view as an apical ‘loop’, and with apical-subapical group of 12 short setae proximally reaching middle of parameral length.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FE5FFDAB45A2" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
Female. Externally similar to male. BL
<quantity id="4CB6E3A3FFED501DFD34FE5FFD2245DE" box="[640,743,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="29" pageNumber="30" unit="mm" value="2.33">2.33 mm</quantity>
; HL
<quantity id="4CB6E3A3FFED501DFCA9FE5FFC4745DE" box="[797,898,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="29" pageNumber="30" unit="mm" value="0.43">0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFED501DFC76FE5EFBED45DE" box="[962,1064,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="29" pageNumber="30" unit="mm" value="0.45">0.45 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFED501DFBDEFE5FFB1445DE" box="[1130,1233,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="29" pageNumber="30" unit="mm" value="1.03">1.03 mm</quantity>
; PL
<quantity id="4CB6E3A3FFED501DFAB6FE5FFAAD45DE" box="[1282,1384,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="mm" value="0.6">0.60 mm</quantity>
, PW
<quantity id="4CB6E3A3FFED501DFF23FE7AFF3B45A2" box="[151,254,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="29" pageNumber="30" unit="mm" value="0.53">0.53 mm</quantity>
; EL 1.30, EW
<quantity id="4CB6E3A3FFED501DFE17FE7BFDCC45A2" box="[419,521,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="29" pageNumber="30" unit="mm" value="0.88">0.88 mm</quantity>
, EI 1.49.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73FE16FC8A4586" box="[199,847,474,501]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BF14E46FFED501DFF73FE16FC8A4586" blockId="29.[151,1437,151,753]" box="[199,847,474,501]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FE16FEA44587" bold="true" box="[199,353,474,500]" pageId="29" pageNumber="30">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFED501DFE75FE17FD8D4586" box="[449,584,475,501]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
(Analamanga Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73FE33FB664682" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BF14E46FFED501DFF73FE33FEDA464E" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FE33FEFE466A" bold="true" box="[199,315,511,537]" pageId="29" pageNumber="30">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFED501DFEF7FE33FD45466A" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,640,511,537]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFED501DFEF7FE33FD45466A" box="[323,640,511,537]" italics="true" pageId="29" pageNumber="30">Anhoraeomorphus barbatus</emphasis>
</taxonomicName>
is unremarkable and examination of aedeagus is necessary to identify this species.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FD8BFB664682" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
<materialsCitation id="3B26441BFFED501DFF73FD8BFB5A4682" collectionCode="NHMW" collectorName="All &amp; Franz" elevation="1500" location="Seehohe" pageId="29" pageNumber="30" specimenCount="6" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="EFDF33B7FFED501DFF73FD8BFEA34612" author="Franz, H." box="[199,358,583,609]" pageId="29" pageNumber="30" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
mentions
<specimenCount id="9D4885CFFFED501DFE6EFD8BFD414612" box="[474,644,583,609]" count="4" pageId="29" pageNumber="30" type="generic">four specimens</specimenCount>
included in the type series, of which
<specimenCount id="9D4885CFFFED501DFB90FD8BFB4C4612" box="[1060,1161,583,609]" count="1" pageId="29" pageNumber="30" type="male">one male</specimenCount>
was fixed as a
<typeStatus id="54F5F0E4FFED501DFA87FD8BFA5C4612" box="[1331,1433,583,609]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
<collectorName id="26BB2B90FFED501DFF23FDA7FF7E46F6" box="[151,187,619,645]" pageId="29" pageNumber="30">All</collectorName>
specimens are preserved at
<collectionCode id="ED5FD683FFED501DFDB1FDA7FDA446F6" box="[517,609,619,645]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="30" type="Museum">NHMW</collectionCode>
, but
<specimenCount id="9D4885CFFFED501DFD28FDA7FC4046F6" box="[668,901,619,645]" count="1" pageId="29" pageNumber="30" type="female" typeStatus="paratype">one paratype female</specimenCount>
is clearly different from remaining specimens and does not belong in
<taxonomicName id="4C4E35C5FFED501DFE19FD43FDF146DA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[429,564,655,681]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFED501DFE19FD43FDF146DA" box="[429,564,655,681]" italics="true" pageId="29" pageNumber="30">A. barbatus</emphasis>
</taxonomicName>
; it must be excluded from the type series. “Plateau du Tampoketsa” is the only locality given in label data, but in the original description
<collectorName id="26BB2B90FFED501DFCD6FD7FFC6746BE" box="[866,930,691,717]" pageId="29" pageNumber="30">Franz</collectorName>
adds more detailed information: “Plateau du Tampokeza {sic!} d’Ankazobe nächst der Forststation Manonkazo im ca.
<quantity id="4CB6E3A3FFED501DFC60FD1AFBEC4682" box="[980,1065,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">
<elevation id="0063A975FFED501DFC60FD1AFBEC4682" box="[980,1065,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</elevation>
</quantity>
<location id="8E91189DFFED501DFB84FD1BFB504682" LSID="urn:lsid:plazi:treatment:03E7FF50FFEB501DFF23FBD6FB664682:8E91189DFFED501DFB84FD1BFB504682" box="[1072,1173,727,753]" name="Seehohe" pageId="29" pageNumber="30">Seehöhe</location>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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